Gabrius fauveli, Smetana, 1963: 2

Taphylininae, Iochini, Aorothiini, Thiini, Latyprosopini, Taphylinini, Mblyopinina, Nisolinina, Yptiomina, Hilonthina & Herman, Lee H., 2001, Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina), Bulletin of the American Museum of Natural History 2441 (265), pp. 2435-3021 : 2709-2740

publication ID

https://doi.org/ 10.1206/0003-0090.265.1.5

persistent identifier

https://treatment.plazi.org/id/1D1E87B6-FEC0-FEB5-FF60-4EDFFC9E487F

treatment provided by

Marcus

scientific name

Gabrius fauveli
status

 

fauveli Smetana, 1963: 2 View in CoL ( Neobisnius ; Type locality: Louisiane).

— Frank, 1981: 20 ( Neobisnius ; synonym of ludicrus ).

maculicollis Scheerpeltz, 1972: 107 ( Neobisnius ; Type locality: Argentinien: Buenos Aires, Ezeiza ).

- DISTRIBUTION: Argentina.

madangensis Last, 1981 , see: Philonthus .

margianus Bordoni, 1975 , see: procerulus Gravenhorst, 1806 .

margipallens Jacquelin du Val, 1857, see: humilis Erichson, 1840 .

maximus Bernhauer, 1910b: 383 ( Neobisnius ; Type locality: Mexiko: Vera Cruz , Cordoba).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; catalog).

— Frank, 1981: 37 ( Neobisnius ; characters; Veracruz).

- DISTRIBUTION: Mexico.

mediopolitus Lea, 1929: 203 ( Neobisnius ; Type locality: Victoria: Melbourne ).

— Scheerpeltz, 1933: 1327 ( Neobisnius ; catalog).

- DISTRIBUTION: Australia.

mixtus Sharp, 1885: 460 ( Actobius ; Type locality: San Jerónimo, Baja Verapaz, Guatemala; [Note: See lectotype designation by Frank, 1981]. Originally cited from Mexico, Cordova, Vera Cruz, Tehuacan; Guatemala, San Geronimo).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; catalog).

— Bierig, 1933: 56 ( Neobisnius ; characters; Panama).

Frank, 1981: 24 ( Neobisnius ; lectotype designation: San Jerónimo , Baja Verapaz, Guatemala; characters; Puebla ; Veracruz; San Luis Potosi; Guatemala) .

- DISTRIBUTION: Mexico, Guatemala, Panama.

moestus Sharp, 1887: 790 ( Actobius ; Type locality: Mexico, near the city).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; catalog).

— Frank, 1981: 45 ( Neobisnius ; characters; Mexico).

- DISTRIBUTION: Mexico.

mundus Last, 1981 , see: Philonthus .

musonoiensis Levasseur, 1969: 881 ( Neobisnius ; Type locality: Katanga: Musonoie, région de Kolwezi).

- DISTRIBUTION: Congo.

nigripes Bernhauer, 1941a: 227 ( Neobisnius ; Type locality: Fukien: Kwangtseh; [Note: See lectotype designation by Gusarov, 1995a]).

— Coiffait, 1974: 331 ( Neobisnius ; characters; China).

— Gusarov, 1995a: 92 [= 1996: 55] ( Neobisnius ; lectotype designation; characters).

- DISTRIBUTION: China.

nigritulus Cameron, 1922 : [Note: Nomen nudum, Articles 12.1, 13.1.1; Scheerpeltz, 1933: 1327, and Blackwelder, 1944: 131, listed this species as described by Cameron, 1922: 119; I cannot find the species there or elsewhere.

nigrocoeruleus Cameron, 1922: 118 ( Neobisnius ; Type locality: Haiti).

— Scheerpeltz, 1933: 1327 ( Neobisnius ; cited as nigrocaeruleus ; catalog).

— Blackwelder, 1943: 441 ( Erichsonius ; synonym of humilis ).

— Blackwelder, 1944: 131 ( Erichsonius ; synonym of humilis ).

— Frank, 1981: 26 ( Neobisnius ; valid species; characters; Jamaica; Cuba; Haiti; Dominican Republic).

- DISTRIBUTION: West Indies.

carbonarius Bierig, 1933: 49 ( Neobisnius ; Type locality: Cuba: Prov. P. del Rio, Sierra del Rosario , Cayajabos).

— Blackwelder, 1943: 441 ( Erichsonius ; synonym of humilis ).

— Blackwelder, 1944: 131 ( Erichsonius ; synonym of humilis ).

— Frank, 1981: 26 ( Neobisnius ; synonym of nigrocoeruleus ).

nitidulus Sharp, 1885: 460 ( Actobius ; Type locality: Guatemala City, Department of Guatemala, Guatemala; [Note: See lectotype designation by Frank, 1981]. Originally cited from Guatemala, near the city, San Geronimo).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; catalog).

Frank, 1981: 34 ( Neobisnius ; lectotype designation: Guatemala City, Department of Guatemala, Guatemala; characters; Veracruz; Guatemala; El Salvador) .

- DISTRIBUTION: El Salvador, Guatemala, Mexico.

politus Sharp, 1885: 461 ( Actobius ; Type locality: Mexico).

— Sharp, 1887: 790 ( Actobius ; Guatemala).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; catalog).

— Frank, 1981: 34 ( Neobisnius ; synonym of nitidulus ).

nothocreatus Frank, 1981: 11 ( Neobisnius ; Type locality: California: Calaveras Co.: Mokelumne Hill ).

- DISTRIBUTION: USA.

nouristanicus Coiffait, 1979 , see: orbus Kiesenwetter, 1850 .

novus Last, 1981: 126 ( Neobisnius ; Type locality: Goroka, Kami River).

- DISTRIBUTION: New Guinea.

occidentoides Frank, 1981: 47 ( Neobisnius ; Type locality: USA: Texas, San Patricio Co.,

Lake Corpus Christi St. Pk. Also cited from Manitoba; Alberta; Vermont; New York; Ohio; Virginia; North Carolina; Kentucky; Tennessee; Missouri; Alabama; Mississippi; Illinois; Minnesota; Kansas; Oklahoma; Arkansas; Louisiana; North Dakota; Montana; Idaho; Washington; South Dakota; Nebraska; Wyoming; Colorado; Utah; Nevada; Texas; New Mexico; Arizona; California; Mexico).

— Downie and Arnett, 1996: 375 ( Neobisnius ; characters; USA).

— Schmidt, 1999: 108 ( Neobisnius ; food for adults and larvae used for rearing species; rearing methods).

- DISTRIBUTION: Canada, USA, Mexico.

ocreatus Horn, 1884 , see: sobrinus Erichson, 1840 .

oculatus Fauvel, 1905b: 178 ( Actobius ; Type locality: Diego-Suarez; forêts au nord de

Fort Dauphin. Afrique orientale anglaise: Baie de Kavirondo, Victoria Nyanza. Congo: Chiloango; Boma. Gabon: Loango).

— Fauvel, 1907: 37 ( Actobius ; Kavirondo Bay, Victoria-Nyanza; Congo; Gabon; Madagascar).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; catalog).

— Bernhauer and Chapman, 1939: 64 ( Neobisnius ; Kenya).

— Scheerpeltz, 1974: 134 ( Neobisnius ; South Africa; South West Africa).

— Lecoq, 1990: 203 ( Neobisnius ; Mascarene Islands).

- DISTRIBUTION: Mascarene Islands, Madagascar, Kenya, Congo, Gabon, Namibia, South Africa.

omnirufus Frank, 1981: 31 ( Neobisnius ; Type locality: Argentina: Mendoza).

- DISTRIBUTION: Argentina.

orbus Kiesenwetter, 1850: 220 ( Philonthus ; Type locality: Perpiniacum, Gerona).

— Kiesenwetter, 1851a: 418 ( Philonthus ; characters; France; Spain).

— Fairmaire and Laboulbène, 1856: 531 ( Philonthus ; characters; France).

— Kraatz, 1857d: 621 ( Philonthus ).

— Baudi, 1870: 387 ( Philonthus ; characters).

— Fauvel, 1874: 204 [= 1874a: 430] ( Erichsonius ; characters; France; Italy; Egypt; Madeira; Canary Islands).

— Fauvel, 1876a: 72 [= 1876: 257] ( Actobius ; Spain; Corsica; Syria).

— Mulsant and Rey, 1876: 592 [= 1877: 448] ( Bisnius ; characters; France).

— Fauvel, 1878: 121 [= 1878a: 41] ( Actobius ; Algeria; Egypt).

— Fauvel, 1886: 52 [= 1886a: 44] ( Actobius ; Morocco; Egypt; Algeria).

— Fauvel, 1902b: 107 ( Actobius ; Morocco; Algeria; Tunisia; Egypt; Canaries).

— Sainte-Claire Deville, 1906: 94 ( Neobisnius ; Corsica).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; catalog).

— Portevin, 1929: 356 ( Neobisnius ; characters; France).

— Peyerimhoff, 1931: 31 ( Neobisnius ; Algeria).

— Scheerpeltz, 1933: 1327 ( Neobisnius ; catalog).

— Gridelli, 1943: 125 ( Neobisnius ; characters; Egypt; Morocco; Italy).

— Last, 1948: 149 ( Neobisnius ; characters in key and aedeagal illustration).

— Cameron, 1952d: 824 ( Neobisnius ; Mauritania).

— Jarrige, 1958: 92 ( Neobisnius ; Algeria).

— Coiffait, 1966b: 338 ( Neobisnius ; Mauritania; Hoggar).

Smetana, 1967i: 551 ( Neobisnius ; Turkey) .

— Coiffait, 1968a: 136 ( Neobisnius ; Chad).

— Coiffait, 1974: 328 ( Neobisnius ; characters; Mediterranean region; Spain; North Africa; Italy; Anatolia; Egypt; Mauritania; Canary Islands; Pyrenees; Corsica).

— Maachi, 1991: 250 ( Neobisnius ; Morocco).

— Ciceroni and Zanetti, 1995: 27 ( Neobisnius ; Italy).

— Gusarov, 1997: 287 ( Neobisnius ; Saudi Arabia).

— Assing and Wunderle, 2001: 36 ( Neobisnius ; Cyprus).

- DISTRIBUTION: Afghanistan, Syria, Turkey, Italy, France, Spain, Madeira, Canary Islands, Cape Verde Islands, Mauritania, Morocco, Algeria, Tunisia, Egypt, Chad, Saudi Arabia, Madagascar, Réunion.

tenellus Wollaston, 1864: 576 ( Philonthus ; Type locality: Teneriffe: Barranco Santo near Santa Cruz).

— Wollaston, 1865: 495 ( Philonthus ; habitat; Canary Islands).

— Wollaston, 1867: 240 ( Philonthus ; characters; notes; Cape Verde Islands).

— Fauvel, 1874: 204 [= 1874a: 430] (Erichsoninus; synonym of orbus ).

— Fauvel, 1902b: 107 ( Actobius ; synonym of orbus ).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; synonym of orbus ).

— Gridelli, 1943: 125 ( Neobisnius ; synonym of orbus ).

— Coiffait, 1974: 329 ( Neobisnius ; synonym of orbus ).

putiphar Saulcy, 1864: 640 ( Philonthus ; Type locality: Kafr-Zaïat , au bord du Nil).

— Baudi, 1870: 387 ( Philonthus ; characters).

— Fauvel, 1874: 204 [= 1874a: 430] (Erichsoninus; synonym of orbus ).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; synonym of orbus ).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; synonym of orbus ).

— Gridelli, 1943: 125 ( Neobisnius ; synonym of orbus ).

— Coiffait, 1974: 329 ( Neobisnius ; synonym of orbus ).

nouristanicus Coiffait, 1979d: 559 ( Neobisnius ; subspecies of orbus ; Type locality: gorges de Chikari, entre Djelalabad et Kaboul).

sanguinicollis Lecoq, 1990: 205 ( Neobisnius ; subspecies of orbus ; Type locality: Madagascar Centre: Isalo. Also cited from Réunion Island).

orientalis Cameron, 1933d: 386 ( Neobisnius ; Type locality: Sumatra: Fort de Kock ).

- DISTRIBUTION: Indonesia.

paederoides LeConte, 1863a: 38 ( Philonthus ; [Note: Preoccupied secondary homonym in Philonthus ]; Type locality: “Central Valley”, Illinois, Missouri, etc., USA; [Note:

See lectotype designation by Frank, 1981]. Originally cited from Western States;

Lake Superior ; Fort Yuma and San Diego, California ).

— Fauvel, 1874: 201 [= 1874a: 427] ( Erichsonius ).

— Horn, 1884: 231 ( Actobius ; characters; Canada to Florida and westward to the Pacific coast).

