Solanum tuberosum L.

OVCHINNIKOVA, ANNA, KRYLOVA, EKATERINA, GAVRILENKO, TATJANA, SMEKALOVA, TAMARA, ZHUK, MIKHAIL, KNAPP, SANDRA & SPOONER, DAVID M., 2011, Taxonomy of cultivated potatoes (Solanum section Petota: Solanaceae), Botanical Journal of the Linnean Society 165 (2), pp. 107-155 : 121-139

4. Solanum tuberosum L. Sp. Pl. 185. 1753.

Battata tuberosa Hill , Hort. Kew. 148. 1768, Lycopersicon tuberosum (L.) Mill., Gard. Dict. ed. 8, no. 7. 1768, Solanum esculentum Neck ., Delic. Gallo-Belg. 1: 119. 1768, Solanum tuberosum L. var. vulgare Hook. f., Bot. Antarct. Voy. I. (Fl. Antarct.) 2: 329. 1846, Solanum tuberosum L. var. cultum A.DC. Arch. Sci. Phys. Nat. ser. 3, 15: 437. 1886, Solanum tuberosum L. var.? vulgare Macloskie, Rep. Princeton Univ. Exped. Patagonia 8: 707. 1905, Solanum cultum (A.DC.) Berthault, Recherc. Bot. var. Cult. Solanum tuberosum , etc. 127, 128. 1911.

Type: cultivated in Uppsala, Anon . (lectotype, LINN! [ LINN 248.12 , BH neg. 6799 ], designated by Hawkes, 1956b: 106). [C] Figure 4 .

Solanum sinense Blanco, Fl. Filip. ed. 1: 137. 1837.

Type: Philippines. Sin. loc. cultivated, F. Blanco s.n. (no specimens found). [C].

Solanum tuberosum L. var. leonhardianum Alef., Landw. Fl. 137. 1866.

Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. melanoceras Alef., Landw. Fl. 137. 1866.

Type: cultivated in Darmstadt, Germany (no specimens found, lectotype, Putsch, 1819, Monographie der Kartoffel, f. 29 , designated here). [C ].

Solanum tuberosum L. var. californicum Alef., Landw. Fl. 138. 1866.

Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. erythroceras Alef., Landw. Fl. 138. 1866.

Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. holsaticum Alef., Landw. Fl. 138. 1866.

Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. cucumerinum Alef., Landw. Fl. 139. 1866.

Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. julianum Alef., Landw. Fl. 139. 1866.

Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. menapianum Alef., Landw. Fl. 139. 1866.

Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. platyceras Alef., Landw. Fl. 139. 1866.

Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. reniforme Alef ., Landw. Fl. 139. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. tener Alef ., Landw. Fl. 139. 1866. Type: cultivated in Darmstadt, Germany (no specimens found).

Solanum tuberosum L. var. brachyceras Alef ., Landw. Fl. 140. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. helenanum Alef. , Landw. Fl. 140. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. laurentianum Alef ., Landw. Fl. 140. 1866. Type: no specimens found (lectotype, Putsch, 1819, Monographie der Kartoffel, f. 2, designated here). [C].

Solanum tuberosum L. var. putscheanum Alef ., Landw. Fl. 140. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. sesquimensale Alef ., Landw. Fl. 140. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. xanthoceras Alef. , Landw. Fl. 140. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. bertuchii Alef ., Landw. Fl. 141. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. saccharatum Alef ., Landw. Fl. 141. 1866. Type: cultivated in Darmstadt, Germany (no specimens found, lectotype, Putsch, 1819, Monographie der Kartoffel, f. 19 , designated here). [C].

Solanum tuberosum L. var. schnittspahnii Alef ., Landw. Fl. 141. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. strobilinum Alef ., Landw. Fl. 141. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. borsdorfianum Alef ., Landw. Fl. 142. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. cordiforme Alef ., Landw. Fl. 142. 1866. Type: cultivated in Darmstadt, Germany (no specimens found, lectotype, Putsch, 1819, Monographie der Kartoffel, f. 3, designated here). [C] .

Solanum tuberosum L. var. nobile Alef. , Landw. Fl. 142. 1866. Type: cultivated in Darmstadt, Germany (no specimens found, lectotype, Putsch, 1819, Monographie der Kartoffel, f. 10 , designated here). [C ].

Solanum tuberosum L. var. album Alef ., Landw. Fl. 143. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. batatinum Alef ., Landw. Fl. 143. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. nucinum Alef ., Landw. Fl. 143. 1866. Type: no specimens found (lectotype, Putsch, 1819, Monographie der Kartoffel, f. 21 , designated here). [C] .

Solanum tuberosum L. var. peruvianum Alef ., Landw. Fl. 143. 1866. Type: cultivated in Darmstadt, Germany (no specimens found, lectotype, Putsch, 1819, Monographie der Kartoffel, f. 23 , designated here). [C].

Solanum tuberosum L. var. alaudinum Alef ., Landw. Fl. 144. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. anglicum Alef ., Landw. Fl. 144. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. bufoninum Alef ., Landw. Fl. 144. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. hispanicum Alef ., Landw. Fl. 144. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. pecorum Alef ., Landw. Fl. 144. 1866. Type: cultivated in Darmstadt, Germany (no specimens found, lectotype, Putsch, 1819, Monographie der Kartoffel, f. 31, designated here). [C].

Solanum tuberosum L. var. rugiorum Alef ., Landw. Fl. 144. 1866. Type: cultivated in Darmstadt, Germany (no specimens found, lectotype, Putsch, 1819, Monographie der Kartoffel, f. 30, designated here). [C] .

Solanum tuberosum L. var. conocarpum Alef ., Landw. Fl. 145. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. praecox Alef. , Landw. Fl. 145. 1866. Type: cultivated in Darmstadt, Germany (no specimens found, lectotype, Putsch, 1819, Monographie der Kartoffel, f. 6, designated here). [C].

Solanum tuberosum L. var. praedicandum Alef ., Landw. Fl. 145. 1866. Type: no specimens found (lectotype, Putsch, 1819, Monographie der Kartoffel, f. 26, designated here).

Solanum tuberosum L. var. rockii Alef ., Landw. Fl. 145. 1866. Type: cultivated in Darmstadt, Germany (no specimens found, lectotype, Putsch, 1819, Monographie der Kartoffel, f. 9 , designated here). [C].

Solanum tuberosum L. var. drakeanum Alef. , Landw. Fl. 146. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. elongatum Alef ., Landw. Fl. 146. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. tinctorium Alef ., Landw. Fl. 146. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. vuchefeldicum Alef ., Landw. Fl. 146. 1866. Type: cultivated in Darmstadt, Germany (no specimens found, lectotype, Putsch, 1819, Monographie der Kartoffel, f. 27 , designated here). [C].

Solanum tuberosum L. var. aethiopicum Alef ., Landw. Fl. 147. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. hassicum Alef., Landw. Fl. 147. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. kaunitzii Alef ., Landw. Fl. 147. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. merceri Alef ., Landw. Fl. 147. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. norfolcicum Alef ., Landw. Fl. 147. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. rossicum Alef ., Landw. Fl. 147. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. ulmense Alef ., Landw. Fl. 147. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. cepinum Alef ., Landw. Fl. 148. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. fragariinum Alef ., Landw. Fl. 148. 1866. Solanum tuberosum L. convar. fragariinum (Alef.) Danert , Kulturpβanze 4: 126. 1956. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. palatinatum Alef ., Landw. Fl. 148. 1866. Type: cultivated in Darmstadt, Germany (no specimens found, lectotype, Putsch, 1819, Monographie der Kartoffel, f. 5 , designated here). [C] .

Solanum tuberosum L. var. versicolor Alef ., Landw. Fl. 148. 1866. Type: cultivated in Darmstadt, Germany (no specimens found, lectotype, Putsch, 1819, Monographie der Kartoffel, f. 7 , designated here). [C] .

Solanum tuberosum L. var. corsicanum Alef ., Landw. Fl. 149. 1866. Type: cultivated in Darmstadt, Germany (no specimens found). [C].

Solanum tuberosum L. var. salamandrinum Alef ., Landw. Fl. 149. 1866. Type: cultivated in Darmstadt, Germany (no specimens found, lectotype, Putsch, 1819, Monographie der Kartoffel, f. 8 , designated here). [C].

Solanum tuberosum L. var. chiloense A.DC ., Arch. Sci. Phys. Nat. ser. 3, 15: 437. 1886. Solanum chiloense (A.DC.) Berthault , Ann. Sci. Agron. Franç. Étrangère, ser. 3, 6: 180. 1911. Type: Chile. Region X (Los Lagos): Isla Chiloé, 1862, R.A. Philippi s.n. (holotype, G! [ G 00070236 ]). [C].

Solanum maglia Schltdl. var. guaytecarum Bitter, Repert. Spec. Nov Regni Veg. 12: 2. 1913. Solanum tuberosum L. var. guaytecarum (Bitter) Hawkes , Proc. Linn. Soc. Lond. 166: 130. 1956. Type: Chile. Region X (Los Lagos): Prov. Chiloé, Guaytecas Archipelago , 6.iii.1857, N. Funck 102C (holotype, P! [ P 00384639 ]). [C].

Solanum tenuifilamentum Juz. & Bukasov , Trudy Vsesoyuzn. S”ezda Gen. Selekts. Semenov. Plemen. Zhivotnov. 3: 603. 1929. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Chinchero, S. Juzepczuk 1355), viii.1929, S. Juzepczuk [1255] (lectotype, LE!, designated by Ovchinnikova et al., 2009: 586). [A].

Solanum phureja Juz. & Bukasov , Trudy Vsesoyuzn. S”ezda Gen. Selekts. Semenov. Plemen. Zhivotnov. 3: 604. 1929. Type: cultivated in Leningrad from tuber accession 4855/3587 collected in Bolivia (La Paz: Sorata, S. Juzepczuk 1654), 1929, S. Juzepczuk [1654] (lectotype, WIR! [ WIR-36885 ], designated by Korovina , Belozor & Chernomorskaja , 1985: 19 ; isolectotype, K! [ K000658022 ]). [A] .

Solanum goniocalyx Juz. & Bukasov , Trudy Vsesoyuzn. S”ezda Gen. Selekts. Semenov. Plemen. Zhivotnov. 3: 605. 1929. Solanum stenotomum Juz. & Bukasov subsp. goniocalyx (Juz. & Bukasov) Hawkes , Rep. (Annual) Scott. Pl. Breed. Sta. 1963: 157. 1963. Type: cultivated in Leningrad from tuber accession collected in Peru ( Cerro de Pasco, S. Juzepzcuk 571), 1929, S. Juzepczuk [571] (lectotype, LE!, designated by Gorbatenko & Hawkes , 1996: 552 ; isolectotype, LE!). [A].

Solanum rybinii Juz. & Bukasov , Trudy Vsesoyuzn. S”ezda Gen. Selekts. Semenov. Plemen. Zhivotnov. 3: 606. 1929. Type: cultivated in Leningrad from tuber accession collected in Colombia (Cundinamarca: near Bogotá, S. Bukasov 46), ix.1929, S. Juzepczuk s.n. (lectotype, LE!, designated here). [A] .

