Philonthus indicus, Kraatz, 1859: 113

Taphylininae, Iochini, Aorothiini, Thiini, Latyprosopini, Taphylinini, Mblyopinina, Nisolinina, Yptiomina, Hilonthina & Herman, Lee H., 2001, Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina), Bulletin of the American Museum of Natural History 2441 (265), pp. 2435-3021 : 2445-2449

indicus Kraatz, 1859: 113 ( Diochus ; Type locality: India orientalis).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; synonym of antennatus ).

— Cameron, 1931: 360 ( Diochus ; New Guinea).

— Cameron, 1932: 46 ( Diochus ; synonym of antennatus ).

— Cameron, 1934d: 79 ( Diochus ; synonym of antennatus ).

apicipennis Cameron, 1922 , see: nanus Erichson, 1839 .

astutus Casey, 1906: 433 ( Diochus ; Type locality: South Africa: Wellington ).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

- DISTRIBUTION: South Africa.

bicolor Scheerpeltz, 1974: 131 ( Diochus ; Type locality: Cape Prov.: Cape Peninsula ,

Cape Point Nature Reserve).

- DISTRIBUTION: South Africa.

bicoloripennis Coiffait, 1981f: 333 ( Diochus ; Type locality: South Andaman, Jolly Boy Island).

- DISTRIBUTION: Andaman Islands.

borneensis Cameron, 1933b: 344 ( Diochus ; Type locality: British North Borneo: Kenokok, 3300 feet).

- DISTRIBUTION: Borneo.

brevipennis Casey, 1906 , see: schaumii Kraatz, 1860 .

brunniventris Cameron, 1938: 8 ( Diochus ; Type locality: Belgian Congo: Eala ).

- DISTRIBUTION: Congo.

communis Coiffait, 1968: 159 ( Diochus ; Type locality: Gabon: Belinga ).

- DISTRIBUTION: Gabon.

conicollis Motschulsky, 1858: 658 ( Rhegmatocerus ; Type locality: Indes orientales).

— Bernhauer, 1902b: 32 ( Diochus ; Ceylon).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Bernhauer, 1922c: 231 ( Diochus ; Formosa).

— Cameron, 1932: 45 ( Diochus ; characters; Ceylon; India).

— Scheerpeltz, 1933: 1324 ( Diochus ; catalog).

— Shibata, 1973a: 131 ( Diochus ; Taiwan).

— Coiffait, 1977b: 206 ( Diochus ; Bhutan).

— Coiffait, 1981f: 329 ( Diochus ; Andaman Islands).

— Coiffait, 1982: 27 ( Diochus ; Nepal).

— Frank, 1982: 35 ( Diochus ; list of parasites).

- DISTRIBUTION: “East Indies”, Sri Lanka, India, Bhutan, Andaman Islands, Taiwan. major Kraatz, 1859: 113 ( Diochus ; Type locality: India orientalis; Ceylan).

— Bernhauer, 1902b: 32 ( Diochus ; synonym of conicollis ).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; synonym of conicollis ).

— Cameron, 1932: 45 ( Diochus ; synonym of conicollis ).

— Cameron, 1934d: 79 ( Diochus ; synonym of conicollis ).

cristobalensis Coiffait, 1981 g: 295 ( Diochus ; Type locality: Ile Galapagos : Ile San Cristobal, montagne près Progreso).

- DISTRIBUTION: Galapagos Islands.

debilis Coiffait, 1968: 159 ( Diochus ; Type locality: Gabon: Belinga ).

- DISTRIBUTION: Gabon.

dilutus Reitter, 1908 , see: staudingeri Kraatz, 1860 .

divisus Fauvel, 1877: 235 ( Diochus ; Type locality: Australie, Nouvelle Galles du Sud).

— Olliff, 1887: 473 ( Diochus ; characters; Australia).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Lea, 1925: 220 ( Diochus ; characters; Australia).

- DISTRIBUTION: Australia.

dundoensis Cameron, 1959: 115 ( Diochus ; Type locality: Angola: Dundo ; Sierra Leone: Freetown).

— Tottenham, 1961: 203 ( Diochus ; Senegal).

- DISTRIBUTION: Angola, Sierra Leone, Senegal.

flandrianus Bernhauer, 1932b: 145 ( Diochus ; Type locality: Equateur: Flandria ).

— Scheerpeltz, 1933: 1324 ( Diochus ; catalog).