— Fall and Cockerell, 1907: 166 ( Actobius ; New Mexico).

— Blatchley, 1910: 393 ( Actobius ; characters; Indiana).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; catalog).

— Notman, 1920: 17 ( Actobius ; New York).

— Notman, 1920a: 180 ( Actobius ; New York).

— Hatch, 1957: 175 ( Neobisnius ; characters; British Columbia; Washington; Idaho).

Frank, 1981: 17 ( Neobisnius ; lectotype designation: “ Central Valley ”, Illinois, Missouri , etc., USA; characters; New York; Maryland; Ohio; Wisconsin; Illinois; Kansas; North Carolina; South Carolina; Georgia; Florida ; Alabama; Mississippi; Texas) .

— Downie and Arnett, 1996: 374 ( Neobisnius ; characters; USA).

- DISTRIBUTION: Canada, USA.

palmula Gravenhorst, 1802 , see: villosulus Stephens, 1833 .

parcepunctatus Bernhauer, 1912e: 176 ( Neobisnius ; Type locality: Argentinien: Chaunar Region ; [Note: According to Frank, 1981: 12, 13 this locality is probably Chañar Region, La Rioja]).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; catalog).

— Frank, 1981: 12 ( Neobisnius ; characters; Argentina).

- DISTRIBUTION: Argentina.

piger Horn, 1884 , see: senilis Horn, 1884 .

planatus Gravenhorst, 1806 , see: procerulus Gravenhorst, 1806 .

politus Sharp, 1885 , see: nitidulus Sharp, 1885 .

praelongus Gemminger and Harold, 1868: 591 ( Philonthus ; replacement name for longulus Kraatz ).

— Fauvel, 1874: 201 [= 1874a: 427] ( Erichsonius ).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; catalog).

— Bernhauer, 1922a: 150 ( Neobisnius ; Cambodia).

Cameron, 1932: 56 ( Neobisnius ; characters; Bihar; Assam; India) .

— Scheerpeltz, 1933: 1327 ( Neobisnius ; catalog).

— Cameron, 1933d: 386 ( Neobisnius ; Sumatra).

— Cameron, 1937: 4 ( Neobisnius ; Java).

— Scheerpeltz, 1954: 113 ( Neobisnius ; India).

— Scheerpeltz, 1957c: 233 ( Neobisnius ; Sumba).

— Last, 1961: 306 ( Neobisnius ; Thailand).

— Scheerpeltz, 1965: 106 ( Neobisnius ; Burma).

— Watanabe and Shibata, 1976: 332 ( Neobisnius ; Malaysia).

— Coiffait, 1981f: 330 ( Neobisnius ; Andaman Islands).

— Barrion, Litsinger, and Last, 1991: 1078 ( Neobisnius ; associated with rice agroecosystem).

— Shibata, 1993a: 318 ( Neobisnius ; Taiwan).

— Ito, 1996a: 84 ( Neobisnius ; Japan; Thailand).

- DISTRIBUTION: Sri Lanka, India, Andaman Islands, Myanmar, Thailand, Cambodia, Vietnam, Malaysia, Indonesia, Philippines, Taiwan, Japan.

longulus Kraatz, 1859: 99 ( Philonthus ; [Note: When this name was replaced it was congeneric with Philonthus longulus Motschulsky which had been described in Gabrius and now is in Bisnius ]; Type locality: Ceylan; India orientalis).

— Fauvel, 1878f: 257 ( Actobius ).

— Fauvel, 1886b: 150 ( Actobius ; Philippines; Ceylon; East Indies; Annam; Sumatra; Celebes).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; synonym of praelongus ).

— Cameron, 1932: 57 ( Neobisnius ; synonym of praelongus ).

— Cameron, 1933d: 386 ( Neobisnius ; synonym of praelongus ).

procerulus Gravenhorst, 1806: 95 ( Staphylinus ; Type locality: Brunsvigae).

— Mannerheim, 1830: 34 ( Gyrohypnus ; Finland; Sweden).

— Mannerheim, 1831: 448 ( Gyrohypnus ; Finland; Sweden).

— Lacordaire, 1835: 409 ( Staphylinus ; characters; France).

— Nordmann, 1837: 120 ( Gyrohypnus ; characters; Germany; Finland).

— Erichson, 1839a: 479 ( Philonthus ; characters; Germany).

— Heer, 1839: 272 ( Philonthus ; characters; Switzerland).

— Erichson, 1840: 541 ( Philonthus ; characters; Germany).

— Kolenati, 1846: 15 ( Leptacinus ; Elisabethopol).

— Hochhuth, 1849: 146 ( Philonthus ; characters).

— Luff, 1996: 30 ( Neobisnius ; collecting notes; Isle of Man).

- DISTRIBUTION: Madagascar, Congo, Kenya, Ethiopia, Egypt, Tunisia, Algeria, Morocco, Canary Islands, Madeira, Azores, Europe, Russia, Azerbaijan, Turkey, Syria, Afghanistan, Uzbekistan, Australia, USA (?).

planatus Gravenhorst, 1806: 108 ( Staphylinus ; Type locality: Borussia).

— Mannerheim, 1830: 34 ( Gyrohypnus ; Russia).

— Mannerheim, 1831: 448 ( Gyrohypnus ; Russia).

— Erichson, 1840: 511 ( Philonthus ; characters; Germany).

— Kraatz, 1857d: 624 ( Philonthus ; synonym of procerulus ).

— Bernhauer and Schubert, 1914: 324 ( Neobisnius ; synonym of procerulus ).

— Smetana, 1958: 137 ( Neobisnius ; synonym of procerulus ).

cinerascens Stephens, 1833: 252 ( Gabrius ; Type locality: Devonshire).

— Stephens, 1839: 401 ( Gabrius ; characters; England).

— Gemminger and Harold, 1868: 591 ( Philonthus ; synonym of procerulus ).

— Bernhauer and Schubert, 1914: 323 ( Neobisnius ; synonym of procerulus ).

— Smetana, 1958: 137 ( Neobisnius ; synonym of procerulus ).

filiformis Wollaston, 1854: 585 ( Philonthus ; Type locality: Madeira: Santa Anna ).

— Wollaston, 1857: 192 ( Philonthus ; habitat; Madeira Islands).

— Wollaston, 1865: 496 ( Philonthus ; habitat; notes; Madeira Islands).

— Wollaston, 1871: 304 ( Philonthus ; notes; characters; Madeira).

— Fauvel, 1874: 201 [= 1874a: 427] ( Erichsonius ).

— Fauvel, 1902b: 107 ( Actobius ; synonym of procerulus ).

— Bernhauer and Schubert, 1914: 324 ( Neobisnius ; variety of procerulus ).

— Smetana, 1958: 137 ( Neobisnius ; synonym of procerulus ).

xantholinoides Wollaston, 1864: 577 ( Philonthus ; Type locality: Teneriffe : Santa Cruz).

— Wollaston, 1865: 496 ( Philonthus ; habitat; Canary Islands).

— Fauvel, 1874: 206 [= 1874a: 432] (Erichsoninus; synonym of prolixus ).

— Fauvel, 1902b: 107 ( Actobius ; synonym of procerulus ).

— Bernhauer and Schubert, 1914: 324 ( Neobisnius ; synonym of procerulus ).

— Smetana, 1958: 137 ( Neobisnius ; synonym of procerulus ).

decolor Coiffait, 1974: 325 , 330 ( Neobisnius ; subspecies of procerulus ; Type locality:

Nord Afghanistan: Bala Murghab, 470 m ; [Note: This subspecies and Neobisnius procerulus margianus have the same type locality]).

margianus Bordoni, 1975a: 151 ( Neobisnius ; subspecies of procerulus ; Type locality: N-Afghanistan: Prov. Herat: Bala Murghab, 470 m).

prolixus Erichson, 1840: 510 ( Philonthus ; Type locality: Austria).

— Redtenbacher, 1849: 707 ( Philonthus ; characters; Austria).

— Kiesenwetter, 1851a: 418 ( Philonthus ; France; Spain; Dalmatia).

— Fairmaire and Laboulbène, 1856: 530 ( Philonthus ; characters; France).

— Kraatz, 1857d: 625 ( Philonthus ; characters; Germany).

— Redtenbacher, 1857: 198 ( Philonthus ; characters; Austria).

— Hochhuth, 1862: 65 ( Philonthus ; Russia).

— Fauvel, 1865a: 297 [= 1865b: 47] ( Philonthus ; habitat; France).

— Fauvel, 1874: 206 [= 1874a: 432] ( Erichsonius ; characters; France; Britain; Algeria; Caucasus; Iran; Canary Islands).

— Redtenbacher, 1874: 216 ( Philonthus ; characters; Austria).

— Mulsant and Rey, 1876: 589 [= 1877: 445] ( Bisnius ; characters; France).

— Fauvel, 1878: 121 [= 1878a: 41] ( Actobius ; Algeria).

— Fauvel, 1886: 53 [= 1886a: 45] ( Actobius ; Algeria).

— Fowler, 1888: 286 ( Actobius ; characters; Britain).

— Ganglbauer, 1895: 465 ( Neobisnius ; characters; middle Europe; Mediterranean region).

— Fauvel, 1902b: 107 ( Actobius ; synonym of procerulus ).

— Reitter, 1909: 123 ( Neobisnius ; characters; Germany).

— Bernhauer and Schubert, 1914: 324 ( Neobisnius ; catalog).

— Watanabe, 1961: 353 ( Leptacinus ; Izu Islands).

— Nakane, 1963a: 90 ( Neobisnius ; characters; habitus photograph; Japan).

— Cho, 1996a: 162 ( Neobisnius ; characters; Korea).

- DISTRIBUTION: Taiwan, Japan, Korea.

putiphar Saulcy, 1864 , see: orbus Kiesenwetter, 1850 .

richteri Bernhauer, 1927e: 245 ( Neobisnius ; Type locality: Argentinien: Prov. Buenos Aires).

— Scheerpeltz, 1933: 1327 ( Neobisnius ; catalog).

— Frank, 1981: 35 ( Neobisnius ; characters; Argentina; Brazil).

- DISTRIBUTION: Argentina, Brazil.

rubripennis Gridelli, 1943 , see: lathrobioides Baudi, 1848 .

rufipennis Cameron, 1919a: 252 ( Neobisnius ; Type locality: Ceylon: Nuwera Eliya ,

alt. 6230–8000 feet).

— Cameron, 1932: 58 ( Neobisnius ; characters; Ceylon).

— Scheerpeltz, 1933: 1327 ( Neobisnius ; catalog).

— Scheerpeltz, 1954: 113 ( Neobisnius ; Ceylon).

- DISTRIBUTION: Sri Lanka.

sanguinicollis Lecoq, 1990 , see: orbus Kiesenwetter, 1850 .

sanguinipennis Scheerpeltz, 1972: 110 ( Neobisnius ; Type locality: Argentinien: Rio Negro ,

El Bolson, 350 m).

- DISTRIBUTION: Argentina.

scutellaris Bernhauer, 1908d: 336 ( Neobisnius ; Type locality: Paraguay).

— Bernhauer and Schubert, 1914: 324 ( Neobisnius ; catalog).

— Scheerpeltz, 1933: 1328 ( Neobisnius ; catalog; Argentina).

— Scheerpeltz, 1972: 17 ( Neobisnius ; Argentina).

— Frank, 1981: 30 ( Neobisnius ; characters; Bolivia; Brazil; Paraguay; Argentina; Uruguay).

- DISTRIBUTION: Brazil, Bolivia, Paraguay, Uruguay, Argentina.

semipunctatus Fairmaire and Germain, 1861: 434 ( Othius ; Type locality: Santiago).

— Fauvel, 1866a: 346 [= 1866b: 97] ( Philonthus ; characters; Chile).

— Fauvel, 1874: 205 [= 1874a: 431] ( Erichsonius ; characters; France; Spain; Algeria; Chile; Australia [Note: This species only occurs in Chile according to Fagel, 1954, so all but Chile

of the preceding citations are erroneous]).

— Fauvel, 1876a: 72 [= 1876: 257] ( Actobius ; Ethiopia [Note: This species only occurs in Chile according to Fagel, 1954, so the preceeding citation is erroneous]).

— Mulsant and Rey, 1876: 842 [= 1877: 698] ( Bisnius ; characters; France [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).

— J. Sahlberg, 1876: 23 ( Bisnius ; characters; Finland [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).

— Fauvel, 1878: 121 [= 1878a: 41] ( Actobius ; Algeria [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).

— Fauvel, 1878g: 545 ( Actobius ; characters; Australia [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).

— Horn, 1884: 227 ( Actobius ; characters; Colorado; Nevada [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).