Solanum stenotomum Juz. & Bukasov , Trudy Vsesoyuzn. S”ezda Gen. Selekts. Semenov. Plemen. Zhivotnov. 3: 609. 1929. Type: cultivated in Leningrad from tuber accession 4870/3537 collected in Bolivia (La Paz, S. Juzepczuk 1681), 1929, S. Juzepczuk [1681] 1552 (lectotype, WIR! [ WIR-18777 ], designated here). [A ].

Solanum andigenum Juz. & Bukasov , Trudy Vsesoyuzn. S”ezda Gen. Selekts. Semenov. Plemen. Zhivotnov. 3: 609. 1929, Solanum tuberosum L. subsp. andigenum (Juz. & Bukasov) Hawkes , Proc. Linn. Soc. Lond. 166: 130. 1956. Type: cultivated in Leningrad from tuber accession collected in Peru (Pasco: Cerro de Pasco, 1927, S. Juzepczuk 598), ix.1929, S. Juzepczuk [598] (lectotype, LE! [unnumbered photograph mounted on K000658023 !], designated here ; isolectotypes, K! [ K000065896 , K000658097 , K000658098 , K000658099 ]). [A ]. Figure 5 .

Solanum boyacense Juz. & Bukasov , Trudy Vsesoyuzn. S”ezda Gen. Selekts. Semenov. Plemen. Zhivotnov. 3: 609. 1929. Solanum rybinii Juz. & Bukasov var. boyacense (Juz. & Bukasov) Hawkes , Potato Collect. Exped. Mexico & S. Amer. 2 Syst. Classific. Collect. 2, 70. 1944. Type: cultivated in Leningrad from tuber accession collected in Colombia (Boyacá: Chochonta, near Tunja , S. Bukasov 20), ix.1929, S. Juzepczuk s.n. (lectotype, LE!, designated here). [A].

Solanum chaucha Juz. & Bukasov , Trudy Vsesoyuzn. S”ezda Gen. Selekts. Semenov. Plemen. Zhivotnov. 3: 609. 1929. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: near Cusco, San Jeronimo, S. Juzepczuk 1010), viii.1929, S. Juzepczuk [1010] (lectotype, LE!, designated by Ovchinnikova et al., 2009: 583 ; isolectotypes, K! [ K000658008 , K000658177 ], LE! [2 additional sheets]). [A].

Solanum mamilliferum Juz. & Bukasov , Trudy Vsesoyuzn. S”ezda Gen. Selekts. Semenov. Plemen. Zhivotnov. 3: 609. 1929. Type: cultivated in Leningrad from tuber accession 4899/3415 collected in Peru (Cusco: Ppisac, S. Juzepczuk 1001), 1928, S. Juzepczuk 1498 (lectotype, WIR! [ WIR-18611 ], designated by Ovchinnikova et al., 2009: 584 ; isolectotype, K! [ K000658176 ]). [A].

Solanum andigenum Juz. & Bukasov var. mexicanum Bukasov , Trudy Prikl. Bot. Suppl. 47: 202, 516. 1930, Solanum andigenum Juz. & Bukasov forma tolucanum Bukasov, Trudy Prikl. Bot. Suppl. 47: 202, 516. 1930. Type: cultivated in Leningrad from tuber accession 4001/1 collected in Mexico ( Mexico: Cuautzingo, Anon . [S. Bukasov] 1), 1929, S. Juzepczuk s.n. (lectotype, WIR! [ WIR-38364 ], designated here). [A].

Solanum andigenum Juz. & Bukasov forma chalcoense Bukasov, Trudy Prikl. Bot. Suppl. 47: 204, 517. 1930. Type: cultivated in Leningrad from tuber accession collected in Mexico (Toluca: Cuatzingo (near Chalco), S. Bukasov 3 & 4; DF: Cocayan and Ajusco, M.C. Antipovich s.n.) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov forma guatemalense Bukasov, Trudy Prikl. Bot. Suppl. 47: 205, 518. 1930. Type: cultivated in Leningrad from tuber accession 4018 collected in Guatemala (Escuintla, Anon . [S. Bukasov] 13), 1929, S. Bukasov [13] (lectotype, WIR! [ WIR-38370 ], designated here) . [A].

Solanum andigenum Juz. & Bukasov var. colombianum Bukasov , Trudy Prikl. Bot. Suppl. 47: 205. 1930, Solanum andigenum Juz. & Bukasov forma tocanum Bukasov, Trudy Prikl. Bot. Suppl. 47: 205. 1930. Type: cultivated in Leningrad from tuber accession collected in Colombia (Cundinamarca, S. Juzepczuk 44), 1930, S. Juzepczuk [44] (lectotype, WIR! [ WIR-38416 ], designated here). [A].

Solanum andigenum Juz. & Bukasov var. cuzcoense Bukasov & Lechn ., Trudy Prikl. Bot. Suppl. 47: 205. 1930. Solanum andigenum Juz. & Bukasov convar. longiacuminatum Lechn ., Trudy Prikl. Bot. 79: 47. 1983. Type: cultivated in Leningrad from tuber accession 4355/3291 collected in Peru (Cusco: Paucartambo, S. Juzepczuk 1348), no date, V.S. Lekhnovich s.n. (lectotype, WIR! [ WIR-37225 ], designated here) . [A].

Solanum andigenum Juz. & Bukasov forma caiceda Bukasov, Trudy Prikl. Bot. Suppl. 47: 208. 1930. Type: cultivated in Leningrad from tuber accession 4039 collected in Colombia (Cundinamarca: Chipaque , Anon . [S. Bukasov] 49b), 1931, S. Juzepczuk [49b] (lectotype, WIR! [ WIR-38498 ], designated here) . [A].

Solanum andigenum Juz. & Bukasov forma lisarassa Bukasov, Trudy Prikl. Bot. Suppl. 47: 209, 520. 1930. Type: cultivated in Leningrad from tuber accession 4083 collected in Colombia (Cundinamarca: near Bogotá, Chipaque , S. Juzepczuk 35), 1929, S. Juzepczuk [35] 1449 (lectotype, WIR! [ WIR-18661 ], designated here). [A].

Solanum andigenum Juz. & Bukasov forma rosada Bukasov, Trudy Prikl. Bot. Suppl. 47: 210, 250, fig. 103. 1930. Type: cultivated in Leningrad from tuber accession 4169 collected in Colombia (Pasto: Popayán, Anon . [S. Bukasov] 98a), 1929, S. Juzepczuk [98a] (lectotype, WIR! [ WIR-38372 ], designated here) . [A].

Solanum andigenum Juz. & Bukasov var. hederiforme Bukasov , Trudy Prikl. Bot. Suppl. 47: 521. 1930. Type: cultivated in Leningrad from tuber accession collected in Colombia (Caldas: Manizales, Anon. [S. Bukasov] 84), 1928, S. Bukasov [84] (lectotype, WIR! [ WIR-38381 ], designated here) . [A].

Solanum stenotomum Juz. & Bukasov var. ccami Bukasov , Trudy Prikl. Bot. Suppl. 58: 33, 164. 1933. Solanum stenotomum Juz. & Bukasov forma ccami (Juz. & Bukasov) Hawkes, Potato Collect. Exped. Mexico & S. Amer., 2, Syst. Classific. Collect. 55. 1944. Type: cultivated in Leningrad from tuber accession 4678/3605 collected in Bolivia (La Paz: Titicaca, Tiahuanaco , S. Juzepczuk 1645), 1929, S. Juzepczuk [1645] (lectotype, WIR!, designated here ). [A].

Solanum stenotomum Juz. & Bukasov var. ppitiquina Bukasov , Trudy Prikl. Bot. Suppl. 58: 33. 1933. Type: cultivated in Leningrad from tuber accession 4862/3446 collected in Peru (Cusco, S. Juzepczuk 1319), 1929, S. Juzepczuk [1319] (lectotype, WIR!, designated here). [A].

Solanum andigenum Juz. & Bukasov forma lilacinoflorum Bukasov, Trudy Prikl. Bot. Suppl. 58: 38. 1933. Type: cultivated in Leningrad from tuber accession 4106 collected in Colombia (Cundinamarca: Chipaque, Anon . [S. Bukasov] 57a), 1929, S. Juzepczuk s.n. (lectotype, WIR! [ WIR-38390 ], designated here) . [A].

Solanum andigenum Juz. & Bukasov subsp. ayapapa Bukasov & Lechn ., Trudy Prikl. Bot. Suppl. 58: 45, 164. 1933. Type: cultivated in Leningrad from tuber accessions collected in Ecuador (sin. loc., no collector cited) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov var. juninum Bukasov , Trudy Prikl. Bot. Suppl. 58: 46. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Junín: sin. loc., S. Juzepczuk 670 & 787) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov forma nodosum Bukasov, Trudy Prikl. Bot. Suppl. 58: 49. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Pasco: Cerro de Pasco market from Titlicaen, S. Juzepczuk 572) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov subsp. tarmense Bukasov & Lechn ., Trudy Prikl. Bot. Suppl. 58: 50. 1933. Solanum andigenum Juz. & Bukasov subsp. tarmense Bukasov & Lechn ., Trudy Prikl. Bot. 79: 46. 1983, nomen superfl. Type: cultivated in Leningrad from tuber accession collected in Peru (Pasco: Cerro de Pasco, 1929, S. Juzepczuk 587b), 1929, V. S. Lekhnovich [4320] (lectotype, WIR!, designated by Lekhnovich , 1983: 46). [A].

Solanum andigenum Juz. & Bukasov var. platyantherum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 50. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Junín: Carhuamayo, 4100 m, S. Juzepczuk 732) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov forma tenue Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 51. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Pasco: Chinchao, S. Juzepczuk 587 & 603) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov var. latius Bukasov & Lechn ., Trudy Prikl. Bot. Suppl. 58: 51. 1933. Type: sin. loc., Anon 179a? (no specimens or collector cited). [A].

Solanum andigenum Juz. & Bukasov var. ckellohuaccoto Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 53, fig. 12. 1933. Type: cultivated in Leningrad from tuber accession 4894/3324 collected in Peru (Cusco: Yuncaypampa , S. Juzepczuk 1228), 1929, S. Juzepczuk [1228] (lectotype, WIR! [ WIR-39571 ], designated here) . [A].

Solanum andigenum Juz. & Bukasov forma alccaihuarmi Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 54, fig. 13. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Ccorau , S. Juzepczuk 1390), 1928, S. Juzepczuk [1390] (lectotype, WIR! [ WIR-37218 ], designated here). [A].

Solanum andigenum Juz. & Bukasov forma huamanuma Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 54, tab. 2, fig. 6. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (‘south Peru group’, S. Juzepczuk 1138) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov forma sunchchu Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 55. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Poroy , S. Juzepczuk 1397), 1928, S. Juzepczuk [1397] (lectotype, WIR! [ WIR-37267 ], designated here). [A].

Solanum andigenum Juz. & Bukasov forma ccompetillo Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 56. 1933. Type: cultivated in Leningrad from tuber accession 4479/3481 collected in Peru (Cusco: San Jeronimo , S. Juzepczuk 1352), 1929, S. Juzepczuk [1352] (lectotype, WIR! [ WIR-37523 ], designated here). [A].

Solanum andigenum Juz. & Bukasov var. huairuru Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 57, tab. 2, fig. 4. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Yuncaypampa, S. Juzepczuk 1231), 1928, S. Juzepczuk [1231] (lectotype, WIR! [ WIR-37376 ], designated here). [A].