- DISTRIBUTION: Congo.

flavicans Sharp, 1876 , see: nanus Erichson, 1839 .

formicetorum Bernhauer, 1927e: 244 ( Diochus ; Type locality: Argentinien: Prov. Santa Fe, Fives Lilles ).

— Scheerpeltz, 1933: 1324 ( Diochus ; catalog).

- DISTRIBUTION: Argentina.

franzi Coiffait, 1981 g: 294 ( Diochus ; Type locality: Ile Galapagos: Ile Santiago).

- DISTRIBUTION: Galapagos Islands.

gabonicus Coiffait, 1968: 157 ( Diochus ; Type locality: Gabon: Makokou ).

- DISTRIBUTION: Gabon.

indicus Kraatz, 1859 , see: antennatus Motschulsky, 1858 .

inornatus Sharp, 1885 , see: nanus Erichson, 1839 .

isabelae Coiffait, 1981 g: 293 ( Diochus ; Type locality: Ile Galapagos: Ile Isabela,

Sierra Negra).

- DISTRIBUTION: Galapagos Islands.

japonicus Cameron, 1930c: 206 ( Diochus ; Type locality: Kobe).

— Scheerpeltz, 1933: 1324 ( Diochus ; catalog).

— Nakane, 1963a: 90 ( Diochus ; characters; habitus photograph; Japan).

— J. Li, 1993: 32 ( Diochus ; China).

- DISTRIBUTION: Japan, China.

libanoticus Fagel, 1966a: 43 ( Diochus ; Type locality: Liban: Kartaba, vallée du Nahr Ibrahim, 1000 m).

— Coiffait, 1972: 370 ( Diochus ; characters; Lebanon).

- DISTRIBUTION: Lebanon.

longicornis Sharp, 1876 , see: nanus Erichson, 1839 .

longicornis Cameron, 1952 , see: petilus Herman, 2001 .

longus Lea, 1929: 203 ( Diochus ; Type locality: Western Australia: Bunbury ).

— Scheerpeltz, 1933: 1324 ( Diochus ; catalog).

- DISTRIBUTION: Australia.

machadoi Cameron, 1959: 115 ( Diochus ; Type locality: Angola: Dundo ; Vila Luso;

Luachimo Forest).

- DISTRIBUTION: Angola.

maculicollis Fauvel, 1891 , see: nanus Erichson, 1839 .

major Kraatz, 1859 , see: conicollis Motschulsky, 1858 .

montguilloni Coiffait, 1987: 497 ( Diochus ; replacement name for testaceus Coiffait ).

- DISTRIBUTION: Galapagos Islands.

testaceus Coiffait, 1981 g: 296 ( Diochus ; [preoccupied]; Type locality: Ile Galapagos:

Ile Santiago, Montagne).

— Coiffait, 1987: 497 ( Diochus ; synonym of montguilloni ).

nanus Erichson, 1839b: 301 ( Diochus ; Type locality: Columbia; [Note: See lectotype designation by Smetana, 1977c]. Also cited from Americae meridionalis regiones).

— Chevrolat and Fauvel, 1863: 430 ( Diochus ; characters; Cuba, Colombia).

— Solsky, 1872: 308 ( Diochus ; Cayenne).

— Lynch, 1884: 172 ( Diochus ; characters; Argentina).

— Sharp, 1885: 466 ( Diochus ; notes; Guatemala; Panama; Colombia).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Blackwelder, 1943: 455 ( Diochus ; characters; Cuba; Jamaica; Dominican Republic; Puerto Rico; St. Croix; Guadeloupe; St. Lucia; Barbados; St. Vincent; Grenadines; Grenada; Trinidad). — Smetana, 1977c: 347 ( Diochus ; lectotype designation: Colombia; characters; notes).

— Frank, 1982: 35 ( Diochus ; list of parasites).

- DISTRIBUTION: Argentina, Brazil, French Guiana, Venezuela, Colombia, Panama, Nicaragua, Guatemala, Mexico, West Indies.

parvulus Kraatz, 1860: 27 ( Diochus ; Type locality: Bahia).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

longicornis Sharp, 1876: 184 ( Diochus ; Type locality: Tapajos).

— Sharp, 1887: 791 ( Diochus ; Mexico).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

vicinus Sharp, 1876: 185 ( Diochus ; Type locality: Tapajos).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

tarsalis Sharp, 1876: 185 ( Diochus ; Type locality: Tapajos).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Scheerpeltz, 1933: 1324 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

flavicans Sharp, 1876: 185 ( Diochus ; Type locality: Tapajos).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

inornatus Sharp, 1885: 466 ( Diochus ; Type locality: Nicaragua, Chontales; Panama, Bugaba 800–1500 feet, Tolé, Peña Blanca 3000–4000 feet).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

vilis Sharp, 1885: 467 ( Diochus ; Type locality: Guatemala, near the city).