— Fauvel, 1886: 53 [= 1886a: 45] ( Actobius ; Algeria [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).

— Ganglbauer, 1895: 465 ( Neobisnius ; synonym of procerulus ).

— Blackburn, 1896: 262 ( Actobius ; Australia [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).

— Fauvel, 1902b: 107 ( Actobius ; synonym of procerulus ).

— Bernhauer and Schubert, 1914: 324 ( Neobisnius ; variety of procerulus ).

— Portevin, 1929: 356 ( Neobisnius ; characters; France [Note: This species only occurs in Chile according to Fagel, 1954, so the citation is erroneous]).

— Scheerpeltz, 1933: 1327 ( Neobisnius ; variety of procerulus ).

— Gridelli, 1943: 127 ( Neobisnius ; characters). — Fagel, 1954 ( Neobisnius ; valid species; characters; separated from procerulus ; all distributional records outside of Chile probably represent different species). — Hatch, 1957: 175 ( Neobisnius ; variety of procerulus ; characters; British Columbia; Washington; Oregon; [Note: This species only occurs in Chile according to Fagel, 1954, so the citation by Hatch is erroneous]). — Coiffait and Saiz, 1968: 361 ( Neobisnius ; characters; Chile). — Saiz, 1971b: 349 ( Neobisnius ; Chile). — Frank, 1981: 10 ( Neobisnius ; characters; Chile). - DISTRIBUTION: Chile.

semirufus Bernhauer, 1921c: 105 ( Neobisnius ; Type locality: Argentinien: Prov. Salta). — Scheerpeltz, 1933: 1328 ( Neobisnius ; catalog). — Frank, 1981: 33 ( Neobisnius ; characters; Argentina; Bolivia). - DISTRIBUTION: Argentina, Bolivia.

senilis Horn, 1884: 226 ( Actobius ; Type locality: Garland, Costilla Co., Colorado; [Note: See lectotype designation by Frank, 1981]. Originally cited from Colorado; Nevada; California). — Fall and Cockerell, 1907: 166 ( Actobius ; New Mexico). — Bernhauer and Schubert, 1914: 325 ( Actobius ; catalog). — Hatch, 1957: 175 ( Neobisnius ; characters; British Columbia; Washington; Oregon; Idaho). — Frank, 1981: 45 ( Neobisnius ; lectotype designation: Garland, Costilla Co., Colorado; characters; British Columbia; Washington; Idaho; Oregon; Colorado; Nevada; New Mexico; California; Mexico). - DISTRIBUTION: Canada, USA, Mexico.

piger Horn, 1884: 219 ( Philonthus ; Type locality: Not cited). — Horn, 1884: 240 ( Actobius ; synonym of senilis ).

simplex Sharp, 1885: 459 ( Actobius ; Type locality: Mexico, Mirador). — Bernhauer and Schubert, 1914: 324 ( Neobisnius ; catalog). — Frank, 1981: 42 ( Neobisnius ; characters; Mexico; Guatemala; El Salvador; Costa Rica; Panama; Brazil). - DISTRIBUTION: Brazil, Panama, Costa Rica, El Salvador, Guatemala, Mexico.

cavifrons Bierig, 1933: 51 ( Neobisnius ; Type locality: Panama: Puerto Armuelles. Also listed from Costa Rica). — Frank, 1981: 42 ( Neobisnius ; synonym of simplex ).

simulator Smetana, 1963 , see: sobrinus Erichson, 1840 .

sobrinus Erichson, 1840: 512 ( Philonthus ; Type locality: Pensylvania; [Note: See lectotype designation by Smetana, 1963]). — Fauvel, 1874: 201 [= 1874a: 427] ( Erichsonius ). — Horn, 1884: 228 ( Actobius ; characters; east of Rocky Mountains; California). — Bernhauer and Schubert, 1914: 324 ( Neobisnius ; catalog). — Notman, 1920: 17 ( Actobius ; New York). — Notman, 1920a: 180 ( Actobius ; New York). — Smetana, 1963: 4, 5 ( Neobisnius ; characters; lectotype designation; Pennsylvania). — Smetana, 1965: 10 ( Neobisnius ; Nova Scotia; Newfoundland). — Frank, 1981: 39 ( Neobisnius ; characters; Newfoundland; New Brunswick; Quebec; Ontario; Maine; New Hampshire; Vermont; Massachusetts; Connecticut; New York; Pennsylvania; Illinois; Wisconsin; Iowa; Nebraska; Oregon; New Jersey; Kentucky; Missouri; Kansas; Colorado; Utah; Maryland; Virginia; West Virginia; North Carolina; Tennessee; South Carolina; Georgia; Florida; Louisiana; Texas; New Mexico; Arizona; California; Mexico; Guatemala; Costa Rica; Venezuela; Colombia; Brazil). — Schmidt, 1994 ( Neobisnius ; egg, larval, and pupal characters; biological notes; Nebraska). — Downie and Arnett, 1996: 375 ( Neobisnius ; characters; USA). — Schmidt, 1999: 108 ( Neobisnius ; food for adults and larvae used for rearing species; rearing methods). - DISTRIBUTION: Canada, USA, Mexico, Guatemala, Costa Rica, Colombia, Venezuela,

Brazil.

agnatus Erichson, 1840: 513 ( Philonthus ; Type locality: valle Araguensi Columbiae; [Note: See lectotype designation by Smetana, 1963]).

— Solsky, 1868a: 137 ( Philonthus ; Mexico).

— Fauvel, 1874: 201 [= 1874a: 427] ( Erichsonius ).

— Horn, 1884: 229 ( Actobius ; characters; Louisiana).

— Sharp, 1885: 459 ( Actobius ; notes; Mexico; Louisiana; Colombia).

— Bernhauer and Schubert, 1914: 322 ( Neobisnius ; catalog).

— Smetana, 1963: 4 ( Neobisnius ; characters; lectotype designation).

— Frank, 1981: 39 ( Neobisnius ; synonym of sobrinus ).

ocreatus Horn, 1884: 228 ( Actobius ; Type locality: southern California; [Note: See lectotype designation by Frank, 1981]. Originally cited from California from the middle regions southward).

— Bernhauer and Schubert, 1914: 325 ( Actobius ; catalog).

— Frank, 1981: 39 ( Neobisnius ; lectotype designation: southern California; synonym of sobrinus ).

arduus Sharp, 1885: 458 ( Actobius ; Type locality: Puebla, Mexico; [Note: See lectotype designation by Frank , 1981]. Originally cited from Mexico, Guanajuato, Cordova , Puebla, Mirador , Tehuacan , Jalapa; Guatemala, near the city, San Geronimo , Torola , Senahu , Balheu , Tactic , Panajachel 5000 feet, Cerro Zunil 4000 feet; Panama, San Miguel in the Pearl Islands).

— Bernhauer and Schubert, 1914: 322 ( Neobisnius ; catalog).

— Frank, 1981: 39 ( Neobisnius ; lectotype designation: Puebla, Mexico; synonym of sobrinus ).

simulator Smetana, 1963: 3 ( Neobisnius ; Type locality: West-Newfoundland: Curling).

— Smetana, 1965: 11 ( Neobisnius ; Newfoundland).

— Frank, 1981: 39 ( Neobisnius ; synonym of sobrinus ).

subaeneus Bernhauer, 1928: 15 ( Neobisnius ; Type locality: Sumatra: Medan ).

— Scheerpeltz, 1933: 1328 ( Neobisnius ; catalog).

— Cameron, 1933d: 386 ( Neobisnius ; Sumatra).

- DISTRIBUTION: Indonesia.

subprolixus Coiffait, 1982: 68 ( Neobisnius ; Type locality: Nepal: Ufer des Tila Khola ,

près Jumla).

— Coiffait, 1981e: 349 ( Neobisnius ; Baltistan).

- DISTRIBUTION: Nepal.

sumbaensis Scheerpeltz, 1957c: 233 , 316 ( Neobisnius ; Type locality: Sumba: Baing).

- DISTRIBUTION: Indonesia.

tenellus Wollaston, 1864 , see: orbus Kiesenwetter, 1850 .

terminalis LeConte, 1863a: 38 ( Philonthus ; Type locality: Maryland; [Note: See lectotype designation by Frank, 1981]).

— Fauvel, 1874: 201 [= 1874a: 427] ( Erichsonius ).

— Horn, 1884: 232 ( Actobius ; characters; Canada to middle States; Texas).

— Bernhauer and Schubert, 1914: 324 ( Neobisnius ; catalog).

— Notman, 1919: 136 ( Actobius ; New York).

— Notman, 1920: 17 ( Actobius ; New York).

— Notman, 1920a: 180 ( Actobius ; New York).

— Bierig, 1933: 56 ( Neobisnius ).

— Smetana, 1965: 11 ( Neobisnius ; Nova Scotia).

Frank, 1981: 13 ( Neobisnius ; lectotype designation; characters; Nova Scotia ; Quebec ; Maine ; New Hampshire ; New York ; Pennsylvania ; Michigan ; Iowa ; Colorado ; Maryland ; Virginia ; Texas ; New Mexico ; Arizona ; California ; Baja California ; Tamaulipas ; Guerrero ; Sinaloa ; Puebla; Oaxaca; Guatemala; Costa Rica) .

— Dearborn and Donahue, 1994: 14 ( Neobisnius ; Maine).

— Downie and Arnett, 1996: 374 ( Neobisnius ; characters; USA).

- DISTRIBUTION: Canada, USA, Mexico, Guatemala, Costa Rica, Panama.

elegantulus Horn, 1884: 232 ( Actobius ; Type locality: Arizona; [Note: See lectotype

designation by Frank, 1981]. Also cited from California).

— Bernhauer and Schubert, 1914: 322 ( Neobisnius ; catalog).

— Frank, 1981: 13 ( Neobisnius ; lectotype designation: Arizona; synonym of terminalis ).

delicatulus Sharp, 1885: 461 ( Actobius ; Type locality: Guatemala, Pantaleon 1700 feet, Paso Antonio 400 feet).

— Bernhauer and Schubert, 1914: 322 ( Neobisnius ; catalog).

— Frank, 1981: 13 ( Neobisnius ; synonym of terminalis ).

adustus Sharp, 1887: 790 ( Actobius ; Type locality: Mexico, La Noria in Sinaloa, Tehuantepec).

— Bernhauer and Schubert, 1914: 322 ( Neobisnius ; catalog).

— Bierig, 1933: 48 ( Neobisnius ; characters; Panama).

— Frank, 1981: 13 ( Neobisnius ; synonym of terminalis ).

transvaalensis Scheerpeltz, 1974: 134 ( Neobisnius ; Type locality: Transvaal: Kruger National Park, Letaba Camp).

- DISTRIBUTION: South Africa.

umbripennis LeConte, 1863 , see: ludicrus Erichson, 1840 .

vigii Frank, 1981: 43 ( Neobisnius ; Type locality: Colombia: Magdalena, Parque Tayrona ,

21 mi. E of Santa Marta).

- DISTRIBUTION: Colombia.

villosulus Stephens, 1833: 251 ( Gabrius ; Type locality: Suffolk).

— Stephens, 1839: 401 ( Gabrius ; characters; England).

— Fauvel, 1874: 204 [= 1874a: 430] ( Erichsonius ; characters; France; Caucasus).

— J. Sahlberg, 1876: 22 ( Bisnius ; Finland).

— Fowler, 1888: 285 ( Actobius ; characters; Britain).

— Ganglbauer, 1895: 464 ( Neobisnius ; characters; Europe).

— Reitter, 1909: 123 ( Neobisnius ; characters; Germany).

— Bernhauer and Schubert, 1914: 324 ( Neobisnius ; catalog).

— Johansen, 1914: 424 ( Bisnius ; characters; Denmark).

— Portevin, 1929: 356 ( Neobisnius ; characters; France).

— Scheerpeltz, 1933: 1328 ( Neobisnius ; catalog).

— Tottenham, 1949: 417 ( Neobisnius ; checklist; Britain).

— V. Hansen, 1952: 30 ( Neobisnius ; characters; Denmark).

— Blackwelder, 1952: 258 ( Neobisnius ; type species of genus).

— Smetana, 1958: 135 ( Neobisnius ; characters; ecology; Czechoslovakia).

— Palm, 1963b: 58 ( Neobisnius ; characters; Sweden; Denmark; Norway; Finland).

— Lohse, 1964: 171 ( Neobisnius ; characters; central Europe).

— Horion, 1965: 114 ( Neobisnius ; Russia; Finland; Sweden; Norway; Britain; Denmark; Holland; Belgium; France to Poland; Romania; Italy; Slovenia; Dalmatia; Bosnia; Germany; Austria).