Solanum andigenum Juz. & Bukasov forma pacus Lechn. ex Bukasov, Trudy Prikl. Bot. Suppl. 58: 57. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Acomayo region, Pomacanchi, S. Juzepczuk 1174 & 1177) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov var. quechuanum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 57. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Jamamachay , S. Juzepczuk 1009a), 1931, S. Juzepczuk [1009a] (lectotype, WIR! [ WIR-37494 ], designated here). [A].

Solanum andigenum Juz. & Bukasov forma chimaco Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 58. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Urubamba, Ollantaytambo , S. Juzepczuk 1064), 1931, S. Juzepczuk [1064] (lectotype, WIR! [ WIR-37468 ], designated here). [A].

Solanum andigenum Juz. & Bukasov var. sihuanum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 58. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Omasbamba , S. Juzepczuk 1256), 1929, S. Juzepczuk [1259] (lectotype, WIR! [ WIR-37027 ], designated here) . [A].

Solanum andigenum Juz. & Bukasov forma llutucruntum Lechn. ex Bukasov, Trudy Prikl. Bot. Suppl. 58: 60. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Acomayo, Pomacanchi , S. Juzepczuk 1249), 1931, S. Juzepczuk [1249] (lectotype, WIR! [ WIR-3740 ], designated here). [A].

Solanum andigenum Juz. & Bukasov forma ppaqui Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 60. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Poroy, S. Juzepczuk 1402), 1931, S. Juzepczuk [1402] (lectotype, WIR! [ WIR-37822 ], designated here). [A].

Solanum andigenum Juz. & Bukasov var. herrerae Bukasov & Lechn. , Trudy Prikl. Bot. Suppl. 58: 60. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Yana Machay , S. Juzepczuk 1283), 1928, S. Juzepczuk [1283] (lectotype, WIR! [ WIR-37793 ], designated here). [A].

Solanum andigenum Juz. & Bukasov forma huallata Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 61. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Prov. Chinchero, 1927, S. Juzepczuk 1438), 1938, J.G. Hawkes s.n. (no specimens cited in protologue; neotype, K! [ K000658026 ], designated here) . [A].

Solanum andigenum Juz. & Bukasov forma leckeumo Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 61. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Ccorna, S. Juzepczuk 1415 and Paucartambo, S. Juzepczuk 1307) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov var. ccusi Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 62, fig. 15. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Yana , S. Juzepczuk 1284), 1929, S. Juzepczuk [1284] (lectotype, WIR! [ WIR-37608 ], designated here) . [A].

Solanum andigenum Juz. & Bukasov forma uncuna Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 63, tab. 2, fig. 5. 1933. as ‘ uncuña ’. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Acomayo, Pomanchanchi, S. Juzepczuk 1175), 1928, S. Juzepczuk [1175] (lectotype, WIR! [WIR-37427], designated here). [A].

Solanum andigenum Juz. & Bukasov forma ancaccmaquin Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 63. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Ppicacc, Huaypun, S. Juzepczuk 1306), no date, S. Juzepczuk [1306] (lectotype, WIR! [WIR-37560], designated here). [A].

Solanum andigenum Juz. & Bukasov forma microstigma Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 63. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (sin. loc., S. Juzepczuk 1304b) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov forma ppaccnacha Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 63. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (sin. loc., S. Juzepczuk 1442) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov var. soccohuaccoto Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 63. 1933. Type: cultivated in Leningrad from tuber accession 4381/3322 collected in Peru (Cusco, S. Juzepczuk 1227), no date, S. Juzepczuk [1227] (lectotype, WIR! [WIR-37243], designated here). [A].

Solanum andigenum Juz. & Bukasov forma pomacanchicum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 63. 1933. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Acomayo, Pomacanchi, S. Juzepczuk 1178c), 1929, S. Juzepczuk [1178c] (lectotype, WIR! [WIR-37590], designated here). [A].

Solanum andigenum Juz. & Bukasov var. imilla Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 66. 1933. Type: no specimens or localities cited. [A].

Solanum andigenum Juz. & Bukasov forma chiarimilla Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 66. 1933. Type: cultivated in Leningrad from tuber accession collected in Bolivia (La Paz, S. Juzepczuk 1582), 1931, S. Juzepczuk [1582] (lectotype, WIR! [WIR-37880], designated here). [A].

Solanum andigenum Juz. & Bukasov forma ccompis Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 67. 1933. Type: cultivated in Leningrad from tuber accession 4508/3500 collected in Peru (Cusco: Tampumachay, S. Juzepczuk 1289), 1928, S. Juzepczuk [1289] (lectotype, WIR!, designated here). [A].

Solanum andigenum Juz. & Bukasov forma huacalayra Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 68. 1933. Type: cultivated in Leningrad from tuber accession collected in Bolivia (La Paz, S. Juzepczuk 1809a) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov forma ovatibaccatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 68. 1933. Type: cultivated in Leningrad from tuber accessions collected in Bolivia (La Paz, S. Juzepczuk 1809a, 1846) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov var. aymaranum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 68. 1933. Type: cultivated in Leningrad from tuber accessions collected in Bolivia (La Paz: Tihuanaco, Lake Titicaca, S. Juzepczuk 1621 & 1661) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov forma conicicolumnatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 68. 1933. Type: cultivated in Leningrad from tuber accession 4615/3545 collected in Bolivia (La Paz, S. Juzepczuk 1521), 1931, S. Juzepczuk [1521] (lectotype, WIR!, designated here). [A].

Solanum andigenum Juz. & Bukasov forma globosum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 69, fig. 17. 1933. Type: cultivated in Leningrad from tuber accession 4621/3547 collected in Bolivia (La Paz, S. Juzepczuk 1583), 1929, S. Juzepczuk [1583] (lectotype, WIR! [WIR-39474], designated here). [A].

Solanum andigenum Juz. & Bukasov forma discolor Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 69. 1933. Type: cultivated in Leningrad from tuber accession collected in Bolivia (La Paz, S. Juzepczuk 1663) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov forma lanciacuminatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 69. 1933. Type: cultivated in Leningrad from tuber accessions collected in Bolivia (La Paz, S. Juzepczuk 1735) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov forma basiscopum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 69. 1933. Type: cultivated in Leningrad from tuber accession 4879/3565 collected in Bolivia (La Paz, S. Juzepczuk 1740), 1929, V.S. Lekhnovich 1428 (lectotype, WIR!, designated here). [A].

Solanum andigenum Juz. & Bukasov forma lapazense Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 70, fig. 18. 1933. Type: cultivated in Leningrad from tuber accession 4662/3260 collected in Bolivia (La Paz: Coni, S. Juzepczuk 1812), 1929, S. Juzepczuk 1394 [1812] (lectotype, WIR! [WIR-18771], designated here). [A].

Solanum andigenum Juz. & Bukasov var. bolivianum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 70, fig. 19. 1933. Type: cultivated in Leningrad from tuber accession 4643/3589 collected in Bolivia (La Paz, S. Juzepczuk 1656), 1929, S. Juzepczuk 1424 [1656] (lectotype, WIR! [WIR-18756], designated here). [A].

Solanum andigenum Juz. & Bukasov forma bifidum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 71. 1933. Type: cultivated in Leningrad from tuber accession 4618/3546 collected in Bolivia (La Paz, S. Juzepczuk 1783), 1931, S. Juzepczuk [1783] (lectotype, WIR!, designated here). [A].

Solanum andigenum Juz. & Bukasov forma incrassatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 71. 1933. Type: cultivated in Leningrad from tuber accession 4618/3546 collected in Bolivia (La Paz, S. Juzepczuk 1583b), 1931, S. Juzepczuk [1583b] (lectotype, WIR! [WIR-1421], designated here). [A].

Solanum andigenum Juz. & Bukasov forma magnicorollatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 71. 1933. Type: cultivated in Leningrad from tuber accession 4653/3575 collected in Bolivia (La Paz, S. Juzepczuk 1811), 1929, S. Juzepczuk [1811] (lectotype, WIR!, designated here). [A].

Solanum andigenum Juz. & Bukasov forma dilatatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 72, fig. 20. 1933. Type: cultivated in Leningrad from tuber accession 4602/3520 collected in Bolivia (La Paz, S. Juzepczuk 1522b), 1931, S. Juzepczuk [1522b] (lectotype, WIR! [WIR-38393], designated here). [A].

Solanum andigenum Juz. & Bukasov forma tiahuanacense Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 72. 1933. Type: cultivated in Leningrad from tuber accession 4669/3599 collected in Bolivia (La Paz: Titicaca, Tiahuanaco, S. Juzepczuk 1640), 1929, S. Juzepczuk 1419 [1640] (lectotype, WIR! [WIR-18779], designated here). [A].

Solanum andigenum Juz. & Bukasov var. longibaccatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 73, fig. 21. 1933. Type: cultivated in Leningrad from tuber accession 4683/3523 collected in Bolivia (La Paz, S. Juzepczuk 1678), 1929, S. Juzepczuk 1400 [1678] (lectotype, WIR! [WIR-18732], designated here).

Solanum andigenum Juz. & Bukasov forma arcuatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 73. 1933. Type: cultivated in Leningrad from tuber accession collected in Bolivia (La Paz, S. Juzepczuk 1522a) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov forma caesium Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 74. 1933. Type: cultivated in Leningrad from tuber accession 4601/3520 collected in Bolivia (La Paz, S. Juzepczuk 1501), 1929, S. Juzepczuk 1395 [1501] (lectotype, WIR! [WIR-18773], designated here). [A].

Solanum andigenum Juz. & Bukasov forma cevallosii Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 74. 1933. Type: cultivated in Leningrad from tuber accession 4613/3543 collected in Bolivia (La Paz, S. Juzepczuk 1659), 1929, S. Juzepczuk 1426 [1659] (lectotype, WIR! [WIR-18764], designated here). [A].

Solanum andigenum Juz. & Bukasov forma pallidum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 74. 1933. Type: cultivated in Leningrad from tuber accession 4612/3541 collected in Bolivia (La Paz, S. Juzepczuk 1525), 1929, S. Juzepczuk 1415 [1525] (lectotype, WIR! [WIR-18758], designated here). [A].

Solanum andigenum Juz. & Bukasov var. stenophyllum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 74. 1933. Type: cultivated in Leningrad from tuber accession collected in Bolivia (La Paz, sin. loc., S. Juzepczuk 1682a) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov var. brevicalyx Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 75, fig. 22. 1933. Type: cultivated in Leningrad from tuber accession 4622/3548 collected in Bolivia (La Paz, S. Juzepczuk 1663b), 1929, S. Juzepczuk [1663b] (lectotype, WIR!, designated here). [A].

Solanum andigenum Juz. & Bukasov forma cryptostylum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 75. 1933. Type: cultivated in Leningrad from tuber accession 4660/3582 collected in Bolivia (La Paz, S. Juzepczuk 1536), 1929, S. Juzepczuk 1429 [1536] (lectotype, WIR! [WIR-18728], designated here). [A].

