Cosmodela, : Naviaux, 1991

Fukuda, Yuki, Ogawa, Ryo & Hori, Michio, 2019, The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia, Zootaxa 4661 (2), pp. 271-308 : 287

Cosmodela sp. cf. separata

( Figs 8A , 13A )

Specimens examined. (N = 1) [ Vietnam] 1♂ ( MHJ), Lao Cai Prov., Sapa , 20-23. V. 2012, M. Hori leg.

Description. Internal sac everted widely and dorsally in lateral view. DA largest in bladders; AL the second largest; B same as large as VA, A, DLL, VLL and VLR.

A thick and cylindrically everted apically. AL roundly right ventro-laterally swollen, vested with fine spines. Medial tooth short, extending ventrally, weakly curved at apex. DLL roundly swollen, vested with fine spines. VA short triangular swollen, everted dorsa-apically. VLR oval and weakly swollen left laterally at base of VA. VLL widely swollen right laterally, with well sclerotized spines, which are forming a narrow band. B slightly and roundly swollen, everted right laterally, numerously with long spines. Flagellum very long, curved and formed S-shaped. Medial tooth short, extending ventrally, weakly curved at apex.

Remarks. This species is clearly distinguished from other members of Cosmodela by the following characteristics: A and VA short; VB non-existent; AL everted ventro-laterally.

Figures

FIGURE 8. Habitus of Cosmodela, Cicindela (Cicindela) and Cylindera.A, Cosmodela sp. (cf. separata); B, C. batesi (Fleutiaux) from Iriomote Is., Japan; C, C. aurulenta (Fabricius) from Penang Is., Malaysia; D, C. virgula (Fleutiaux) from Laos; E, C. duponti duponti (Dejean) from South India; F, Cicindela (Cicindela) japana japana Motschulsky; G, C. (C.) gemmata aino Lewis; H, C. (C.) sachaliensis niohozana Bates; I, C. (C.) lewisii lewisii Bates; J, Cylindera (Eugrapha) elisae elisae (Motschulsky); K, Cicindela (Cicindela) transbaicalica japanensis Chaudoir. Scales: 10 mm.

FIGURE 13. Everted internal sacs of Cosmodela. A, Cosmodela sp. (cf. separata) from Vietnam; B, C. aurulenta (Fabricius) from Penang Is., Malaysia; C, C. virgula (Fleutiaux) from Laos; D, C. duponti duponti (Dejean) from South India. d1, d2, d3, d4, show the connecting of flagellum. Scales: 1.0 mm.