— Szujecki, 1968a: 726 ( Neobisnius ; cited as villosus; Poland).

— Tikhomirova, 1973a: 182 ( Neobisnius ; checklist; Soviet Union).

— Coiffait, 1974: 327 ( Neobisnius ; characters; Europe).

— Moore and Legner, 1975: 18 ( Neobisnius ; type species of genus).

— Pope, 1977: 29 ( Neobisnius ; Britain).

— Muona, 1979: 14 ( Neobisnius ; Finland; Russia; Sweden; Norway; Denmark).

— Moore and Legner, 1979: 25 ( Neobisnius ; type species of genus).

— Burakowski, Mroczkowski, and Stefańska, 1980: 31 ( Neobisnius ; catalog; Poland).

— Frank, 1981: 8 ( Neobisnius ; characters; Quebec; Ontario; New York; Minnesota; Oregon).

— Uhlig and Vogler, 1981: 110 ( Neobisnius ; notes; Germany).

— Tóth, 1984: 35 ( Neobisnius ; characters; Hungary).

— Lucht, 1987: 104 ( Neobisnius ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Gusarov, 1989: 10 ( Neobisnius ; habitat notes; Ukraine).

— Lohse and Lucht, 1989: 160 ( Neobisnius ; notes).

— Zanetti, 1989: 112 ( Neobisnius ; Italy).

— Silfverberg, 1992: 17 ( Neobisnius ; Finland; Russia; Sweden; Norway; Denmark; Estonia;

Lithuania).

— Smetana, 1993b: 48 ( Neobisnius ; Czech Republic; Slovakia).

— Ciceroni and Zanetti, 1995: 27 ( Neobisnius ; Italy).

— Downie and Arnett, 1996: 375 ( Neobisnius ; characters; USA).

— M. Hansen, 1996: 104 ( Neobisnius ; checklist; Denmark).

— Anderson, Nash, and O’Connor, 1997: 18 ( Neobisnius ; Ireland).

— Herman, 2001: 26 ( Neobisnius ; nomen protectum).

- DISTRIBUTION: Europe, Canada, USA.

palmula Gravenhorst, 1802: 49 ( Staphylinus ; [Note: This name is older than villosulus ]; Type locality: Not cited, but presumably Brunsvigae).

— Latreille, 1804: 333 ( Staphylinus ; synonym of cinctus Latreille ).

— Gravenhorst, 1806: 107 ( Staphylinus ).

— Gyllenhal, 1810: 359 ( Staphylinus ; synonym of pilicornis Paykull ).

— Zetterstedt, 1828: 81 ( Xantholinus ; synonym of pilicornis Paykull ).

— Erichson, 1840: 514 ( Philonthus ; characters; Germany).

— Redtenbacher, 1849: 825 ( Philonthus ; characters; Austria).

— Kraatz, 1857d: 625 ( Philonthus ; notes).

— Bernhauer and Schubert, 1914: 324 ( Neobisnius ; synonym of villosulus ).

— Scheerpeltz, 1933: 1328 ( Neobisnius ; synonym of villosulus ).

— Gridelli, 1943: 118 ( Neobisnius ; synonym of elongatulus ).

— Smetana, 1958: 135 ( Neobisnius ; synonym of villosulus ).

— Coiffait, 1974: 327 ( Neobisnius ; synonym of villosulus ).

— Herman, 2001: 26 ( Neobisnius ; nomen oblitum).

cinctus Latreille, 1804: 333 ( Staphylinus ; [preoccupied]; Type locality: Brunswick).

— Erichson, 1840: 514 ( Philonthus ; synonym of palmula ).

— Bernhauer and Schubert, 1914: 324 ( Neobisnius ; synonym of villosulus ).

— Smetana, 1958: 135 ( Neobisnius ; synonym of villosulus ).

elongatulus Erichson, 1839a: 479 ( Philonthus ; Type locality: Mark Brandenburg).

— Heer, 1839: 272 ( Philonthus ; characters; Switzerland).

— Erichson, 1840: 511 ( Philonthus ; characters; Germany).

— Jacobsen, 1849: 706 ( Philonthus ; characters; Denmark).

— Hochhuth, 1849: 146 ( Philonthus ; characters; Mingrelien).

— Redtenbacher, 1849: 707 ( Philonthus ; characters; Austria).

— Fairmaire and Laboulbène, 1856: 531 ( Philonthus ; characters; France).

— Kraatz, 1857d: 623 ( Philonthus ; characters; Germany).

— Redtenbacher, 1857: 197 ( Philonthus ; characters; Austria).

— Thomson, 1859: 25 (type species of Bisnius ).

— Thomson, 1860: 168 ( Bisnius ; characters; Scandinavia).

— Fauvel, 1865a: 297 [= 1865b: 47] ( Philonthus ; habitat; France).

— Fauvel, 1874: 204 [= 1874a: 430] ( Erichsonius ; synonym of villosulus ).

— Redtenbacher, 1874: 216 ( Philonthus ; characters; Austria).

— Seidlitz, 1875: 279 ( Philonthus ; characters; Baltic region).

— Mulsant and Rey, 1876: 594 [= 1877: 450] ( Bisnius ; characters; France).

— Ganglbauer, 1895: 464 ( Neobisnius ; synonym of villosulus ).

— Bernhauer and Schubert, 1914: 324 ( Neobisnius ; synonym of villosulus ).

— Scheerpeltz, 1933: 1328 ( Neobisnius ; synonym of villosulus ).

— Gridelli, 1943: 118 ( Neobisnius ; characters; Italy; Balkan peninsula).

— Last, 1948: 149 ( Neobisnius ; characters in key and aedeagal illustration).

— Smetana, 1958: 135 ( Neobisnius ; synonym of villosulus ).

— Coiffait, 1974: 327 ( Neobisnius ; synonym of villosulus ).

— Lohse and Lucht, 1989: 160 ( Neobisnius ; synonym of villosulus ).

fulvicornis Notman, 1920: 27 ( Actobius ; Type locality: Schoharie, N.Y.).

— Scheerpeltz, 1933: 1329 ( Neobisnius ; catalog).

— Frank, 1981: 8 ( Neobisnius ; synonym of villosulus ).

xantholinoides Wollaston, 1864 , see: procerulus Gravenhorst, 1806 .

Orthidus

[1 species; Palaearctic Region]

Orthidus Mulsant and Rey, 1876: 339 [= 1876: 195] (species included: cribratus ). TYPE SPECIES: Philonthus cribratus Erichson , fixed by monotypy.

— Heyden, Reitter, and Weise, 1891: 102 (list of species of Europe and Caucasus).

— Ganglbauer, 1895: 436 (characters).

— Heyden, Reitter, and Weise, 1906: 168 (list of species of Europe and Caucasus).

— Reitter, 1909: 116, 122 (characters).

— Bernhauer and Schubert, 1914: 361 (world catalog; 1 species).

— Cameron, 1919b: 207 (synonym of Phucobius ).

— Cameron, 1921a: 354, 405 (characters; key to Singapore species; catalog).

— Winkler, 1925: 381 (catalog for Palaearctic region).

— Porta, 1926: 99 (characters; species of Italy).

— Portevin, 1929: 354 ( Orthidus ; characters; France).

— Scheerpeltz, 1933: 1368 (world catalog supplement).

— Normand, 1935: 372 (list of species of Tunisia).

— Blackwelder, 1952: 278 (type species: cribratus ).

— Coiffait, 1956e: 183, 187, 220 (characters; notes).

— Kocher, 1958: 152 (checklist of species; Morocco).

— Smetana, 1959a: 214 (list of species collected in Albania).

— Coiffait, 1963: 8, 20 (characters).

— Tikhomirova, 1973a: 186 (checklist of species of USSR).

— Coiffait, 1974: 350 (characters; species of western Palaearctic region).

— Moore and Legner, 1976: 541 (includes seashore species; notes).

— Smetana, 1995e: 49 (characters).

— Ciceroni and Zanetti, 1995: 27 (list of species of Italy).

atlanticus Coiffait, 1956 , see: cribratus Erichson, 1840 .

cribratus Erichson, 1840: 431 ( Philonthus ; Type locality: Istria; Gallia meridionali; Sardinia; Lusitania ).

— Küster, 1853a: 45 ( Philonthus ; characters; France; Spain; Portugal; Sardinia; Istria; Dalmatia). — Fairmaire and Laboulbène, 1856: 514 ( Philonthus ; characters; France).

— Kraatz, 1857d: 569 ( Philonthus ; characters).

— Redtenbacher, 1857: 189 ( Philonthus ; characters; Austria).

— Fauvel, 1874: 196 [= 1874a: 422] ( Cafius ; characters; France; Austria; Italy; Corsica; Sardinia; Portugal; Algeria).

— Redtenbacher, 1874: 208 ( Philonthus ; characters; Austria).

— Mulsant and Rey, 1876: 341 [= 1877: 197] ( Anodus ; characters; France).

— Fauvel, 1878: 120 [= 1878a: 40] ( Cafius ; Algeria; Tunisia).

— Fauvel, 1886: 52 [= 1886a: 44] ( Cafius ; Morocco; Tunisia; Algeria).

— Ganglbauer, 1895: 436 ( Orthidus ; characters; coast of western Mediterranean region;

Atlantic coast of France).

— Fauvel, 1902b: 106 ( Cafius ; Morocco; Algeria; Tunisia).

— Sainte-Claire Deville, 1906: 95 ( Orthidus ; Corsica).

— Reitter, 1909: 122 ( Orthidus ).

— Bernhauer and Schubert, 1914: 361 ( Orthidus ; catalog).

— Scheerpeltz, 1933: 1368 ( Orthidus ; catalog).

— Coiffait, 1956e: 221 ( Orthidus ; characters).

— Focarile, 1964: 58 ( Orthidus ; Italy).

— Coiffait, 1974: 352 ( Orthidus ; characters; Mediterranean coast of Europe and North Africa).

— Bordoni and Carfi, 1978 ( Orthidus ; note on peg setae of aedeagus).

— Ciceroni and Zanetti, 1995: 27 ( Orthidus ; Italy).

- DISTRIBUTION: Tunisia, Algeria, Morocco, Portugal, Spain, France, Italy, Croatia. atlanticus Coiffait, 1956e: 221 ( Orthidus ; subspecies of cribratus ; Type locality: côtes atlantiques de la Bretagne aux Pyrénées et probablement en Espagne).

— Coiffait, 1974: 352 ( Orthidus ; subspecies of cribratus ; characters; Atlantic coast of Morocco).

angularis Erichson, 1840: 477 ( Philonthus ; Type locality: Mexico).

— Sharp, 1885: 443 ( Paederomimus ; Mexico).

— Bernhauer and Schubert, 1914: 367 ( Paederomimus ; catalog).

— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 62 ( Paederomimus ; necrophilous species; distribution in Mexico).

- DISTRIBUTION: Mexico.

angusticollis Erichson, 1840: 487 ( Philonthus ; Type locality: Mexico).

— Sharp, 1885: 447 ( Paederomimus ; notes; Mexico; Guatemala).

— Bernhauer and Schubert, 1914: 367 ( Paederomimus ; catalog).

- DISTRIBUTION: Mexico, Guatemala.

aterrimus Bernhauer, 1917d: 36 ( Paederomimus ; Type locality: West-Columbien: Umgebung von Cali am Rio Cauca, Cañon del Mte. Tolima, 1700 m).

— Scheerpeltz, 1933: 1376 ( Paederomimus ; catalog).

- DISTRIBUTION: Colombia.

basipennis Sharp, 1885: 450 ( Paederomimus ; Type locality: Panama, Chiriqui).

— Bernhauer and Schubert, 1914: 367 ( Paederomimus ; catalog).

- DISTRIBUTION: Panama.

capitalis Sharp, 1876: 171 ( Philonthus ; Type locality: Ega).

— Sharp, 1885: 438 ( Paederomimus ).

— Bernhauer and Schubert, 1914: 367 ( Paederomimus ; catalog).

- DISTRIBUTION: Brazil.

capitatus Sharp, 1885: 441 ( Paederomimus ; Type locality: Mexico, Jalapa , Playa Vicente).

— Bernhauer and Schubert, 1914: 367 ( Paederomimus ; catalog).

- DISTRIBUTION: Mexico.

cephalax Sharp, 1885: 446 ( Paederomimus ; Type locality: Panama, David ).

— Bernhauer and Schubert, 1914: 367 ( Paederomimus ; catalog).

- DISTRIBUTION: Panama.

chontalensis Sharp, 1885: 440 ( Paederomimus ; Type locality: Nicaragua, Chontales).

— Bernhauer and Schubert, 1914: 367 ( Paederomimus ; catalog).