Solanum tuberosum L. var. chilotanum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 80, fig. 23. 1933. Solanum chilotanum (Bukasov & Lechn.) Hawkes, Potato Collect. Exped. Mexico & S. Amer., 2, Syst. Classif. Collect. 79. 1944, as nom. provis. Type: cultivated in Leningrad from tuber accession collected in Chile (Chiloé Island, Ancud, S. Juzepczuk 1938), 1930, S. Juzepczuk [1938] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma camota Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 81. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, S. Juzepczuk 1954) (no specimens found). [C].

Solanum tuberosum L. forma chaped Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 81. 1933. Type: cultivated in Leningrad from tuber accession 4737 collected in Chile (Ancud, S. Juzepczuk 1944), 1929, S. Juzepczuk [1944] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma indianum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 81. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, S. Juzepczuk 1934) (no specimens found). [C].

Solanum tuberosum L. forma viride Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 81. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy Island, S. Juzepczuk 1962), 1931, S. Juzepczuk [1962] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma mahuinhue Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 82. 1933. Type: cultivated in Leningrad from tuber accession 4823 collected in Chile (San Sebastiano, S. Juzepczuk 2038), 1929, S. Juzepczuk [2038] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma pillicuma Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 82. 1933. Type: cultivated in Leningrad from tuber accession 4778 collected in Chile (Yutuy, S. Juzepczuk 1986), 1929, S. Juzepczuk [1986] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma acuminatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 83. 1933. Type: cultivated in Leningrad from tuber accession 4806 collected in Chile (Chiloé, Mechiaco, S. Juzepczuk 2020), 1929, S. Juzepczuk [2020] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma barmacata Bukasov & Lechn ., Trudy Prikl. Bot. Suppl. 58: 83. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, S. Juzepczuk 1942) (no specimens found). [C].

Solanum tuberosum L. forma brachykalukon Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 83. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, Aguatao, S. Juzepczuk 2002) (no specimens found). [C].

Solanum tuberosum L. forma thalassinum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 83. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Chiloé Island, Mechiaco and Quetalhahue, S. Juzepczuk 2019, 2025a) (no specimens found). [C].

Solanum tuberosum L. forma oculosum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 83. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Puerto Montt, S. Juzepczuk 1919) (no specimens found). [C].

Solanum tuberosum L. forma coriala Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 84. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Santiago to Puerto Montt, Ancud, S. Juzepczuk 1834, 1924) (no specimens found). [C].

Solanum tuberosum L. forma enode Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 84. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, S. Juzepczuk 1932) (no specimens found). [C].

Solanum tuberosum L. forma montticum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 84. 1933. Type: cultivated from tuber accession 4724 collected in Chile / Argentina (Puerto Montt, S. Juzepczuk 1930), 1929, S. Juzepczuk [1930] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma tenuipedicellatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 84. 1933. Type: cultivated from tuber accession 4704 collected in Chile (Santiago, S. Juzepczuk 1840b), 1929, S. Juzepczuk [1840b] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma brevipapillosum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 84. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Santiago, S. Juzepczuk 1835) (no specimens found). [C].

Solanum tuberosum L. var. elegans Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 85. 1933. Type: cultivated in Leningrad from tuber accession 4788 collected in Chile (Yutuy, S. Juzepczuk 1993), 1929, S. Juzepczuk [1993] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma conicum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 86. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, S. Juzepczuk 2000a) (no specimens found). [C].

Solanum tuberosum L. forma latum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 86. 1933. Type: cultivated in Leningrad from tuber accession 4792 collected in Chile (Yutuy, S. Juzepczuk 2006), 1929, S. Juzepczuk [2006] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. var. villaroella Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 86. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Chiloé Island, Mechiaco, S. Juzepczuk 2021), 1929, S. Juzepczuk [2021] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma araucanum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 87. 1933. Type: cultivated in Leningrad from tuber accession 4765 collected in Chile (Chiloé, Yutuy, S. Juzepczuk 1975), 1929, S. Juzepczuk [1975] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. var. recurvatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 87. 1933. Type: cultivated in Leningrad from tuber accession 4800 collected in Chile (Yutuy, Aguatao, S. Juzepczuk 2005), 1929, S. Juzepczuk [2005] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. var. multibaccatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 87. 1933. Type: cultivated in Leningrad from tuber accession 4734 collected in Chile (Ancud, S. Juzepczuk 1940), 1929, S. Juzepczuk [1940] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma elongatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 88. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, S. Juzepczuk 1964) (no specimens found). [C].

Solanum tuberosum L. forma ovatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 88. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, Quetalhahue, S. Juzepczuk 1985) (no specimens found). [C].

Solanum tuberosum L. var. brevipilosum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 88. 1933. Type: cultivated in Leningrad from tuber accession 4749 collected in Chile (Castro, S. Juzepczuk 2010), 1929, S. Juzepczuk [2010] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma contortum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 89. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, S. Juzepczuk 2040) (no specimens found). [C].

Solanum tuberosum L. forma yutuense Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 89. 1933. Type: cultivated from tuber accession 4789 collected in Chile (Yutuy, S. Juzepczuk 1994), 1929, S. Juzepczuk [1994] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. var. crassipedicellatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 89. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, S. Juzepczuk 1970) (no specimens found). [C].

Solanum tuberosum L. var. rubrisuturatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 89. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, Coleo, S. Juzepczuk 1971), 1929, S. Juzepczuk [1971] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma sebastianum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 90. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (San Sebastiano Island near Chiloé, S. Juzepczuk 2034), 1929, S. Juzepczuk [2034] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma infectum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 90. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Chiloé, S. Juzepczuk 2022), 1929, S. Juzepczuk [2022] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma pichunha Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 91. 1933. as ‘ pichuña ’. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, S. Juzepczuk 1996), 1930, S. Juzepczuk [1996] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma auriculatum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 92. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, S. Juzepczuk 2004) (no specimens found). [C].

Solanum tuberosum L. forma crassifilamentum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 92. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Yutuy, S. Juzepczuk 1980), 1931, S. Juzepczuk [1980] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma obliquum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 92. 1933. Type: cultivated in Leningrad from tuber accession 4786 collected in Chile (Yutuy, S. Juzepczuk 1991), 1929, S. Juzepczuk [1991] (lectotype, WIR!, designated here). [C].

Solanum tuberosum L. forma roseum Bukasov & Lechn., Trudy Prikl. Bot. Suppl. 58: 93. 1933. Type: cultivated in Leningrad from tuber accession collected in Chile (Chiloé Island, Ancud, Castro, San Sebastiano Island, S. Juzepczuk 1960, 7863) (no specimens found). [C].

Solanum herrerae Juz., Izv. Akad. Nauk SSSR, Ser. Biol. 2: 310. 1937. Type: Peru. Cusco: Kcaira, Granja del Escuela de Cusco, 18.xi.1932, Kaufmann s.n. (no specimens located, probably grown from cultivated material in Leningrad). [A].

Solanum molinae Juz., Izv. Akad. Nauk SSSR, Ser. Biol. 2: 308. 1937. Type: cultivated in Leningrad from tuber accession collected in Chile ( Isla Chiloé, iv.1928, R. Christie s.n.), 5.ix.1935, S. Juzepczuk 2014 (no specimens found). [C].

Solanum andigenum Juz. & Bukasov var. vallejosii Lechn., Fl. Fauna Peruanas 1: 16. 1940. as ‘ valeejosii ’ an obvious typographical error for vallejosii. Type: cultivated in Leningrad from tuber accession collected in Peru (Ancash: Huaraz, 1927, P.S. Vallejos s.n.), viii.1938, J.G. Hawkes s.n. (no specimens found; neotype, K! [K000658041], designated here; isoneotype, K! [K000658042]). [A].

Solanum andigenum Juz. & Bukasov var. acutifoliatum Lechn., Fl. Fauna Peruanas 1: 17. 1940. Type: cultivated in Leningrad from tuber accessions collected in Peru (Ancash, P.S. Vallejos K7, K11) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov forma elongatipedicellatum Lechn., Fl. Fauna Peruanas 1: 17. 1940. Type: cultivated in Leningrad from tuber accession collected in Peru (Ancash: Huaraz, P.S. Vallejos s.n.) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov forma huarasicum Lechn., Fl. Fauna Peruanas 1: 17. 1940. Type: cultivated in Leningrad from tuber accession collected in Peru (P.S. Vallejos K9) (no specimens found). [A].

Solanum andigenum Juz. & Bukasov var. ancashicum Lechn., Fl. Fauna Peruanas 1: 18. 1940. Type: cultivated in Leningrad from tuber accession collected in Peru (Ancash: Huaraz, P.S. Vallejos K14) (no specimens found). [A].

Solanum subandigenum Hawkes, Potato Collect. Exped. Mexico & S. Amer., 2, Syst. Classific. Collect. 53, 128. 1944. Type: Bolivia. Chuquisaca: Prov. Sucre, on the outskirts of Sucre above the city, 6.iii.1939, E.K. Balls B.1646 (lectotype, BM! [BM000935105], designated here). [A].

Solanum churuspi Hawkes, Potato Collect. Exped. Mexico & S. Amer., 2, Syst. Classific. Collect. 64, 129. 1944. Type: cultivated in Cambridge ( UK) from tuber accession EPC-675 collected in Peru (Cusco: Paucartambo, 1939, L.A. Yabar 25), 27.viii.1941, Anon. [J.G. Hawkes] s.n. (lectotype, K! [K000658003], designated here). [A].

Solanum yabari Hawkes, Potato Collect. Exped. Mexico & S. Amer., 2, Syst. Classific. Collect. 65, 129. 1944. Type: cultivated in Cambridge ( UK) from tuber accession EPC-698 collected in Peru (Cusco: Prov. Paucartambo, 1939, L.A. Yabar 44), 17.viii.1942, Anon. [J.G. Hawkes] s.n. (lectotype, K! [K000658002], designated here). [A].

Solanum cardenasii Hawkes, Potato Collect. Exped. Mexico & S. Amer., 2, Syst. Classific. Collect. 67, 129. 1944. Type: cultivated in Cambridge ( UK) from tuber accession EPC-284 collected in Bolivia (Cochabamba: near Cochabamba, 16.iii.1939, E.K. Balls 6399), 25.v.1944, Anon. [J.G. Hawkes] s.n. (lectotype, K! [K000658021], designated here) [βowers only]. [A].

Solanum phureja Juz. & Bukasov var. pujeri Hawkes, Potato Collect. Exped. Mexico & S. Amer., 2, Syst. Classific. Collect. 68, 130. 1944. Type: cultivated in Cambridge ( UK) from tuber accession collected in Bolivia (La Paz: Prov. Omasuyos: Finca de Huatajata c. 12 500 ft, 14.iv.1939, E.K. Balls 6510), 22.ix.1944, Anon. [J.G. Hawkes] s.n. (lectotype, K! [K000658019]). [A].

Solanum ascasabii Hawkes, Potato Collect. Exped. Mexico & S. Amer., 2, Syst. Classific. Collect. 69, 130. 1944. Type: cultivated in Cambridge ( UK) from tuber accession collected in Ecuador (Napo: Oyacachi, 18.vii.1939, L. Ascasabi s.n.), 16.viii.1941 Anon. [J.G. Hawkes] s.n. (lectotype, K! [K000658013]; isolectotypes K! [K000658014, K000658015]). [A].