— Bernhauer and Schubert, 1914: 320 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

maculicollis Fauvel, 1891: 106 ( Diochus ; proposed for the species Kraatz misidentified as nanus ; Type locality: Caracas; Colonie Tovar).

— Kraatz, 1860: 27 ( Diochus ; [Note: Misidentification: Bernhauer and Schubert, 1914: 319, cited nanus Kraatz as a synonym of maculicollis ; Kraatz did not describe the species as new, he attributed it to Erichson]; characters; Venezuela).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

perplexus Cameron, 1922: 116 ( Diochus ; Type locality: Grenada).

— Scheerpeltz, 1933: 1324 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

apicipennis Cameron, 1922: 116 ( Diochus ; Type locality: St. Vincent).

— Scheerpeltz, 1933: 1323 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

antennalis Cameron, 1922: 117 ( Diochus ; Type locality: Grenada).

— Scheerpeltz, 1933: 1323 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

pumilio Bernhauer, 1929 h: 193 ( Diochus ; Type locality: Mexiko: Veracruz).

— Scheerpeltz, 1933: 1324 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

ochraceus Cameron, 1926b: 347 ( Diochus ; Type locality: Siwaliks, Mothronwala).

— Cameron, 1932: 47 ( Diochus ; characters; India).

— Scheerpeltz, 1933: 1324 ( Diochus ; catalog).

- DISTRIBUTION: India.

octavii Fauvel, 1877: 235 ( Diochus ; Type locality: Australe , Victoria).

— Fauvel, 1878g: 535 ( Diochus ; Australia).

— Olliff, 1887: 472 ( Diochus ; characters; Australia).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Bernhauer, 1920: 16 ( Diochus ; Australia).

— Lea, 1925: 220 ( Diochus ; characters; Australia).

- DISTRIBUTION: Australia.

pallidiceps Casey, 1906 , see: schaumii Kraatz, 1860 .

papuanus Cameron, 1939b: 145 ( Diochus ; Type locality: New Guinea: Kokoda, altitude

1200 feet).

- DISTRIBUTION: New Guinea.

parvulus Kraatz, 1860 , see: nanus Erichson, 1839 .

perplexus Cameron, 1922 , see: nanus Erichson, 1839 .

petilus Herman, 2001: 43 ( Diochus ; replacement name for longicornis Cameron ).

- DISTRIBUTION: Congo.

longicornis Cameron, 1952a: 328 ( Diochus ; [preoccupied]; Type locality: Belgian Congo: Katanga, galerie forestière de la Kisanga).

— Herman, 2001: 43 ( Diochus ; synonym of petilus ).

proximus Cameron, 1952a: 328 ( Diochus ; Type locality: Belgian Congo: Massif des Kundelungu ).

- DISTRIBUTION: Congo.

pubiventris Lea, 1929: 203 ( Diochus ; Type locality: North Australia: Darwin; New South Wales: Tamworth ).

— Scheerpeltz, 1933: 1324 ( Diochus ; catalog).

— Scheerpeltz, 1935a: 649 ( Diochus ; Australia).

- DISTRIBUTION: Australia.

pulchellus Cameron, 1918: 84 ( Diochus ; Type locality: Singapore: Sembawang ).

— Cameron, 1921a: 375, 404 ( Diochus ; characters; Singapore).

— Scheerpeltz, 1933: 1324 ( Diochus ; catalog).

- DISTRIBUTION: Singapore.

pumilio Bernhauer, 1929 , see: nanus Erichson, 1839 .

punctipennis Motschulsky, 1858: 659 ( Rhegmatocerus ; Type locality: Indes orientales).

— Fauvel, 1898a: 117 ( Diochus ; Seychelles).

— H. Kolbe, 1910: 20 ( Diochus ; Seychelles).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Bernhauer, 1922b: 176 ( Diochus ; Seychelles).

— Cameron, 1932: 46 ( Diochus ; characters; India).

— Scheerpeltz, 1933: 1324 ( Diochus ; catalog).

— Cameron, 1934d: 79 ( Diochus ; type examined).

— Cameron, 1937: 4 ( Diochus ; Java).

— Scheerpeltz, 1957c: 233 ( Diochus ; Sumba).

— Coiffait, 1981f: 329 ( Diochus ; Andaman Islands).