- DISTRIBUTION: Nicaragua.

cognatus Sharp, 1876: 169 ( Philonthus ; [preoccupied]; Type locality: Ega).

— Sharp, 1885: 438 ( Paederomimus ).

— Bernhauer and Schubert, 1914: 367 ( Paederomimus ; catalog).

— Herman, 2001: 46 (Paederominus; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: Brazil.

confertus Sharp, 1885: 449 ( Paederomimus ; Type locality: Panama, David, Volcan de Chiriqui about 4000 feet).

— Bernhauer and Schubert, 1914: 367 ( Paederomimus ; catalog).

- DISTRIBUTION: Panama.

conformis Sharp, 1876: 175 ( Philonthus ; [Note: Preoccupied as secondary homonym in Philonthus ]; Type locality: Amazons).

— Sharp, 1885: 438 ( Paederomimus ).

— Bernhauer and Schubert, 1914: 367 ( Paederomimus ; catalog).

- DISTRIBUTION: Brazil.

contractus Sharp, 1885: 448 ( Paederomimus ; Type locality: Guatemala, San Geronimo , Cubilguitz ; Cahabon, Chacoj; Nicaragua).

— Bernhauer and Schubert, 1914: 367 ( Paederomimus ; catalog).

- DISTRIBUTION: Guatemala, Nicaragua.

corrosus Sharp, 1885: 451 ( Paederomimus ; Type locality: Panama, Bugaba 800 to 1500 feet). — Bernhauer and Schubert, 1914: 367 ( Paederomimus ; catalog).

- DISTRIBUTION: Panama.

cribratus Sharp, 1885: 450 ( Paederomimus ; Type locality: Panama, Volcan de Chiriqui

2000 to 3000 feet).

— Bernhauer and Schubert, 1914: 367 ( Paederomimus ; catalog).

- DISTRIBUTION: Panama.

cribricollis Erichson, 1840: 487 ( Philonthus ; Type locality: Columbia ; Mexico).

— Solsky, 1875: 15 ( Philonthus ; Peru).

— Sharp, 1885: 448 ( Paederomimus ; notes; Mexico; Colombia).

— Fauvel, 1891: 118 ( Paederomimus ; Venezuela).

— Fauvel, 1901: 87 ( Paederomimus ; Colombia; Venezuela; Antilles).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

— Blackwelder, 1943: 396 ( Paederomimus ; characters; Grenada; Trinidad).

— Marcuzzi, 1977: 12 ( Paederomimus , cited as Paederimimus; Venezuela).

- DISTRIBUTION: West Indies, Mexico, Venezuela, Colombia, Peru.

trinidadensis Bernhauer, 1915 l: 296 ( Paederomimus ; Type locality: Trinidad: Caparo ).

— Scheerpeltz, 1933: 1376 ( Paederomimus ; catalog).

— Blackwelder, 1943: 397 ( Paederomimus ; synonym of cribricollis ).

cupreonitens Schubert, 1909: 294 ( Paederomimus ; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d’Orizaba).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Mexico.

densiceps Bernhauer, 1908d: 332 ( Paederomimus ; Type locality: Brasilien: Petropolis ).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Brazil.

difformiceps Sharp, 1885: 445 ( Paederomimus ; Type locality: Guatemala, near the city, Capetillo ).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Guatemala.

distans Sharp, 1885: 450 ( Paederomimus ; Type locality: Guatemala, Zapote ).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Guatemala.

flavoguttatus Bernhauer, 1908d: 332 ( Paederomimus ; Type locality: Brasilien).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Brazil.

gentilis Sharp, 1885: 439 ( Paederomimus ; Type locality: Mexico, Cerro de Plumas , Jalapa).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 62 ( Paederomimus ; necrophilous species; distribution in Mexico).

- DISTRIBUTION: Mexico.

hilaris Sharp, 1885 , see: sharpi Bernhauer and Schubert, 1914 .

hilaris Fauvel, 1874: 246 [= 1874a: 472] ( Philonthus ; replacement name for laetus Erichson ). — Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Colombia, Brazil.

laetus Erichson, 1840: 477 ( Philonthus ; [preoccupied]; Type locality: Columbia; Brasilia). — Sharp, 1885: 438 ( Paederomimus ).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; synonym of hilaris ).

imitator Bierig, 1937: 198 ( Paederomimus ; Type locality: Panama: Volcan de Chiriqui, 1500 m). - DISTRIBUTION: Panama.

insolitus Sharp, 1885: 447 ( Paederomimus ; Type locality: Panama, Bugaba 800 to 1500 feet). — Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Panama.

insularis Bernhauer, 1908 , see: lustralis Erichson, 1840 .

interjectus Bernhauer, 1908d: 333 ( Paederomimus ; Type locality: St. Vincent: W. J [I?]. Windward side).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

— Blackwelder, 1943: 397 ( Paederomimus ; characters; St. Vincent).

- DISTRIBUTION: West Indies.

smithi Cameron, 1922: 121 ( Paederomimus ; Type locality: St. Vincent).

— Scheerpeltz, 1933: 1376 ( Paederomimus ; catalog).

— Blackwelder, 1943: 397 ( Paederomimus ; synonym of interjectus ).

klimschi Bernhauer, 1917d: 36 ( Paederomimus ; Type locality: Brasilien: S. Catharina).

— Scheerpeltz, 1933: 1376 ( Paederomimus ; catalog).

- DISTRIBUTION: Brazil.

laetus Erichson, 1840 , see: hilaris Fauvel, 1874 .

longipes Sharp, 1876: 178 ( Philonthus ; Type locality: Obydos).

— Sharp, 1885: 438 ( Paederomimus ).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Brazil.

lustralis Erichson, 1840: 489 ( Philonthus ; Type locality: Puerto Rico).

— Sharp, 1885: 438 ( Paederomimus ).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

— Blackwelder, 1943: 398 ( Paederomimus ; characters; Puerto Rico; St. Thomas).

- DISTRIBUTION: West Indies.

insularis Bernhauer, 1908d: 333 ( Paederomimus ; Type locality: Ins. St. Thomas).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

— Blackwelder, 1943: 398 ( Paederomimus ; synonym of lustralis ).

lustrator Sharp, 1876: 172 ( Philonthus ; Type locality: Para).

— Sharp, 1885: 438 ( Paederomimus ).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Brazil.

maculipes Sharp, 1885: 445 ( Paederomimus ; Type locality: Mexico, Cordova , Jalapa).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Mexico.

mandibularis Sharp, 1885: 439 ( Paederomimus ; Type locality: Guatemala, Capetillo ).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Guatemala.

mexicanus Sharp, 1885: 444 ( Paederomimus ; Type locality: Mexico, Jalapa ).

— Scheerpeltz, 1933: 1376 ( Paederomimus ; catalog).

- DISTRIBUTION: Mexico.

nigrinus Sharp, 1885: 442 ( Paederomimus ; Type locality: Nicaragua, Chontales).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Nicaragua.

nigripennis Sharp, 1885: 446 ( Paederomimus ; Type locality: Panama, Bugaba 800 to

1500 feet, Volcan de Chiriqui from 2000 to 4000 feet).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Panama.

nigrocyaneus Schubert, 1909: 294 ( Paederomimus ; Type locality: südöstliches Mexico, Gegend von Jalapa, nördlich vom Pic d’Orizaba).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Mexico.

nobilis Sharp, 1885: 442 ( Paederomimus ; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Panama.

sydowi Bernhauer, 1917d: 37 ( Paederomimus ; Type locality: Brasilien: S. Paulo, Umgebung von Ribeirao Preto Fundao).

— Scheerpeltz, 1933: 1376 ( Paederomimus ; catalog).

- DISTRIBUTION: Brazil.

tetricus Blackwelder, 1943: 395 ( Paederomimus ; Type locality: Trinidad: Port of Spain).

- DISTRIBUTION: West Indies.

traili Sharp, 1876: 170 ( Philonthus ; Type locality: Ananá).

— Sharp, 1885: 438 ( Paederomimus ).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Brazil.

trinidadensis Bernhauer, 1915 , see: cribricollis Erichson, 1840 .

vicinus Sharp, 1885: 444 ( Paederomimus ; Type locality: Panama, Volcan de Chiriqui 2000 to 3000 feet).

— Bernhauer and Schubert, 1914: 368 ( Paederomimus ; catalog).

- DISTRIBUTION: Panama.

Paracraspedomerus

[1 species; Oceanic Region]

Paracraspedomerus Moore, 1960: 99 (species included: speculifrons ). TYPE SPECIES: Cafius speculifrons Fauvel , fixed by original designation and monotypy.

Xystridiogaster Scheerpeltz, 1966e: 402 (species included: speculifrons ). TYPE SPECIES: Cafius speculifrons Fauvel , fixed by original designation (p. 405).

— Moore, 1972: 218 (synonym of Paracraspedomerus ).

speculifrons Fauvel, 1877: 259 ( Cafius ; Type locality: Nouvelle Calédonie).

— Fauvel, 1889b: 263 ( Cafius ; notes; New Caledonia).

— Fauvel, 1903d: 269 ( Hesperus ; New Caledonia).

— Bernhauer and Schubert, 1914: 364 ( Hesperus ; catalog).

— Scheerpeltz, 1933: 1374 ( Hesperus ; catalog).

— Moore, 1960: 99 ( Paracraspedomerus ; New Caledonia).

— Scheerpeltz, 1966e: 402 ( Xystridiogaster ; characters; New Caledonia).

— Legner and Moore, 1977: 176 ( Paracraspedomerus ; found under bark).

- DISTRIBUTION: New Caledonia.

Pescolinus

[2 species; Neotropical Region]

Pescolinus Sharp, 1885: 453 (species included: palmatus ). TYPE SPECIES: Pescolinus palmatus Sharp , fixed by monotypy.

— Bernhauer and Schubert, 1914: 369 (world catalog; 1 species).

— Bierig, 1937: 191 (characters).

— Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and

the West Indies).

— Blackwelder, 1952: 299 (type species: palmatus ).

palmatus Sharp, 1885: 453 ( Pescolinus ; Type locality: Panama, Volcan de Chiriqui 4000 to 6000 feet).

— Bernhauer and Schubert, 1914: 369 ( Pescolinus ; catalog).

- DISTRIBUTION: Panama.

schmidti Bierig, 1937: 193 ( Pescolinus ; Type locality: Costa Rica: Vara Blanca , 2000 m).

- DISTRIBUTION: Costa Rica.

Peucoglyphus

[1 species; Oriental Region]

Peucoglyphus Bernhauer, 1926f: 317 (species included: corporaali ). TYPE SPECIES: Peucoglyphus corporaali Bernhauer , fixed by monotypy.

— Scheerpeltz, 1933: 1371 (world catalog supplement).

— Blackwelder, 1952: 299 (type species: corporaali ).

corporaali Bernhauer, 1926f: 318 ( Peucoglyphus ; Type locality: Buru).

— Scheerpeltz, 1933: 1371 ( Peucoglyphus ; catalog).

- DISTRIBUTION: Indonesia.

Phileciton

[2 species; Neotropical Region]

Phileciton Wasmann, 1894: 211 (species included: badariottii ). TYPE SPECIES: Phileciton badariottii Wasmann , fixed by monotypy.

— Bernhauer and Schubert, 1914: 326 (world catalog; 2 species).

— Blackwelder, 1944: 132 (checklist of species from Mexico, Central and South America, and

the West Indies).

— Blackwelder, 1952: 300 (type species: badariottii ).

— Seevers, 1965: 328 (doubtfully distinct from Erichsonius and Neobisnius ; myrmecophilus species; list of species).

badariottii Wasmann, 1894: 212 ( Phileciton ; Type locality: Sales. bei Lorena, Prov. S. Paulo).

— Bernhauer and Schubert, 1914: 326 ( Phileciton ; catalog).

— Seevers, 1965: 328 ( Phileciton ; notes; Brazil).

- DISTRIBUTION: Brazil.

wasmanni Bernhauer, 1908d: 334 ( Phileciton ; Type locality: Brasilien: S. Paulo).

— Bernhauer and Schubert, 1914: 326 ( Phileciton ; catalog).

— Seevers, 1965: 328 ( Phileciton ; doubtfully distinct from badariottii ).

- DISTRIBUTION: Brazil.