Solanum rybinii Juz. & Bukasov var. popayanum Hawkes, Potato Collect. Exped. Mexico & S. Amer., 2, Syst. Classific. Collect. 71, 130. 1944. Type: cultivated in Cambridge ( UK) from tuber accession EPC-546 collected in Colombia (Cauca: market in Popayán, A.H.G. Alston s.n.), 16.viii.1941, Anon. [J.G. Hawkes] s.n. (lectotype, K! [K000658016], designated here; isolectotype, K! [K000658174]; additional βowering material from same tuber accession collected 1946 K! [K000658017, K000658018]). [A].

Solanum rybinii Juz. & Bukasov var. pastoense Hawkes, Potato Collect. Exped. Mexico & S. Amer., 2, Syst. Classific. Collect. 71, 131. 1944. Type: cultivated at the John Innes Institute in Norwich from tuber accession EPC-979 collected in Colombia (Nariño: Pasto, E.K. Balls 7475 [EPC 979]), 1961, Anon. s.n. (neotype, K! [K000658020], designated here). [A].

Solanum chaucha Juz. & Bukasov forma purpureum Hawkes, Potato Collect. Exped. Mexico & S. Amer., 2, Syst. Classific. Collect. 75, 131. 1944. Type: cultivated in Cambridge ( UK) from tuber accession EPC-1071 collected in Peru (Puno: Prov. Puno, J. Soukup s.n.), 19.viii.1942, Anon. [J.G. Hawkes] s.n. (lectotype, K! [K000658009], designated here). [A].

Solanum coeruleiflorum Hawkes, Potato Collect. Exped. Mexico & S. Amer., 2, Syst. Classific. Collect. 76, 131. 1944. Type: cultivated in Cambridge ( UK) from tuber accession EPC-412 collected in Peru (Ayacucho: Ayacucho market, 25.v.1939, E.K. Balls 6941), 17.viii.1942, Anon. [J.G. Hawkes] s.n. (lectotype, K! [K000658007], designated here; isolectotype, K! [K000658008]). [A].

Solanum stenotomum Juz. & Bukasov forma canasense Vargas, Papas Sudper. 2: 7. 1956 [‘1954’]. Type: Peru. Cusco: Prov. Canas, Checka, 3570 m, C. Vargas 136 (holotype, CUZ!). [A].

Solanum stenotomum Juz. & Bukasov forma puca Vargas, Papas Sudper. 2: 7. 1956 [‘1954’]. Type: Peru. Apurímac: Prov. Grau, Hacienda Récord, 3650 m, C. Vargas 92 (holotype, CUZ!). [A].

Solanum stenotomum Juz. & Bukasov forma yurrac Vargas, Papas Sudper. 2: 7. 1956 [‘1954’]. Type: Peru. Apurímac: Prov. Andahuaylas, 3550 m, C. Vargas 76 (holotype, CUZ!). [A].

Solanum stenotomum Juz. & Bukasov var. poccoya Vargas, Papas Sudper. 2: 7. 1956 [‘1954’]. Type: Peru. Apurímac: Prov. Andahuaylas, 3550 m, C. Vargas 76 (holotype, CUZ!). [A].

Solanum andigenum Juz. & Bukasov var. carhua Vargas, Papas Sudper. 2: 49, fig. 5. 1956 [‘1954’]. Type: Peru. Apurimac: Prov. Grau, Maras, 3600 m, C. Vargas 842, 848 (no specimens found). [A].

Solanum andigenum Juz. & Bukasov var. digitotuberosum Vargas, Papas Sudper. 2: 49, fig. 4. 1956 [‘1954’] as ‘ digitotubatrosum ’. Type: Peru. Apurimac: Prov. Andahuaylas, 3050 m, C. Vargas 77 (no specimens found). [A].

Solanum apurimacense Vargas, Papas Sudper. 2: 58. 1956 [‘1954’]. Type: Peru. Apurímac: Prov. Grau, Pamparackay, 3150 m, Papas Cult. 811 (holotype, CUZ!). [A].

Solanum tuberosum L. var. luteocarnosum Danert , Kulturpβanze 4: 123. 1956. Type: ‘Parnassia’ cultivated in Gatersleben Nr. 21118/55, origin Gross Lüsewitz (Knehden), 21.vii.1955, Anon. (lectotype, GAT! [GAT-7102], designated here; tuber collection from same accession collected 23.ix.1955, GAT! [GAT-7116]).

Solanum tuberosum L. var. albiflorum Danert , Kulturpβanze 4: 124. 1956. Type: ‘Wekaragis’ cultivated in Gatersleben Nr. 20959/55, origin Gross Lüsewitz (Gross Lüsewitz), 21.vii.1955, Anon. (lectotype, GAT! [GAT-7091], designated here).

Solanum tuberosum L. var. utillissimum Danert , Kulturpβanze 4: 124. 1956. Type: ‘Primula’ Gross cultivated in Gatersleben Nr. 20853/55, origin Gross Lüsewitz (Gross Lüsewitz), 20.vii.1955, Anon. (lectotype, GAT! [GAT-7079], designated here).

Solanum tuberosum L. var. alatum Danert , Kulturpβanze 4: 124. 1956. Type: ‘Urtica’ cultivated in Gatersleben Nr. 21116/55, origin Gross Lüsewitz, 21.vii.1955, Anon. (lectotype, GAT! [GAT-7101], designated here; tuber collection from same accession collected 23.ix.1955, GAT! [GAT-7119]).

Solanum tuberosum L. var. lutescens Danert , Kulturpβanze 4: 124. 1956. Type: ‘Fichtelgold’ cultivated in Gatersleben Nr. 20986/55, origin Gross Lüsewitz, 21.vii.1955, Anon. (lectotype, GAT! [GAT-7095], designated here).

Solanum tuberosum L. var. rubrum Danert , Kulturpβanze 4: 124. 1956. Type: ‘Sickingen’ cultivated in Gatersleben Nr. 20970/55, origin Gross Lüsewitz (Wentow), 21.vii.1955, Anon. (lectotype, GAT! [GAT-7093], designated here).

Solanum tuberosum L. var. caesium Danert , Kulturpβanze 4: 125. 1956. Type: no plant material cited (no specimens cited).

Solanum tuberosum L. var. aflorum Danert , Kulturpβanze 4: 125. 1956. Type: no plant material cited (no specimens cited).

Solanum tuberosum L. var. violaceum Danert , Kulturpβanze 4: 125. 1956. Type: no plant material cited (no specimens cited).

Solanum tuberosum L. var. ibbelianum Danert , Kulturpβanze 4: 125. 1956. Type: no plant material cited (no specimens cited).

Solanum tuberosum L. var. deficiens Danert , Kulturpβanze 4: 126. 1956. Type: no plant material cited (no specimens cited).

Solanum tuberosum L. var. uniforme Danert , Kulturpβanze 4: 126. 1956. Type: no plant material cited (no specimens cited).

Solanum tuberosum L. var. flavescens Danert , Kulturpβanze 4: 126. 1956. Type: no plant material cited (no specimens cited).

Solanum tuberosum L. var. variegatum Danert , Kulturpβanze 4: 126. 1956. Type: no plant material cited (no specimens cited).

Solanum tuberosum L. var. tigrinum Danert , Kulturpβanze 4: 126. 1956. Type: no plant material cited (no specimens cited).

Solanum tuberosum L. var. coloratum Danert , Kulturpβanze 4: 126. 1956. Type: no plant material cited (no specimens cited).

Solanum tuberosum L. var. marmoratum Danert , Kulturpβanze 4: 126. 1956. Type: no plant material cited (no specimens cited).

Solanum diemii Brücher, Darwiniana 13: 108. 1964. Solanum tuberosum L. var. diemii (Brücher) Gorbat., Potato Sp. S. Amer. 375. 2006. Type: Argentina. Río Negro/ Neuquén: Parque Nacional Nahuel Huapí, 760 m, 10.ii.1957, O. Brücher & H. Brücher 900 (lectotype, SI!, designated here; isolectotype, SI!). [C].

Solanum sanmartinense Brücher, Darwiniana 13: 111, fig. 2. 1964. Solanum tuberosum L. var. sanmartinense (Brücher) Gorbat., Potato Sp. S. Amer. 375. 2006. Type: Argentina. Neuquén: San Martín de los Andes, Estancia Collun-Có, 15.ii.1957, O. Brücher & E. Brücher 1095 (holotype, SI!).

Solanum oceanicum Brücher, Repert. Spec. Nov. Regni Veg. 70: 137. 1965. Type: Chile. Region X (Los Lagos): Prov. Chiloé, Chiloé, west coast, S of Río Medina, between Gueno Goique and Laya de los Tigres, 1–2 m, 4.iv.1958, H. Brücher 1196 (no herbaria cited, no specimens yet found). [A].

Solanum tascalense Brücher, Repert. Spec. Nov. Regni Veg. 73: 104. 1966. Type: Argentina. Jujuy: Department. Tumbala, Valle de Tascal, 3600 m, ii.1958, H. Brücher & O. Brücher 1209 (no herbaria cited, no specimens yet found). [A].

Solanum zykinii Lechn., Trudy Prikl. Bot. Genet. Selek. 62(1): 44. 1978. Type: cultivated in Russia from tuber accession collected in Chile (Chiloé Island, C. Ochoa s.n.), A.G. Zykin s.n. (holotype, WIR!). [C].

Solanum ochoanum Lechn., Trudy Prikl. Bot. Genet. Selek. 62(1): 48. 1978. Type: cultivated in Leningrad from tuber accession collected in Chile ( Isla Guaiteca, C. Ochoa s.n. [K-11289]), 25.viii.1978, L.M. Turuleva s.n. (holotype, WIR!). [C].

Solanum parvicorollatum Lechn., Bjull. Vsesojuzn. Inst. Rasteniev. 105: 12. 1980. Type: cultivated in Leningrad from seeds collected in Peru (Junín: Prov. Tarma, Viljarika, Univ Central de Peru), ix.1979, L.M. Turuleva N-3 (holotype, WIR! [WIR-41763]). [A].

Solanum estradae L.E.López, Mutisia 55: 5, figs. 3, 4. 1983. Solanum phureja Juz. & Bukasov var. estradae (L.E.López) Hawkes, Potato , Evolution, Biodivers.& Gen. Res. 179. 1990. Type: Colombia. Tolima: Quindío, Mun. Pijao, Vereda La Quiebra, 1850 m, 15.xi.1973, L.E. López J. CCC-4702 (holotype, COL!; isotype, ICA!). [A].

Solanum andigenum Juz. & Bukasov subsp. ecuatorianum Lechn., Trudy Prikl. Bot. 79: 46. 1983. Type: cultivated in Leningrad from tuber accession collected in Ecuador (Riobamba, N. I. Vavilov K-50), 1933, V.S. Lekhnovich s.n. (holotype, WIR! [WIR-38663]). [A].

Solanum andigenum Juz. & Bukasov subsp. centraliperuvianum Lechn., Trudy Prikl. Bot. 79: 46. 1983. Type: cultivated in Leningrad from tuber accession 4216/3147 collected in Peru (Huánuco: Tarma, S. Juzepczuk 191c), sin dat., V.S. Lekhnovich s.n. (holotype, WIR! [WIR-38955]). [A].