— Lecoq, 1990: 189 ( Diochus ; Mascarene Islands).

- DISTRIBUTION: Mascarene Islands, Seychelles, Andaman Islands, India, Indonesia. santiagus Coiffait, 1981 g: 294 ( Diochus ; Type locality: Ile Galapagos: Ile Santiago).

- DISTRIBUTION: Galapagos Islands.

schaumii Kraatz, 1860: 27 ( Diochus ; Type locality: Louisiana; [Note: See lectotype desig-

nation by Smetana, 1982]. Also cited from Georgia).

— Sharp, 1885: 467 ( Diochus ; Louisiana; Guatemala; Nicaragua).

— Casey, 1906: 431 ( Diochus ; characters; Rhodes Island ; North Carolina; Missouri; Texas) .

— Blatchley, 1910: 400 ( Diochus ; characters; Indiana).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Notman, 1920b: 704 ( Diochus ; Florida).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

— Smetana, 1977c: 348 ( Diochus ; valid species; characters).

— Smetana, 1982: 29 ( Diochus ; lectotype designation: Louisiana; characters; Ontario; Quebec; Alabama; Arkansas; Connecticut; Florida; Georgia; Illinois; Indiana; Iowa; Kansas; Kentucky; Louisiana; Maine; Maryland; Massachusetts; Michigan; Mississippi; Missouri; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; Rhode Island; South Carolina; Texas; Virginia; West Virginia; Wisconsin) .

— Campbell and Davies, 1991: 123 ( Diochus ; checklist of species; Canada).

— Downie and Arnett, 1996: 395 ( Diochus ; characters; USA).

— Schmidt, 1999: 108 ( Diochus ; food for adults and larvae used for rearing species; rearing methods).

- DISTRIBUTION: Canada, USA, Guatemala, Nicaragua.

brevipennis Casey, 1906: 431 ( Diochus ; Type locality: Missouri; [Note: See lectotype designation by Smetana, 1982]. Also cited from Virginia; North Carolina, Asheville ).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

— Smetana, 1982: 29 ( Diochus ; lectotype designation: Mo. [Missouri]; synonym of schaumii ).

pallidiceps Casey, 1906: 432 ( Diochus ; Type locality: Texas, Austin; [Note: See lectotype designation by Smetana, 1982]).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

— Smetana, 1982: 29 ( Diochus ; lectotype designation; synonym of schaumii ).

thoracicus Casey, 1906: 432 ( Diochus ; Type locality: New York, Long Island; [Note:

See lectotype designation by Smetana, 1982]).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Blackwelder, 1943: 456 ( Diochus ; synonym of nanus ).

— Smetana, 1982: 29 ( Diochus ; lectotype designation; synonym of schaumii ).

senegalensis Fauvel, 1900b: 72 ( Diochus ; Type locality: Congo: Kinchassa ; Boma. Haut-Senegal: Badumbe).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

- DISTRIBUTION: Mali, Congo.

staudingeri Kraatz, 1860: 27 ( Diochus ; Type locality: Andalusia).

— Redtenbacher, 1874: 223 ( Diochus ; characters).

— Fauvel, 1878: 115 [= 1878a: 35] ( Diochus ; Algeria).

— Fauvel, 1886: 47 [= 1886a: 39] ( Diochus ; Algeria).

— Fauvel, 1897d: 289 ( Diochus ; Algeria).

— Fauvel, 1902b: 98 ( Diochus ; Algeria; Tunisia).

— Reitter, 1908: 108 ( Baptolinus ; characters; Andalusia).

— Bernhauer and Schubert, 1914: 319 ( Diochus ; catalog).

— Gridelli, 1931a: 157 ( Diochus ; characters; Algeria).

— Scheerpeltz, 1933: 1324 ( Diochus ; catalog).

— Coiffait, 1972: 369 ( Diochus ; characters; references; North Africa; Spain; France).

- DISTRIBUTION: Portugal, Spain, Algeria, Tunisia.

dilutus Reitter, 1908: 108 [= 1908a: 11] ( Diochus ; variety of staudingeri ; Type locality: Andalusien).

— Scheerpeltz, 1933: 1324 ( Diochus ; cited as dilutior ; synonym of staudingeri ).

— Coiffait, 1972: 369 ( Diochus ; cited as dilutior ; variety of staudingeri ; characters; Portugal).

tahitiensis Coiffait, 1976: 225 ( Diochus ; Type locality: Tahiti: Vallée Papenoo).

- DISTRIBUTION: Society Islands.