Philonthus

[1255 species (19 doubtful); Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, and Oceanic Regions]

Philonthus Stephens, 1829a: 23 (species included: laminatus , aeneus , splendens , politus , decorus , atratus , fimetarius , marginatus , concinnus , obscurus , varians , simplex , aterrimus , opacus , nitens , agilis , sanguinolentus , bimaculatus , cor[r]uscus, micans , and punctus along with 25 unavailable names; [Note: There is some dispute concerning the authorship of Philonthus . The name was used in 1827 and again in 1829. Westwood (1827: 58) described the larva of “ Philonthus politus ?” and presented fairly detailed characters and illustrations for the species. However, since he was unsure that the larva was identified accurately, we can reject as unavailable that use of Philonthus (Articles 12.2.5). In 1829 Philonthus was published in separate catalogs by both Curtis and Stephens. Tottenham (1962: 153) wrote that W. D. Hincks had informed him “... that he has seen definite evidence that the relevant [for date of Philonthus ] part of Curtis’ work was published in June...”. Earlier in the same paragraph Tottenham stated an August, 1829 publication date for Stephens’ work. According to a letter from N. D. Riley, a former Keeper at the British Museum (Natural History), sent to J. G. Franclemont in 1949 there is evidence to support a June 1, 1829 publication of Stephens’ “Nomenclature...” Unfortunately neither Tottenham nor Riley documented their opinions. Smetana (1995e: 51, 56) concluded no dates had been established and fixed the date of publication for both as December 31, 1829. According to evidence presented by D. Fletcher, 1979: xv, the date of publication was June 1, 1829 (see

note attached to the citation for Stephens, 1829a in bibliography)]). TYPE SPECIES: Staphylinus splendens Fabricius , fixed by subsequent designation by Curtis, 1836:

pl. 610.

— Curtis, 1829: 25 (catalog; Britain).

— Stephens, 1829: 279 (catalog; Britain).

— Stephens, 1832: 200, 226 (characters).

— Curtis, 1836: pl. 610 (characters; type species; Britain).

— Westwood, 1838a: 16 (characters; type species).

— Erichson, 1839a: 445 (characters).

— Stephens, 1839: 385 (characters).

— Heer, 1839: 257 (characters).

— Erichson, 1840: 426 (characters).

— Schaum, 1852: 26 (list of species; Europe).

— Lacordaire, 1854: 80 (characters; notes; list of species).

— Wollaston, 1854: 580 (characters; notes).

— Fairmaire and Laboulbène, 1856: 513 (characters).

— Jacquelin du Val, 1856: 35 (characters).

— Giebel, 1856: 72 (fossil species).

— Gistel, 1856: 388 (list of species).

— Kraatz, 1857d: 565 (characters).

— Redtenbacher, 1857: 188 (characters; key to Austrian species).

— Thomson, 1858: 30 (characters).

— G. Waterhouse, 1858: 24 (catalog of British species).

— Thomson, 1859: 24 (characters; type species: splendens ).

— Schaum, 1859: 26 (catalog; European species).

— Thomson, 1860: 153 (characters).

— LeConte, 1861a: 65 (characters).

— Gredler, 1863: 104 (list of species from Tirol).

— LeConte, 1863: 24 (list of North American species).

— Fauvel, 1866a: 341 [= 1866b: 92] (characters).

— Wencker and Silbermann, 1866: 29 (list of species; collecting notes; France).

— Gemminger and Harold, 1868: 584 (catalog).

— Jacquelin du Val, 1868: 68 (catalog; European species).

— Sharp, 1871d: 11 (list of British species).

— Fauvel, 1874: 207 [= 1874a: 433] (characters; key to species; larval characters).

— Redtenbacher, 1874: 207 (characters; key to Austrian species).

— Oustalet, 1874: 167 (characters; description of fossil species).

— Fauvel, 1875a: XXIX [= 1875b: 231] (catalog).

— Mulsant and Rey, 1876: 344 [= 1877: 200] (characters; key to French species).

— Sharp, 1876: 165 (characters; notes).

— Fauvel, 1877: 262 (notes; key to species).

— Provancher, 1877: 248, 730 (characters; key to species).

— Fauvel, 1878f: 257 (characters; notes; key to New Guinea species).

— Broun, 1880: 110 (characters).

— Heyden, 1880: 73 (list of species of Siberian region).

— Duvivier, 1883: 144 (catalog).

— Lynch, 1884: 146 (characters; key for Argentine species).

— Horn, 1884: 179 (characters; key to North American species).

— Sharp, 1885: 395 (characters; notes; Mexican and Central American species).

— Fowler, 1888: 258 (notes; larval characters; key to British species).

— Seidlitz, 1891: 382 (key to some European species).

— Seidlitz, 1891a: 407 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 103 (list of species of Europe and Caucasus).

— Heyden, 1893: 42 (list of species of Siberian region).

— Ganglbauer, 1895: 437 (adult and larval characters; key to middle European species).

— Heyden, 1896: 32 (list of species of Siberian region).

— Fauvel, 1897d: 299 (catalog of species of Barbary; Azores; Salvages; Madeira; Canaries).

— Germain, 1903: 414 (characters; notes).

— Fauvel, 1903d: 270 (key to New Caledonia species).

— Heyden, Reitter, and Weise, 1906: 161 (list of species of Europe and Caucasus).

— Reitter, 1909: 116 (characters).

— Blatchley, 1910: 379 (characters; key to Indiana species).

— Petri, 1912: 71 (list of species of Siebenbürgen).

— Bernhauer and Schubert, 1914: 326 (world catalog; 574 species).

— Johansen, 1914: 388 (characters; key to Danish species).

— Leng, 1920: 106 (catalog of North American species).

— Moore and Legner, 1976: 541 (includes seashore species; notes).

— Pope, 1977: 29 (list of British species).

— Moore and Miller, 1978: 38 (two unnamed species from Pleistocene asphalt deposits; California).

— Topp, 1978: 312 (larval characters).

— E. Kangas, 1979: 101 (key to species of varians group).

— Moore and Legner, 1979: 28 (characters; notes).

— Muona, 1979: 14 (list of Scandinavian species).

— Uhlig, 1979: 252 (some species collected in Germany).

— M. Dvořák, 1979: 115 (some species collected in Slovakia).

— Uhlig, Vogel, and Sieber, 1980: 246 (some species collected in Germany).

— Burakowski, Mroczkowski, and Stefańska, 1980: 38 (catalog; Poland).

— Uhlig and Vogler, 1981: 111 (list of some species collected in Germany; habitat and collecting notes).

— Spahr, 1981: 96 (references to specimens from amber and copal).

— Boller, 1983 (larval characters; larval key to some species).

— Dettner, 1983 (comparative anatomy of defensive glands and defensive secretions).

— Shibata, 1983: 90 (checklist of species of Japan).

— Hammond, 1984: 206 (checklist of species of Borneo).

— Tóth, 1984: 41 (characters; habitus; key to species of Hungary).

— Yuh, Paik, Kwon, and Lee, 1985: 240 (checklist of species of Korea).

— Frank, 1986a: 370 (checklist of species of Florida).

— Muona and Viramo, 1986: 13 (list of species of northeastern Finland).

— Segers, 1986: 43 (checklist of species; Belgium).

— Drugmand 1987b (comments on distribution of Belgian species).

— Ádám, 1987: 149 (list of 33 species with collecting notes; Hungary).

— Lohse and Lucht, 1989: 163 (supplement to key to species of Lohse, 1964).

— Lecoq, 1990: 202 (key to species of Mascarene Islands).

— Angelini, 1991: 192 (list of species from high elevations of Calabria, Italy).

— M. Hansen, Kristensen, Mahler, Pedersen, 1991: 107 (list of Danish species).

— Smetana, 1991a: 227 (key to species of furvus complex of Mexico and Central America).

— Campbell and Davies, 1991: 117 (checklist of species; Alaska and Canada).

— Silfverberg, 1992: 16 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Navarrete-Heredia and Márquez-Luna, 1993 (cave collections; Mexico).

— Siitonen, 1993: 230 (list of Finnish species).

— Hayashi, 1993: 81 (key to species of prolatus group).

— Burakowski, 1993: 22, 23 (rearing techniques).

— Welch, 1993: 228 (ovariole structure and development).

— Smetana, 1993b: 48 (list of species of Czech Republic and Slovakia).

— Hayashi, 1994: 120, 126 (notes on classification of Philonthus and related genera; diagnostic characters; subgeneric characters).

— Hernández, Outerelo, and Gamarra, 1994: 188 (checklist of species; Canary Islands).

— Smetana, 1995e: 51 (characters; classification and revision of and key to species of USA and Canada; phylogeny of species groups; author of genus is Stephens).

— Hu and Frank, 1995b (surface structure of eggs of five species and key for identification of eggs [p.587]).

— Navarrete-Heredia, 1995: 96 (list of species of Mexico).

— M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 28 (additional records to Danish list of species).

— Ciceroni and Zanetti, 1995: 27 (list of species of Italy).

— M. Hansen, 1996: 104 (list of Danish species).

— M. Hansen, Mahler, Palm, and Pedersen, 1996: 243 (additions to list of Danish species).

— Lauterer and Opravil, 1996: 244 (subfossil; 16th century; Czech Republic).

— Downie and Arnett, 1996: 375 (characters; key to species of Northeastern North America).

— Lecoq, 1997: 280 (key to “yellow” species of Madagascar).

— Schillhammer, 1998 (revision of and key to species of cyanipennis group; notes).

— Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).

— M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (list of species collected in 1997; Denmark).

— Assing, Frisch, Kahlen, et al., 1998: 137 (supplement to key to species in Lohse, 1964; notes).

— Schillhammer, 1999: 57 (key to species of spinipes and cinctulus species groups).

— Assing and Schülke, 1999: 14 (authorship of Philonthus ; notes; synonymic discussion).

— Schillhammer, 2000: 113 (revision of and key to species of politus complex; species group listing; correction on page 159 of scale bars in Schillhammer, 1999).

— Newton, Thayer, Ashe, and Chandler, 2000: 396 (112 Nearctic species; notes; characters in key).

— Hayashi, 2000: 23 (notes on empodial setae).

— Naomi, Kuranishi, Saito, and Maruyama, 2000: 109 (collected from Kuril Islands and Kamchatka).

— Jimenez-Sanchez, Navarrete-Heredia, and Padilla-Ramirez, 2000: 62 (necrophilous species

in Mexico).

Laxobates Gistel, 1834: 8 (species included: coenosus , cyanipennis , splendens , laminatus , aeneus , metallicus , cephalotes , decorus , politus , atratus , carbonarius , ebeninus cited as ebeneninus, varians , varius , immundus , fuscus , sanguinolentus , tenuis , fimetarius , agilis , splendidulus along with 12 unavailable names). TYPE SPECIES: Staphylinus splendens Fabricius , fixed by subsequent designation by Blackwelder, 1952: 212.

— Smetana, 1958: 143 (synonym of Philonthus ).

— Smetana, 1995e: 51 (synonym of Philonthus ).

Eccoptolonthus Bernhauer, 1912f: 206 (subgenus of Philonthus ; species included: conradti ). TYPE SPECIES: Philonthus (Eccoptolanthus) conradti Bernhauer , fixed by monotypy.

— Bernhauer and Schubert, 1914: 326 (synonym of Philonthus ).

— Cameron, 1932: 62 (subgenus of Philonthus ).

— Blackwelder, 1943: 399 (subgenus of Philonthus ; type species: conradti ).

— Blackwelder, 1952: 138 (subgenus of Philonthus ; type species: conradti ).

Pseudophilonthus Bernhauer, 1915 k: 302 (subgenus of Philonthus ; species included: bicoloripennis ). TYPE SPECIES: Philonthus (Pseudophilonthus) bicoloripennis Bernhauer , fixed by monotypy.

— Bernhauer, 1932b: 147 (valid genus).

— Scheerpeltz, 1933: 1330 (subgenus of Philonthus ).

— Blackwelder, 1943: 399 (subgenus of Philonthus ; type species: bicoloripennis ).

— Blackwelder, 1952: 329 (subgenus of Philonthus ; type species: bicoloripennis ).

Onychophilonthus Neresheimer and Wagner, 1924: 156 (subgenus of Philonthus ; species included: marginatus ). TYPE SPECIES: Staphylinus marginatus Ström , fixed by monotypy.

— Scheerpeltz, 1933: 1330 (subgenus of Philonthus ).

— Blackwelder, 1943: 399 (subgenus of Philonthus ; type species: marginatus ).

— Tottenham, 1949: 372, 419 (type species: marginatus Ström ; subgenus of Philonthus ; checklist of British species).

— Blackwelder, 1952: 276 (subgenus of Philonthus ; type species: marginatus ).

— Smetana, 1958: 143, 226 (subgenus of Philonthus ; characters; type species).

— Smetana, 1959: 170 (subgenus of Philonthus ; characters).

— Smetana, 1963d: 402 (subgenus of Philonthus ; characters; key to species).

— Coiffait, 1974: 116, 120, 190, 305 (subgenus of Philonthus ; characters; key to species of western Palaearctic region).