Solanum andigenum Juz. & Bukasov convar. brevipilosum Lechn., Trudy Prikl. Bot. 79: 46. 1983. Type: cultivated in Leningrad from tuber accession 8427 collected in Peru (Huánuco: Huancayo, 1929, S. Juzepczuk 632), 1929, V.S. Lekhnovich s.n. (holotype, WIR! [WIR-39580]). [A].

Solanum andigenum Juz. & Bukasov convar. stenon Lechn., Trudy Prikl. Bot. 79: 46. 1983. Type: cultivated in Leningrad from tuber accession 4249/3227 collected in Peru (Huánuco: Huancayo, Mejorada [as Mejarada]), 1929, S. Juzepczuk 603c), 1929, V.S. Lekhnovich s.n. (holotype, WIR!). [A].

Solanum andigenum Juz. & Bukasov convar. pucamata Lechn., Trudy Prikl. Bot. 79: 46. 1983. Type: cultivated in Leningrad from tuber accession 4204/ 3136 collected in Peru (Huánuco: Tarma, Hacienda Florida, 1928, S. Juzepczuk 155), sin. dat., V.S. Lekhnovich s.n. (holotype, WIR!). [A].

Solanum andigenum Juz. & Bukasov subsp. australiperuvianum Lechn., Trudy Prikl. Bot. 79: 46. 1983. Type: cultivated in Leningrad from tuber accession 4398/3461 collected in Peru (Cusco: Huaccotto, 1928, S. Juzepczuk 1341b), sin. dat., V.S. Lekhnovich s.n. (holotype, WIR! [WIR-37232]). [A].

Solanum andigenum Juz. & Bukasov convar. curtibaccatum Lechn., Trudy Prikl. Bot. 79: 47. 1983. Type: cultivated in Leningrad from tuber accession 4479/3481 collected in Peru (Cusco: San Jeronimo, 1929, S. Juzepczuk 1352), sin. dat., V.S. Lekhnovich s.n. (holotype, WIR!). [A].

Solanum andigenum Juz. & Bukasov convar. obtusiacuminatum Lechn., Trudy Prikl. Bot. 79: 47. 1983. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Ccorau, 1931, S. Juzepczuk 1400), sin. dat., V.S. Lekhnovich s.n. (holotype, WIR! [WIR-37592]). [A].

Solanum andigenum Juz. & Bukasov convar. microstigmatum Lechn., Trudy Prikl. Bot. 79: 47. 1983. Type: cultivated in Leningrad from tuber accession 4373/3312 collected in Peru (Cusco: Sihua, 1929, S. Juzepczuk 1260), sin. dat., V.S. Lekhnovich s.n. (holotype, WIR!). [A].

Solanum andigenum Juz. & Bukasov convar. acutifolium Lechn., Trudy Prikl. Bot. 79: 47. 1983. Type: cultivated in Leningrad from tuber accession 4381/ 3322 collected in Peru (Cusco: Yuncaipampa, S. Juzepczuk 1227), sin. dat. V.S. Lekhnovich s.n. (holotype, WIR!). [A].

Solanum andigenum Juz. & Bukasov convar. titicacense Lechn., Trudy Prikl. Bot. 79: 47. 1983. Type: cultivated in Leningrad from tuber accession 4579/3436 collected in Peru (Cusco: Ollantaytambo, S. Juzepczuk 1078), sin. dat., V.S. Lekhnovich s.n. (holotype, WIR! [WIR-380023]). [A].

Solanum andigenum Juz. & Bukasov convar. nudiculum Lechn., Trudy Prikl. Bot. 79: 47. 1983. Type: cultivated in Leningrad from tuber accession 4548/ 3373 collected in Peru (Cusco: Acomayo, Pomacanchi, 1929, S. Juzepczuk 1979), sin. dat., V.S. Lekhnovich s.n. (holotype, WIR!). [A].

Solanum andigenum Juz. & Bukasov convar. adpressipilosum Lechn., Trudy Prikl. Bot. 79: 48. 1983. Type: cultivated in Leningrad from tuber accession collected in Peru (Cusco: Paucartambo, Misquilla, 1929, S. Juzepczuk 1468a), sin. dat., V.S. Lekhnovich s.n. (holotype, WIR! [WIR-37056]). [A].

Solanum andigenum Juz. & Bukasov subsp. bolivianum Lechn., Trudy Prikl. Bot. 79: 48. 1983. Type: cultivated in Leningrad from tuber accession collected in Bolivia (La Paz, 1929, S. Juzepczuk 1740), sin. dat., V.S. Lekhnovich [1437a] (holotype, WIR!). [A].

Solanum andigenum Juz. & Bukasov convar. macron Lechn., Trudy Prikl. Bot. 79: 48. 1983. Type: cultivated in Leningrad from tuber accession collected in Bolivia (La Paz, 1928, S. Juzepczuk 1783), sin. dat., V.S. Lekhnovich s.n. (holotype, WIR!). [A].

Solanum andigenum Juz. & Bukasov convar. brevicalyces Lechn., Trudy Prikl. Bot. 79: 48. 1983. Type: cultivated in Leningrad from tuber accession collected in Bolivia (La Paz, 1929, S. Juzepczuk 11660c), sin. dat., V.S. Lekhnovich s.n. [1409a] (holotype, WIR!). [A].

Solanum andigenum Juz. & Bukasov convar. brachistylum Lechn., Trudy Prikl. Bot. 79: 48. 1983. Type: cultivated in Leningrad from tuber accession collected in Bolivia (La Paz: Cohi, 1929, S. Juzepczuk 1536), sin. dat., V.S. Lekhnovich s.n. [1429a] (holotype, WIR!). [A].

Solanum andigenum Juz. & Bukasov convar. elongatibaccatum Lechn., Trudy Prikl. Bot. 79: 48. 1983. Type: cultivated in Leningrad from tuber accession collected in Bolivia (La Paz, 1928, S. Juzepczuk 1526), sin. dat., V.S. Lekhnovich s.n. (holotype, WIR! [WIR-38661]). [A].

Solanum andigenum Juz. & Bukasov subsp. argentinicum Lechn., Trudy Prikl. Bot. 79: 48. 1983. Type: cultivated in Leningrad from tuber accession collected in Argentina (Tucumán: Lilio, 1951, R.D. Schechai 2186-3), sin. dat., V.S. Lekhnovich s.n. [2898] (holotype, WIR!). [A].

Solanum hygrothermicum Ochoa, Econ. Bot. 38: 129. 1984. Solanum phureja Juz. & Bukasov subsp. hygrothermicum (Ochoa) Hawkes, Potato , Evolution, Biodivers. & Gen. Res. 179. 1990. Type: Peru. San Martín: Cumbasa River, 600 m, C. Ochoa 1194 (holotype, CUZ! [original citation herb. Ochoa]; isotype, US! [US-2971891]).

Solanum phureja Juz. & Bukasov var. caeruleum Ochoa, Phytologia 65: 103. 1988. as ‘ caeruleus ’. Type: Bolivia. La Paz: Prov. Franz Tamayo, Queara, 3350 m, C. Ochoa & A. Salas 14973 (holotype, CUZ! [original citation herb. Ochoa]; isotypes, CIP!, F!). [A].

Solanum phureja Juz. & Bukasov var. sanguineum Ochoa, Phytologia 65: 103. 1988. as ‘ sanguineus ’. Type: Bolivia. La Paz: Prov. Inquisivi, Quime, 2900 m, C. Ochoa & A. Salas 15515 (holotype, CUZ! [original citation herb. Ochoa]; isotype, CIP!). [A].

Solanum phureja Juz. & Bukasov forma sayhuanimayo Ochoa, Phytologia 65: 103. 1988. Type: Bolivia. La Paz: Prov. Franz Tamayo, Sayhuanimayo, C. Ochoa & A. Salas 15001 (holotype, CUZ! [original citation herb. Ochoa]; isotype, CIP!). [A].

Solanum phureja Juz. & Bukasov forma viuda Ochoa, Phytologia 65: 103. 1988. Type: Bolivia. La Paz: Prov. Larecaja, Sorata, 2700 m, C. Ochoa 3536 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum phureja Juz. & Bukasov var. flavum Ochoa, Phytologia 65: 103. 1988. Type: Bolivia. La Paz: Prov. Franz Tamayo, C. Ochoa & A. Salas 14970 (holotype, CUZ! [original citation herb. Ochoa]; isotype, CIP!). [A].

Solanum phureja Juz. & Bukasov forma orbiculatum Ochoa, Phytologia 65: 104. 1988. as ‘ orbiculata ’. Type: Bolivia. La Paz: Prov. Franz Tamayo, near Puina, between Ayamachay and Sayhuanimayo, 3200 m, C. Ochoa & A. Salas 15009 (holotype, CUZ! [original citation herb. Ochoa]; isotypes, CIP!, F!). [A].

Solanum phureja Juz. & Bukasov var. rubro-roseum Ochoa, Phytologia 65: 104. 1988. as ‘ rubro-rosea ’. Type: Bolivia. La Paz: Prov. Franz Tamayo, Queara, 3400 m, C. Ochoa & A. Salas 14971 (holotype, CUZ! [original citation herb. Ochoa]; isotype, CIP!). [A].

Solanum phureja Juz. & Bukasov forma timusi Ochoa, Phytologia 65: 104. 1988. Type: Bolivia. La Paz: Prov. Larecaja, Timusi, C. Ochoa 3910 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum phureja Juz. & Bukasov var. janckophureja Ochoa, Phytologia 65: 104. 1988. Type: Bolivia. La Paz: Prov. Larecaja, Sorata, 2600 m, C. Ochoa & A. Salas 15472 (holotype, CUZ! [original citation herb. Ochoa]; isotype, F!).

Solanum stenotomum Juz. & Bukasov forma chiarckati Ochoa, Phytologia 65: 104. 1988. Type: Bolivia. La Paz: Prov. Murillo, C. Ochoa 3925 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum stenotomum Juz. & Bukasov forma chiarphinu Ochoa, Phytologia 65: 104. 1988. Type: Bolivia. La Paz: Prov. Murillo, La Paz, 3650 m, C. Ochoa 3924 (holotype, herb. Ochoa, not found). [A].

Solanum stenotomum Juz. & Bukasov forma pacajes Ochoa, Phytologia 65: 104. 1988. Type: Bolivia. La Paz: Prov. Pacajes, Caquiaviri, C. Ochoa 3551 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum stenotomum Juz. & Bukasov forma alkkapitiquina Ochoa, Phytologia 65: 105. 1988. Type: Bolivia. La Paz: Prov. Los Andes, Palculo, 3800 m, C. Ochoa 3184 (holotype, herb. Ochoa, not found). [A].

