Cicindela (Cicindela) lewisii lewisii Bates, 1873

Fukuda, Yuki, Ogawa, Ryo & Hori, Michio, 2019, The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia, Zootaxa 4661 (2), pp. 271-308 : 301

Cicindela (Cicindela) lewisii lewisii Bates, 1873

( Figs 8I , 15D )

Cicindela lewisii Bates, 1873: 226 .

Cicindela lewisii kagoshimanensis Iga, 1955:15 [synonymy by Nakane 1976:3].

Cicindela (Cicindela) lewisii: Rivalier 1950:225 .

Specimens examined. (N = 3) [ Japan] 1♂ ( YFJ), Tokushima, Yoshinogawa-kakô, Tokushima-City , 31. VII. 2012, Y. Fukuda leg ; 1♂ ( EUMJ), Tokushima, Estuary of Yoshinogawa River , Tokushima city, 26-28. VIII. 1996, K. Aita leg ; 1♂ ( EUMJ), Tokushima, Kago, Ohbara-chô , 17. VIII. 1985, H. Iwasaki leg.

Description. Internal sac everted dors-apically in lateral view. VA largest in bladders; DA the second largest; B and VLR the third largest; DLL and VLL smallest in bladders.

DLL roundly swollen, vested with fine spines. DA well swollen basally. VA triangular swollen, everted left laterally, vested numerously with short and weakly chitinous spines. VLR swollen at base of VA, everted left laterally. VLL roundly swollen, everted right laterally numerously with well sclerotized spines. B roundly well swollen, everted right laterally, numerously with short spines, which are forming a band. Flagellum well short, curved. Medial tooth long, separated at middle.

Remarks. This species is clearly distinguished from other members of the subgenus Cicindela by sharing the characteristics of the position of each bladder but differs by the following characteristics: VA triangular swollen, everted left laterally; medial tooth separated at middle.

Figures

FIGURE 8. Habitus of Cosmodela, Cicindela (Cicindela) and Cylindera.A, Cosmodela sp. (cf. separata); B, C. batesi (Fleutiaux) from Iriomote Is., Japan; C, C. aurulenta (Fabricius) from Penang Is., Malaysia; D, C. virgula (Fleutiaux) from Laos; E, C. duponti duponti (Dejean) from South India; F, Cicindela (Cicindela) japana japana Motschulsky; G, C. (C.) gemmata aino Lewis; H, C. (C.) sachaliensis niohozana Bates; I, C. (C.) lewisii lewisii Bates; J, Cylindera (Eugrapha) elisae elisae (Motschulsky); K, Cicindela (Cicindela) transbaicalica japanensis Chaudoir. Scales: 10 mm.

FIGURE 15. Everted internal sacs of Cicindela (Cicindela). A, Cicindela (Cicindela) japana japana Motschulsky; B, C. (C.) transbaicalica japanensis Chaudoir; C, (C.) sachalinensis niohozana Bates; D, C. (C.) lewisii lewisii Bates; E, C. (C.) gemmata aino Lewis. Scales: 1.0 mm.