Nephronthus Bernhauer, 1932b: 147 (subgenus of Pseudophilonthus ; species included: grandis ). TYPE SPECIES: Pseudophilonthus (Nephronthus) grandis Bernhauer ,

fixed by monotypy.

— Scheerpeltz, 1933: 1330 (subgenus of Philonthus ).

— Blackwelder, 1943: 399 (subgenus of Philonthus ; type species: grandis ).

— Blackwelder, 1952: 261 (subgenus of Philonthus ; type species: grandis ).

Philonthopsis Cameron, 1932: 261 (species included: antennalis ). TYPE SPECIES: Philonthopsis antennalis Cameron, 1932 , fixed by monotypy.

— Scheerpeltz, 1933: 1418 (world catalog supplement).

— Blackwelder, 1952: 301 (type species: sabulosus ).

— Scheerpeltz, 1965: 279 (characters; type species: antennalis ).

— Schillhammer, 1994: 74 (synonym of Philonthus ).

Cephalonthus Blackwelder, 1952: 96 (subgenus of Philonthus ; for species included see Bernhauer, 1940b). TYPE SPECIES: Philonthus caffer Boheman , fixed by original designation (see Herman, 2001).

— Bernhauer, 1940b: 635 (subgenus of Philonthus ; species included: lewisius , caffer , ustus , kochianus ; characters; [Note: This name is unavailable because Bernhauer did not designate

a type species]).

— Herman, 2001: 9 (name unavailable when proposed by Bernhauer, 1940, made available by Blackwelder, 1952; Blackwelder author of name, not Bernhauer).

Philonthoblerius Tottenham, 1949b: 300 (subgenus of Philonthus ; species included: schoutedeni ). TYPE SPECIES: Philonthus schoutedeni Bernhauer , fixed by original designation and monotypy.

— Blackwelder, 1952: 301 (subgenus of Philonthus ; type species: schoutedeni ).

Raucalius Tottenham, 1949b: 304 (subgenus of Philonthus ; species included: rugulipennis , peripateticus , madianus ). TYPE SPECIES: Philonthus (Raucalius) peripateticus Tottenham , fixed by original designation.

— Blackwelder, 1952: 337 (subgenus of Philonthus ; type species: peripateticus ).

— Levasseur, 1967b: 1624 (notes; characters).

Spatulonthus Tottenham, 1955: 154 (subgenus of Philonthus ; species included: longicornis , kasaiensis , vestigator , ugandae , bestialis , bisignatus , deleterius , delusor , ridens , minutus , sanamus , nitidicollis , incognitus , peregrinus , igacus , labdanus , maskinius , ponderosus , tachyoryctides, limulus , bishanus ). TYPE SPECIES: Philonthus longicornis Stephens , fixed by original designation.

— Tottenham, 1955: 155 (key to species).

— Coiffait, 1963: 8, 18 (characters; notes).

— Coiffait and Saiz, 1968: 348, 360 (characters; notes).

— Bordoni, 1973j: 707 (list of species from islands near Sicily).

— Coiffait, 1974: 308 (adult and larval characters; key to species of western Palaearctic region).

— Burakowski, Mroczkowski, and Stefańska, 1980: 36 (catalog; Poland).

— Segers, 1986: 46 (checklist of species; Belgium).

— Drugmand 1987b (comments on distribution of Belgian species).

— Ádám, 1987: 151 (list of three species with collecting notes; Hungary).

— Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Smetana, 1995e: 51 (synonym of Philonthus ).

Kenonthus Coiffait, 1960: 361 (subgenus of Philonthus ; species included: montivagus , aerosus , laevicollis , nimbicola ). TYPE SPECIES: Philonthus montivagus Heer , fixed

by original designation.

— Coiffait, 1974: 116, 120, 186, 298 (subgenus of Philonthus ; characters; key to species of western Palaearctic region).

Trionthus Coiffait, 1963: 14 (subgenus of Philonthus ; species included: lepidus ). TYPE SPECIES: Staphylinus lepidus Gravenhorst , fixed by monotypy.

— Coiffait, 1974: 116, 120, 193 (subgenus of Philonthus ; characters; key to species of western Palaearctic region).

— Ádám, 1987: 151 (cited as valid genus; list of one species with collecting notes; Hungary).

— Smetana, 1995e: 51 (synonym of Philonthus ).

Paragabrius Coiffait, 1963: 12 (species included: micans , micanthoides, furcifer , palustris fulvipes , virgo , fagniezi , truquii , mimus ). TYPE SPECIES: Staphylinus micans Gravenhorst , fixed by original designation.

— Coiffait, 1974: 84 (characters; key to species of western Palaearctic region).

— Pace and Zanetti, 1977: 301 (characters; notes; relationships).

— Burakowski, Mroczkowski, and Stefańska, 1980: 34 (catalog; Poland).

— Segers, 1986: 43 (checklist of species; Belgium).

— Drugmand 1987b (comments on distribution of Belgian species).

— Ádám, 1987: 151 (cited as valid genus; list of three species with collecting notes; Hungary).

— Angelini, 1991: 192 (list of species from high elevations of Calabria, Italy).

— Silfverberg, 1992: 16 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Smetana, 1995e: 51 (synonym of Philonthus ).

Kirschenblatia Bolov and Kryzhanovskij, 1969: 512 (species included: kabardensis ). TYPE SPECIES: Kirschenblatia kabardensis Bolov and Kryzhanovskij , fixed by original designation.

— Bolov and Kryzhanovskij, 1969a: 327 (translation of Bolov and Kryzhanovskij, 1969).

— Schülke and Uhlig, 1989: 247 (synonym of Philonthus ).

Palaeophilonthus Coiffait, 1972: 50 (proposed for Coiffait’s [1965g: 616] misidentification of Sectophilonthus ; species included: rossicus ). TYPE SPECIES: Sectophilonthus rossicus Coiffait , fixed by monotypy.

— Coiffait, 1965g: 616 (characters; type species; [Note: Misidentified as Sectophilonthus according to Coiffait, 1972: 50]).

— Coiffait, 1974: 5 (characters; species of western Palaearctic region).

— Smetana, 1995e: 51 (synonym of Philonthus ).

Metagabrius Coiffait, 1974: 84 , 86 (subgenus of Paragabrius ; species included: juvenilis , fuscatus ). TYPE SPECIES: Philonthus juvenilis Peyerimhoff , fixed by original designation.

— Smetana, 1995e: 51 (synonym of Philonthus ).

Parambigus Marcuzzi, 1977: 12 (subgenus of Philonthus ; without description; [Note: Nomen nudum, Article 13.1.1]; species included: lucidus Fauvel ).

Glaphyronthus Scheerpeltz, 1978: 220 (subgenus of Philonthus ; species included: tenellicornis ). TYPE SPECIES: Philonthus (Glaphyronthus) tenellicornis Scheerpeltz , fixed by original designation and monotypy.

Paralionthus Ádám, 1996: 236 (species included: punctus ). TYPE SPECIES: Staphylinus punctus Gravenhorst , fixed by original designation and monotypy.

— Assing and Schülke, 1999: 14 (synonym of Philonthus ).

Piezarthrus Lecoq, 1998: 63 (subgenus of Philonthus ; species included: pollux , steineri , pauliani , viettei , marojejy , humbloti , castor , rectus ). TYPE SPECIES: Philonthus pollux Fauvel , fixed by original designation.

abactus Sharp, 1876 , see: Paederomimus .

abauriae Gridelli, 1924 , see: Rabigus .

abavus † Scudder, 1900: 57 ( Philonthus ; fossil; Type locality: Florissant , Colorado) .

— Handlirsch, 1907: 726 ( Philonthus ; catalog; Miocene; Colorado).

- DISTRIBUTION: USA.

abdicans Tottenham, 1949b: 313 ( Philonthus ; Type locality: South Africa).

— Tottenham, 1962: 163 ( Philonthus ; characters).

- DISTRIBUTION: South Africa.

abdominalis Erichson, 1847: 89 ( Philonthus ; Type locality: Republica Peruana).

— Bernhauer and Schubert, 1914: 326 ( Philonthus ; catalog).

- DISTRIBUTION: Peru.

aberdarensis Bernhauer, 1939 , see: Gabrius .

aberrans Sharp, 1876 , see: Paederomimus .

aberrans Cameron, 1932: 111 ( Philonthus ; [preoccupied]; Type locality: Mussoorie district; Mossy Falls; Keyarkuli. Assam: Naga Hills , Laimatak ).

— Scheerpeltz, 1933: 1330 ( Philonthus ; catalog).

— Herman, 2001: 46 ( Philonthus ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: India.

aberti Hatch, 1957: 191 ( Philonthus ; Type locality: Oregon: L. Abert).

— Smetana, 1995e: 307 ( Philonthus ; characters; California; Nevada; Oregon).

- DISTRIBUTION: USA.

abkhazensis Coiffait, 1963 , see: rufipes Stephens, 1832 .

abnormalis Sharp, 1885: 397 ( Philonthus ; Type locality: Mexico, Oaxaca).

— Bernhauer and Schubert, 1914: 326 ( Philonthus ; catalog).

— Navarrete-Heredia, 1995: 96 ( Philonthus ; Mexico; Guatemala).

- DISTRIBUTION: Guatemala, Mexico.

abyssinus Fauvel, 1881a: 203 ( Belonuchus ; Type locality: Scioa, Mahal-Uonz; Let Marefia). — Fauvel, 1907: 39 ( Philonthus ; Naivasha; Nairobi; Kilimandjaro; Kiboscho; Abyssinia; Gallas; Choa; Usambara; Mozambique; Mashunaland; Cape; Angola).

— Eichelbaum, 1913: 130 ( Philonthus ; Tanzania).

— Bernhauer and Schubert, 1914: 326 ( Philonthus ; catalog).

— Rambousek, 1922: 1008 ( Philonthus ; Ethiopia).

— Scheerpeltz, 1933: 1330 ( Philonthus ; catalog).

— Tottenham, 1949b: 315 ( Philonthus ).

— Cameron, 1950g: 38 ( Philonthus ; Belgian Congo).

— Tottenham, 1962: 164 ( Philonthus ; characters).

— Frank, 1982: 38 ( Philonthus ; list of parasites).

- DISTRIBUTION: Ethiopia, Kenya, Congo, Tanzania, Mozambique, Zimbabwe,

South Africa, Angola.

holomelas Eppelsheim, 1895: 201 ( Philonthus ; Type locality: Ganale Gudda, Arussi Galla). — Eppelsheim, 1895: 213 ( Philonthus ; synonym of abyssinus ).

— Fauvel, 1907: 39 ( Philonthus ; synonym of abyssinus ).

— Bernhauer and Schubert, 1914: 327 ( Philonthus ; synonym of abyssinus ).

acamas Smetana, 1995e: 281 ( Philonthus ; Type locality: Oregon: Austin House , Jct. Hwys.

26 & 7, 1005 m. Also cited from California, Colorado, Nevada, Oregon).

- DISTRIBUTION: USA.

accedens Sharp, 1885: 403 ( Philonthus ; Type locality: Guatemala, Capetillo ).

— Bernhauer and Schubert, 1914: 327 ( Philonthus ; catalog).

— Frank, 1982: 38 ( Philonthus ; list of parasites).

- DISTRIBUTION: Guatemala.

acceptus Last, 1973: 429 ( Philonthus ; Type locality: N.E. New Guinea: Karkar Island ).

— Last, 1987: 31 ( Philonthus ; Papua New Guinea).

- DISTRIBUTION: New Guinea.

aciculatus Stephens, 1832 , see: sanguinolentus Gravenhorst, 1802 .

acroleucus Kraatz, 1859: 91 ( Philonthus ; Type locality: Ceylan; India orientalis).

— Bernhauer, 1902b: 27 ( Philonthus ; Ceylon).

— Bernhauer and Schubert, 1914: 327 ( Philonthus ; catalog).

— Cameron, 1932: 115 ( Philonthus ; characters; Ceylon).

— Scheerpeltz, 1933: 1330 ( Philonthus ; catalog; Philippines).

— Cameron, 1933d: 387 ( Philonthus ; Sumatra).

— Cameron, 1937: 5 ( Philonthus ; Java).

— Frank, 1982: 38 ( Philonthus ; list of parasites).

- DISTRIBUTION: Sri Lanka, “East Indies”, Indonesia, Philippines.

acromyrmecis Scheerpeltz, 1976b: 135 ( Philonthus ; subgenus Philonthus ; Type locality: Kolonie Nova Teutonia, Brasilien).

- DISTRIBUTION: Brazil.

— Raichev, 1979: 82 ( Philonthus ; Bulgaria).

— Tóth, 1984: 48 ( Philonthus ; characters; Hungary).