Solanum stenotomum Juz. & Bukasov forma janckochojllu Ochoa, Phytologia 65: 105. 1988. Type: Bolivia. La Paz: Prov. Murillo, La Paz, 3650 m, C. Ochoa 3938 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum stenotomum Juz. & Bukasov forma puluwayku Ochoa, Phytologia 65: 105. 1988. Type: Bolivia. Oruro: Prov. Oruro, Oruro, 3650 m, C. Ochoa 10493 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum stenotomum Juz. & Bukasov forma wilachojllu Ochoa, Phytologia 65: 105. 1988. Type: Bolivia. La Paz: Prov. Murillo, La Paz, 3650 m, C. Ochoa 3942 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum stenotomum Juz. & Bukasov var. arichuwa Ochoa, Phytologia 65: 105. 1988. Type: Bolivia. La Paz: Prov. Ingavi, Huatajata, 3850 m, C. Ochoa 3937 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum stenotomum Juz. & Bukasov var. chojllu Ochoa, Phytologia 65: 105. 1988. Type: Bolivia. La Paz: Prov. Murillo, La Paz, 3650 m, C. Ochoa 3912 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum stenotomum Juz. & Bukasov var. kkamara Ochoa, Phytologia 65: 105. 1988. Type: Bolivia. La Paz: Prov. Murillo, La Paz, 3650 m, C. Ochoa 3920 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum stenotomum Juz. & Bukasov var. luru Ochoa, Phytologia 65: 105. 1988. Type: Bolivia. Potosí: Prov. Frías, Tinguipaya, 3900 m, C. Ochoa 3949 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum stenotomum Juz. & Bukasov var. pitiquina Ochoa, Phytologia 65: 105. 1988. Type: Bolivia. La Paz: Prov. Ingavi, Tiahuanaco, C. Ochoa 3535 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum stenotomum Juz. & Bukasov forma azureockati Ochoa, Phytologia 65: 106. 1988. Type: Bolivia. La Paz: Prov. Inquisivi, Quime, 3200 m, C. Ochoa & A. Salas 15514 (holotype, herb. Ochoa, not found; isotype, CIP!). [A].

Solanum stenotomum Juz. & Bukasov forma churipuya Ochoa, Phytologia 65: 106. 1988. Type: Bolivia. La Paz: Prov. Murillo, Walata Chico, 3850 m, C. Ochoa 3152 (holotype, herb. Ochoa, not found). [A].

Solanum stenotomum Juz. & Bukasov forma kkarachipampa Ochoa, Phytologia 65: 106. 1988. Type: Bolivia. Potosí: Prov. Linares, Kkarachipampa, 4000 m, C. Ochoa 10481 (holotype, herb. Ochoa, not found). [A].

Solanum stenotomum Juz. & Bukasov forma laram Ochoa, Phytologia 65: 106. 1988. Type: Bolivia. La Paz: Prov. Murillo, La Paz, 3650 m, C. Ochoa 3926 (holotype, herb. Ochoa, not found). [A].

Solanum stenotomum Juz. & Bukasov forma phitikalla Ochoa, Phytologia 65: 106. 1988. Type: Bolivia. La Paz: Prov. Murillo, La Paz, 3650 m, C. Ochoa 3928 (holotype, herb. Ochoa, not found). [A].

Solanum stenotomum Juz. & Bukasov forma quime Ochoa, Phytologia 65: 106. 1988. Type: Bolivia. La Paz: Prov. Inquisivi, Quime, 2900 m, C. Ochoa & A. Salas 15513 (holotype, herb. Ochoa, not found). [A].

Solanum stenotomum Juz. & Bukasov var. zapallo Ochoa, Phytologia 65: 106. 1988. Type: Bolivia. La Paz: Prov. Ingavi, Tiahuanaco, 3680 m, C. Ochoa 3145 (holotype, herb. Ochoa, not found). [A].

Solanum chaucha Juz. & Bukasov var. ccoesullu Ochoa, Phytologia 65: 107. 1988. Type: Peru. Cusco: Prov. Quispicanchis, Ccatacca, 3680 m, C. Ochoa 7848 (holotype, herb. Ochoa, not found). [A].

Solanum chaucha Juz. & Bukasov var. ckati Ochoa, Phytologia 65: 107. 1988. Type: Bolivia. La Paz: Prov. Murillo, La Paz, 3650 m, C. Ochoa 3929 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum chaucha Juz. & Bukasov var. khoyllu Ochoa, Phytologia 65: 107. 1988. Type: Bolivia. Cochabamba: Prov. Cercado, Cochabamba, brought from Colomi, 3200 m, C. Ochoa 3948 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum chaucha Juz. & Bukasov var. pucasuitu Ochoa, Phytologia 65: 108. 1988. Type: Peru. Junín: Prov. Yauli, Chacapalpa, 3700 m, C. Ochoa 6 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum chaucha Juz. & Bukasov var. surimana Ochoa, Phytologia 65: 108. 1988. Type: Bolivia. Oruro: Prov. Poopó, Urmiri, 3750 m, C. Ochoa 3946 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum tuberosum L. forma alkkaimilla Ochoa, Phytologia 65: 108. 1988. Type: Bolivia. La Paz: Prov. Murillo, La Paz, 3650 m, C. Ochoa 3902 (holotype, herb.Ochoa, not found; isotype, CIP!). [A].

Solanum tuberosum L. forma islaimilla Ochoa, Phytologia 65: 108. 1988. Type: Bolivia. La Paz: Prov. Murillo, La Paz, 3650 m, C. Ochoa 3969 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma janckochockella Ochoa, Phytologia 65: 108. 1988. Type: Bolivia. La Paz: Prov. Ingavi, Sullcatiti, CIP-702616 (holotype, CIP!).

Solanum tuberosum L. forma saniimilla Ochoa, Phytologia 65: 108. 1988. Type: Bolivia. La Paz: Prov. Murillo, Huatajata, 3850 m, C. Ochoa 3972 (holotype, CUZ! [original citation herb. Ochoa]). [A].

Solanum tuberosum L. forma amajaya Ochoa, Phytologia 65: 109. 1988. Type: Bolivia. Potosí: Prov. Frías, Azangaro, 3800 m, C. Ochoa 10508 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma chiarpala Ochoa, Phytologia 65: 109. 1988. Type: Bolivia. La Paz: Prov. Ingavi, Huanccolo, 3680 m, C. Ochoa 3977 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma janckopala Ochoa, Phytologia 65: 109. 1988. Type: Bolivia. Oruro: Prov. Poopó, Urmiri, 3750 m, C. Ochoa 3996 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma wilapala Ochoa, Phytologia 65: 109. 1988. Type: Bolivia. La Paz: Prov. Ingavi, Huanccollo, 3680 m, C. Ochoa 3986 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma wilaimilla Ochoa, Phytologia 65: 109. 1988. Type: cultivated at Colección Peruana de Papa (Peruvian Potato Collection), Universidad Agraria La Molina, Lima, Peru from accession collected in Bolivia (La Paz Prov. Murillo, Chiclaya Grande, 3840), CPP-1810 (holotype, herb. Ochoa, not found).

Solanum tuberosum L. forma chojosajama Ochoa, Phytologia 65: 109. 1988. Type: Bolivia. La Paz: Prov. Omasuyos, Huatajata, 3850 m, C. Ochoa 3971 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma wilamonda Ochoa, Phytologia 65: 109. 1988. Type: Bolivia. La Paz: Prov. Murillo, La Paz, 3650 m, C. Ochoa 3967 (holotype, herb. Ochoa, not found; isotype, CIP!). [A].

Solanum tuberosum L. forma huacazapato Ochoa, Phytologia 65: 110. 1988. Type: cultivated at Colección Peruana de Papa (Peruvian Potato Collection), Univ. Agraria La Molina, Lima, Peru, from accession collected in Bolivia (La Paz: Prov. Los Andes, Pucarani, 3900 m), no date, CPP-1843 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma jancckokkoyllu Ochoa, Phytologia 65: 110. 1988. Type: Bolivia. La Paz: Prov. Murillo, Huancané, 3900 m, CPP-1814 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma kunurana Ochoa, Phytologia 65: 110. 1988. Type: Bolivia. Potosí: Prov. Chayanta, Ravelo, 3500 m, C. Ochoa 12115 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma overita Ochoa, Phytologia 65: 110. 1988. Type: Bolivia. La Paz: Prov. Murillo, La Paz, 3650 m, C. Ochoa 3991 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma surico Ochoa, Phytologia 65: 110. 1988. Type: Bolivia. La Paz: Prov. Camacho, Puerto Acosta, 3850 m, C. Ochoa 2803 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma tinguipapa Ochoa, Phytologia 65: 110. 1988. Type: Bolivia. Potosí: Prov. Frias, Tinguipaya, 3900 m, C. Ochoa 10648 (holotype, herb. Ochoa, not found; isotype, CIP!). [A].

Solanum tuberosum L. forma wilahuaycku Ochoa, Phytologia 65: 110. 1988. Type: Bolivia. La Paz: Prov. Murillo, La Paz, 3650 m, C. Ochoa 3897 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. var. lelekkoya Ochoa, Phytologia 65: 111. 1988. Type: Bolivia. La Paz: Prov. Larecaja, Sorata, 2700 m, C. Ochoa & A. Salas 11792 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. var. malcachu Ochoa, Phytologia 65: 111. 1988. Type: Bolivia. Potosí: Prov. Saavedra, Lequezana, 3300 m, C. Ochoa 3959 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma huichinkka Ochoa, Phytologia 65: 111. 1988. Type: Bolivia. Potosí: Prov. Frias, Gran Pena, 3800 m, C. Ochoa 10671 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma milagro Ochoa, Phytologia 65: 111. 1988. Type: Bolivia. La Paz: Prov. Murillo, Huatajata, 3850 m, CPP-1807 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. var. lelekkoya Ochoa, Phytologia 65: 111. 1988. Type: Bolivia. La Paz: Prov. Larecaja, Sorata, 2700 m, C. Ochoa & A. Salas 11792 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma chiarlelekkoya Ochoa, Phytologia 65: 111. 1988. Type: Bolivia. La Paz: Prov. Larecaja, near Sorata, Curupampa, 2700 m, C. Ochoa & A. Salas 11795 (holotype, herb. Ochoa, not found; isotype, CIP!). [A].

Solanum tuberosum L. forma laramlelekkoya Ochoa, Phytologia 65: 111. 1988. Type: Bolivia. Chuquisaca: Prov. Sud Cinti, Higueras, 2600 m, C. Ochoa 11976 (holotype, herb. Ochoa, not found).

Solanum tuberosum L. var. runa Ochoa, Phytologia 65: 111. 1988. Type: Bolivia. Cochabamba: Cochabamba, brought from Punata, 3200 m, C. Ochoa 2999 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. var. malcachu Ochoa, Phytologia 65: 111. 1988. Type: Bolivia. Potosí: Prov. Saavedra, Lequezana, 3300 m, C. Ochoa 3959 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. var. murukewillu Ochoa, Phytologia 65: 112. 1988. Type: Peru. Cusco: Prov. Calca, between Chumpe and Paro/Paro, 3700 m, C. Ochoa 4020 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. var. sipancachi Ochoa, Phytologia 65: 112. 1988. Type: Bolivia. Potosí: Prov. Saavedra, Lequezana, 3300 m, C. Ochoa 3958 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. var. tacaco Ochoa, Phytologia 65: 112. 1988. Type: Bolivia. La Paz: Prov. Aroma, Huatajata, 3900 m, C. Ochoa 2791 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. var. murukewillu Ochoa, Phytologia 65: 112. 1988. Type: Peru. Cusco: Prov. Calca, between Chumpe and Paro-Paro, 3700 m, C. Ochoa 4020 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma azulruna Ochoa, Phytologia 65: 112. 1988. Type: Bolivia. Potosí: Prov. Saavedra, Lequezana, 3300 m, C. Ochoa 3954 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. var. sicha Ochoa, Phytologia 65: 112. 1988. Type: Bolivia. La Paz: Prov. Inquisivi, Quime, 2700 m, C. Ochoa & A. Salas 15517 (holotype, herb. Ochoa, not found; isotype, CIP!). [A].