— Yuh, Paik, Kwon, and Lee, 1985: 240 ( Philonthus ; Korea).

— Lucht, 1987: 105 ( Philonthus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Schülke and Uhlig, 1988: 11 ( Philonthus ; notes; Germany).

— J. Li, 1993: 36 ( Philonthus ; China).

— Smetana, 1993b: 48 ( Philonthus ; subgenus Philonthus ; Czech Republic; Slovakia).

— Hayashi, 1994: 124 ( Philonthus ; characters; Japan).

— Ciceroni and Zanetti, 1995: 27 ( Philonthus ; Italy).

— Sterrenburg and Schülke, 1997: 18 ( Philonthus ; Netherlands).

— Owen, 1997a: 149 ( Philonthus ; collecting sites; Ireland).

— Schillhammer, 1999a: 61 ( Philonthus ).

— Kashcheev, 1999a: 57 ( Philonthus ; Kazakhstan).

— Schillhammer, 2000: 134 ( Philonthus ; characters; Russia; Mongolia; North Korea; Japan; China). - DISTRIBUTION: Europe, Russia, Kazakhstan, Mongolia, Korea, China, Japan.

temporalisoides Drugmand, 1988b: 249 ( Philonthus ; subgenus Philonthus ; Type locality: U.R.S.S.: lac Baikal, Sibérie).

— Schillhammer, 1999a: 61 ( Philonthus ; synonym of addendus ).

— Schillhammer, 2000: 134 ( Philonthus ; synonym of addendus ).

adjacens Cameron, 1937: 7 ( Philonthus ; Type locality: Java: G. Tangkoeban Prahoe,

alt. 4000–5000 ft).

- DISTRIBUTION: Indonesia.

adscitus Märkel and Kiesenwetter, 1848 , see: laevicollis Lacordaire, 1835 .

adustus Casey, 1915 , see: Gabrius .

adversarius Bernhauer, 1917 g: 46 ( Philonthus ; replacement name for kamerunensis Bernhauer, 1915 ).

— Scheerpeltz, 1933: 1330 ( Philonthus ; catalog).

- DISTRIBUTION: Cameroon.

kamerunensis Bernhauer, 1915 k: 301 ( Philonthus ; [preoccupied]; Type locality: Kamerun).

— Scheerpeltz, 1933: 1330 ( Philonthus ; synonym of adversarius ).

adversus Bernhauer and Schubert, 1914: 327 ( Philonthus ; replacement name for misellus Fauvel ).

— Cameron, 1932: 100 ( Philonthus ; characters; Burma; Assam; Darjeeling).

— Scheerpeltz, 1933: 1330 ( Philonthus ; catalog).

- DISTRIBUTION: Myanmar, India.

misellus Fauvel, 1895: 262 ( Philonthus ; [preoccupied]; Type locality: Bírmanie,

Carin Cheba, 900–1100 m).

— Bernhauer and Schubert, 1914: 327 ( Philonthus ; synonym of adversus ).

— Cameron, 1932: 100 ( Philonthus ; synonym of adversus ).

adversus Cameron, 1925: 59 ( Philonthus ; [preoccupied]; replacement name for misellus Fauvel ).

adversus Cameron, 1925 , see: adversus Bernhauer and Schubert, 1914 .

aeger Eppelsheim, 1895c: 59 ( Philonthus ; Type locality: Simla im Himalaya).

— Bernhauer and Schubert, 1914: 327 ( Philonthus ; catalog).

— Cameron, 1932: 83 ( Philonthus ; characters; India).

— Scheerpeltz, 1933: 1330 ( Philonthus ; catalog).

— Coiffait, 1982a: 233 ( Philonthus ; Bhutan).

— Schillhammer, 2000: 140 ( Philonthus ; characters; India; Nepal; China).

- DISTRIBUTION: China, Nepal, Bhutan, India.

aeolus Herman, 2001: 46 ( Philonthus ; replacement name for temporalis Coiffait ).

- DISTRIBUTION: Bhutan.

temporalis Coiffait, 1977b: 221 ( Paragabrius ; [preoccupied]; Type locality: Bhutan: Dorjula , 2900 m).

— Smetana, 1995e: 51 ( Philonthus ; transferred from Paragabrius by reason of Smetana’s synonymy of the genus with Philonthus ).

— Herman, 2001: 46 ( Philonthus ; synonym of aeolus ).

aemulus Tottenham, 1939d: 233 ( Philonthus ; replacement name for serraticornis Bernhauer ).

— Tottenham, 1949b: 361 ( Philonthus ; characters).

— Tottenham, 1961: 205 ( Philonthus ; Senegal).

— Levasseur, 1966a: 1214 ( Philonthus ; Tanganyika; Senegal; Congo).

— Levasseur, 1968a: 54 ( Philonthus ; Central African Republic).

- DISTRIBUTION: Namibia, Congo, Tanzania, Central African Republic, Senegal.

serraticornis Bernhauer, 1908c: 110 ( Philonthus ; [preoccupied]; Type locality: Deutsch- Südwestafrika: Otjosondu ).

— Tottenham, 1939b: 233 ( Philonthus ; synonym of aemulus ).

antennatus Bernhauer and Schubert, 1914: 328 ( Philonthus ; replacement name for serraticornis Bernhauer ; [preoccupied]).

aeneiceps Sharp, 1876 , see: Paederomimus .

aeneicollis Sharp, 1876 , see: Paederomimus .

aeneipennis Boheman, 1858: 30 ( Philonthus ; Type locality: China).

— Fauvel, 1903: 159 ( Philonthus ; India).

— Fauvel, 1904a: 59 ( Philonthus ; Belgaum; Ceylan; Japan; Celebes; Maurice; China;

New Guinea).

— Bernhauer, 1907: 386 ( Philonthus ; Japan).

— Bernhauer and Schubert, 1914: 327 ( Philonthus ; catalog).

— Bernhauer, 1922c: 232 ( Philonthus ; Formosa).

— Bernhauer, 1928: 16 ( Philonthus ; Sumatra).

— Cameron, 1932: 113 ( Philonthus ; characters; Ceylon; India).

— Scheerpeltz, 1933: 1330 ( Philonthus ; catalog).

— Cameron, 1933d: 386 ( Philonthus ; Sumatra).

— Cameron, 1937: 5 ( Philonthus ; Java).

— Cameron, 1949b: 467 ( Philonthus ; China).

— Scheerpeltz, 1957c: 233 ( Philonthus ; Sumba).

— Nakane and Sawada, 1960: A123 ( Philonthus ; Yakushima Island).

— Scheerpeltz, 1960: 5 ( Philonthus ; Pakistan; Iran; Afghanistan).

— Nakane, 1963a: 91 ( Philonthus ; characters; habitus photograph; Japan).

— Scheerpeltz, 1976a: 87 ( Philonthus ; Nepal).

— Watanabe and Shibata, 1972: 64 ( Philonthus ; Yaku-shima Island).

— Coiffait, 1977b: 206 ( Philonthus ; Bhutan).

— Coiffait, 1981f: 329 ( Philonthus ; Andaman Islands).

— Coiffait, 1982a: 232 ( Philonthus ; India).

— Frank, 1982: 38 ( Philonthus ; list of parasites).

— J. Li, 1993: 36 ( Philonthus ; China).

— Yadav, Chand, and Yadav, 1994: 6 ( Philonthus ; morphology of mitotic and meiotic chromosomes).

— Watanabe, 1998a: 140 ( Philonthus ; Japan).

— Watanabe, 1998b: 262 ( Philonthus ; Japan).

- DISTRIBUTION: China, Japan, Taiwan, New Guinea, Indonesia, Andaman Islands, Bhutan, Nepal, India, Pakistan, Afghanistan, Iran, Yemen, Mauritius, Tanzania, Senegal.

cliens Eppelsheim, 1890b: 277 ( Philonthus ; Type locality: Nagpore , India centralis).

— Fauvel, 1903: 159 ( Philonthus ; variety of aeneipennis ).

— Fauvel, 1904a: 59 ( Philonthus ; variety of aeneipennis ; Khandesh; Kanara; Aden).

— Bernhauer and Schubert, 1914: 327 ( Philonthus ; variety of aeneipennis ).

— Cameron, 1932: 113 ( Philonthus ; synonym of aeneipennis ).

— Scheerpeltz, 1933: 1330 ( Philonthus ; variety of aeneipennis ).

— Gridelli, 1936a: 151 ( Philonthus ; characters; valid species).

— Tottenham, 1961: 204 ( Philonthus ; Senegal).

— Tottenham, 1962: 139, 223, 226 ( Philonthus ; characters; Tanganyika).

— Scheerpeltz, 1976a: 87 ( Philonthus ; synonym of aeneipennis ).

— Coiffait, 1982: 28 ( Philonthus ; subspecies of erythropus ; Nepal).

— Coiffait, 1982a: 233 ( Philonthus ; variety of aeneipennis ; Nepal).

— Coiffait, 1984e: 373 ( Philonthus ; variety of aeneipennis ; Nepal).

punctatissimus Schubert, 1908: 619 ( Philonthus ; Type locality: Pathanka, Himalaya).

— Bernhauer and Schubert, 1914: 352 ( Philonthus ; catalog).

— Cameron, 1932: 113 ( Philonthus ; synonym of aeneipennis ).

— Scheerpeltz, 1933: 1330 ( Philonthus ; synonym of aeneipennis ).

lindemanni Scheerpeltz, 1960: 5 ( Philonthus ; variety of aeneipennis ; Type locality:

West Pakistan: Rawalpindi Umg.).

aeneipennis Cameron, 1932 , see: distincticornis Cameron, 1932 .

aeneoceps Broun, 1880: 111 ( Philonthus ; Type locality: Whangarei Harbour).

— Bernhauer and Schubert, 1914: 327 ( Philonthus ; catalog).

- DISTRIBUTION: New Zealand.

aenescens Solsky, 1870a: 262 ( Philonthus ; Type locality: Mexico).

— Sharp, 1885: 408 ( Philonthus ; Mexico).

— Bernhauer and Schubert, 1914: 327 ( Philonthus ; catalog).

- DISTRIBUTION: Mexico.

aeneus De Geer, 1774 , see: laminatus Creutzer, 1799 .

aeneus Rossi, 1790 , see: politus Linné, 1758 .

aequalis Horn, 1884: 205 ( Philonthus ; Type locality: Canada, Lake Superior region ; [Note: See lectotype designation by Smetana, 1995e]).

— Bernhauer and Schubert, 1914: 327 ( Philonthus ; catalog).

— Smetana, 1963a: 4 ( Philonthus ; characters; Montreal; Illinois; Iowa; Massachusetts; Kansas; Missouri; New Jersey).

— Frank, 1982: 38 ( Philonthus ; list of parasites).

Smetana, 1995e: 368 ( Philonthus ; lectotype designation: Lake Superior ; characters; notes; Manitoba; Ontario; Quebec; Arkansas; Connecticut; Georgia; Illinois; Indiana; Iowa; Kansas; Kentucky; Maine; Maryland; Massachusetts; Michigan; Missouri; Nebraska; New Hampshire; New Jersey; New York; North Carolina; Ohio; Oklahoma; Pennsylvania; Tennessee; Texas; Virginia; West Virginia; Wisconsin) .

— Downie and Arnett, 1996: 380 ( Philonthus ; characters; USA).

- DISTRIBUTION: Canada, USA.

viperinus Horn, 1884: 205 ( Philonthus ; cited as a Fauvel manuscript name without characters or locality; [Note: Nomen nudum, Article 12.1]).

— Scheerpeltz, 1933: 1367 ( Philonthus ; cited as a valid species; [Note: The name is still unavailable]).

— Smetana, 1963a: 3, 4 ( Philonthus ; [Note: Smetana cited “... viperinus Fauvel , in litt.” (p. 3) followed by a long discussion then listed it as a synonym of aequalis ; it is still unavailable]). aeratus Stephens, 1832 , see: intermedius Lacordaire, 1835 .

aeripennis Cameron, 1943: 130 ( Philonthus ; Type locality: Darjeeling: Ghum district:

Tiger Hill, altitude 8500–10,000 feet).

- DISTRIBUTION: India.

aerosus Kiesenwetter, 1851a: 416 ( Philonthus ; Type locality: Alpes de la Styrie).

— Kraatz, 1857d: 587 ( Philonthus ; characters; Alps).

— Redtenbacher, 1857: 191 ( Philonthus ; characters; Austria).

— Fauvel, 1874: 232 [= 1874a: 458] ( Philonthus ; characters; Styria; Moravia; Switzerland;

Tirol; Sicily).

— Redtenbacher, 1874: 210 ( Philonthus ; characters; Austria).

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Gabrius

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

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