Solanum tuberosum L. var. sipancachi Ochoa, Phytologia 65: 112. 1988. Type: Bolivia. Potosí: Prov. Saavedra, Lequezana, 3300 m, C. Ochoa 3958 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma pulo Ochoa, Phytologia 65: 112. 1988. Type: Bolivia. Potosí: Prov. Alonso Ibáñez, Sacaca, CIP-702704 (holotype, herb. Ochoa, not found; isotype, CIP!). [A].

Solanum tuberosum L. forma wilakoyu Ochoa, Phytologia 65: 112. 1988. Type: cultivated in Lima, Peru from collection made in Bolivia [?], CIP-702691 (holotype, herb. Ochoa, not found; isotype, CIP!).

Solanum tuberosum L. var. taraco Ochoa, Phytologia 65: 112. 1988. Type: Bolivia. La Paz: Prov. Aroma, Huatajata, 3900 m, C. Ochoa 2791 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma alkkasilla Ochoa, Phytologia 65: 113. 1988. Type: Bolivia. Potosí: Prov. Saavedra, Esquiri, 2600 m, C. Ochoa 3989 (holotype, herb. Ochoa, not found). [A].

Solanum tuberosum L. forma yuractaraco Ochoa, Phytologia 65: 113. 1988. Type: Bolivia. Potosí: Prov. Frías, Santa Lucía, 3800 m, C. Ochoa 10685 (holotype, herb. Ochoa, not found).

Description: Herbs 0.4–1.4 m tall, ascending to erect or semi-erect, decumbent or prostrate. Stems 5–19 mm in diameter at base of plant, unwinged or with wings to 5 mm, nearly glabrous to densely pubescent, green or purple to green and purple mottled. Sympodial units tri- to plurifoliate, not geminate. Leaves odd-pinnate, the blades 8–22 x 5–13 cm, medium to dark green, membranous to chartaceous, leaf surface dull to shiny, nearly glabrous to densely pubescent adaxially and abaxially, with hairs like those of the stems; lateral leaβet pairs 3–8, only slightly decreasing in size from the apex to the base; most distal lateral leaβets 3–8 x 1.5–5.5 cm, ovate to elliptic to broadly ellipticlanceolate, the apex acute to acuminate or shortly acuminate, the base generally oblique, rounded to cuneate to cordate, rarely truncate; terminal leaβet 3.5–9.0 x 1.0– 5.5 cm, ovate to elliptic to broadly elliptic-lanceolate, the apex acute to acuminate or shortly acuminate, the base generally oblique, rounded to cuneate to cordate, rarely truncate; interjected leaβets 0–45, sessile to short petiolulate, ovate to elliptic to broadly elliptic-lanceolate; petioles 2–6 cm, pubescent as the stems. Pseudostipules 4–25 mm long, auriculate to semi-elliptic, falcate, pubescent with hairs like those of the stem. Inβorescences 5–11 cm, terminal with a subtending axillary bud, generally in distal half of the plant, usually forked, with 0–25 βowers, with all βowers apparently perfect, the axes pubescent with hairs like those of the stem; peduncle 0–22 cm long; pedicels 10–35 mm long in βower and fruit, spaced 1–10 mm apart, articulated in approximately the middle third. Flowers homostylous, pentamerous. Calyx 0–10 mm long, the tube 1–2 mm, the lobes 0–9 mm, short and acute to long attenuate, the acumens 1–8 mm long, with hairs like those of the stem. Corolla 2–6 cm in diameter, rotate to rotatepentagonal, white to pink to lilac to blue to purple to red–purple, uniform or with white acumens or with a secondary colour stippled, in bands, or in the star, adaxially or abaxially or both, the tube 1–2 mm long, the acumens 3–5 mm long, often prominent, the corolla edges βat, not folded dorsally, glabrous abaxially, minutely puberulent adaxially, especially along the midribs, ciliate at the margins, especially at the tips of the corollas. Stamens with the filaments 1–2 mm long; anthers 3–8 mm long, cordate at the base, lanceolate, connivent, yellow, poricidal at the tips, the pores lengthening to slits with age. Ovary glabrous; style 9–13 x 1 mm, exceeding stamens by 7 mm, straight, papillose in the distal half; stigma capitate. Fruit a globose to ovoid berry, 1–4 cm in diameter, green to green tinged with white or purple spots or bands when ripe, glabrous. Seeds from living specimens ovoid and c. 2 mm long, whitish to greenish in fresh condition and drying brownish, with a thick covering of ‘hairlike’ lateral walls of the testal cells that make the seeds mucilaginous when wet, green–white throughout; testal cells honeycomb-shaped when lateral walls removed by enzyme digestion. Chromosome number: 2 n = 2 x = 24 (Ochoa & Salas 14973), 2 n = 3 x = 36 (Ochoa 2202), 2 n = 4 x = 48 (Ochoa 2159); all vouchers at the Herbarium of the International Potato Center, Lima, Peru).

Phenology: Flowering and fruiting from January to May.

Distribution: Landrace populations of S. tuberosum grow from western Venezuela south to northern Argentina (Andean populations), and then, with a gap of distribution in south-central Chile, in the islands of the Chonos and Guaitecas Archipelagos and adjacent mainland Chile (lowland Chilean populations), with the Andean populations growing mainly from 2000 to 4600 m in elevation and the lowland Chilean populations at or near sea level to 500 m. Landrace populations are still maintained out of their natural range, introduced in post-Colombian times, in Mexico and Central America, the Shimla Hills of India and the Canary Islands. The modern cultivated potato (also classified as S. tuberosum ) is cultivated worldwide.

The spelling of the epithet ‘ andigenum ’, as originally used by Juzepczuk & Bukasov (1929), has often been changed by subsequent authors to ‘ andigena ’, and it occasionally appears as ‘ andigena ’ in indices. In this article and on the Solanaceae Source website, all these have been corrected to the original spelling. The name S. andigenum was validly published in 1929 (see discussion under S. curtilobum above), but is often cited as having been coined later (in Bukasov, 1930). Together with S. andigenum , other names treated by us as synonyms of S. tuberosum which were validly published in 1929 and cited in various indices as having been published in 1930 are S. boyacense Juz. & Bukasov , S. chaucha Juz. & Bukasov , S. goniocalyx Juz. & Bukasov , S. mamilliferum Juz. & Bukasov , S. phureja Juz. & Bukasov , S. rybinii Juz. & Bukasov , S. stenotomum Juz. & Bukasov and S. tenuifilamentum Juz. & Bukasov. Descriptions in Latin accompanied these names and new species’ rank epithets in Bukasov (1930), but not those of taxa described at varietal and form rank . It may be that Juzepczuk and Bukasov did not really intend the many names coined at these lower ranks to be of the same sort or equivalent to the species’ level names, but, under the nomenclatural rules of the day, they must be treated as validly published in the absence of evidence to the contrary.

Hawkes & Hjerting (1989: 378) superβuously lectotypified S. phureja with Juzepczuk 1654 at WIR; this lectotypification had already been carried out by Korovina et al. (1985). Hawkes & Hjerting (1989: 388) lectotypified S. chaucha with a specimen of Juzepczuk 1010 said to be collected in Peru in ‘herb. Vavilov Institute’ (WIR), but Ovchinnikova et al. (2009: 583) failed to encounter any such specimen in the WIR herbarium. They did find three sheets of this collection (grown from Juzepczuk 1010 in Russia) at LE, and designated one of these as the lectotype: ( Ovchinnikova et al., 2009). Several isotypes with labels in Bukasov’s handwriting with the tuber accession number Juzepczuk 1010 and the same date of collection as the specimens in LE, which were presumably given to Hawkes by Russian potato taxonomists during his Russian trip in 1938 (see Hawkes, 2004), are today held at K. Hawkes & Hjerting (1989) also cite other specimens in ‘herb. JGH’ that were obtained from Russian potato taxonomists during Hawkes’ visit (e.g. a sheet of Juzepczuk 1166, isotype of S. juzepczukii , now at K [K000585544]).

Gorbatenko & Hawkes (1996) lectotypified S. goniocalyx with a specimen taken from the accession Juzepczuk 571 from Cerro de Pasco ( Peru) that was cited in the original description, because they were unable to locate any specimens of the number cited in the original publication as the type for this name (Juzepczuk 587) in either LE or WIR. A sheet of the type was apparently loaned to C. Ochoa, but has not been located. Should this sheet be located, it would be the obligate lectotype for S. goniocalyx , which would have to be re-lectotypified. Given the uncertain location of much of Ochoa’s herbarium (see synonymy above), we cite Gorbatenko & Hawkes’ (1996) lectotypification here.

In describing S. andigenum var. mexicanum and var. columbianum, Bukasov (1930) did not specify material for these taxa, but instead described them in reference to their component forms. We have followed them in assuming forma tolucanum to be homotypic (and thus probably an illegimate name for forma mexicanum ); they state ‘the typical form’ in reference to forma tolucanum . The same principle has been used for var. colombianum and forma tocanum .

We have neotypified several of the names coined by Bukasov and Lekhnovich ( Bukasov, 1933) with material collected in Leningrad by Hawkes in 1938 (see discussion above), as no original material was found in any of the St. Petersburg herbaria.

We have used specimens made of βowering stems of one of the many cultivars cited by Danert in his original description, which were cultivated in Gatersleben in 1955, as lectotypes for five of these names; these are the only names for which material has been found in GAT (K. Pistrick, 26.x.2010).

No original material of S. rybinii var. pastoense has been found in the Hawkes’ material held at K, and so this name has been neotypifed with material of the type accession (collected in the market in Pasto, Colombia by E. K. Balls as tubers and grown as EPC accession 979) grown at the John Innes Institute in Norwich ( UK), near the original cultivation site of Cambridge. Solanum subandigenum is not represented in the Hawkes’ herbarium material at K, but a specimen collected in the field is held in BM; this sheet, consisting of several sterile seedlings, has been designated the lectotype in the absence of other herbarium material. The only herbarium material we have found of the type collection of S. cardenasii (Balls 6299) is a βower dissection (prepared in the manner usually undertaken by Hawkes) at K; this has been designated the lectotype, although material with leaves would have been preferable. No specimens were cited by Hawkes (1944) in his description of S. yabari , but specimens were cited as belonging to the two varieties (neither validly published because of a lack of Latin diagnoses, Article 36.1, see above). Two of these were cited in the Latin diagnosis (but not in the English description) of S. yabari: Yabar 44 (EPC-698) and EPC-1077 without a collector. We have selected a sheet of Yabar 44 at K, collected from plants grown at Cambridge from EPC-698, as the lectotype for this species.

In his 1983 paper describing infraspecific variation in Andean potatoes, V. S. Lekhnovich used the infraspecific rank ‘convarietas’, a rank not explicitly recognized in ICBN (Article 4.1), but Article 4.3 states that additional ranks may be added provided that confusion is not introduced. Lekhnovich (1983) appears to have deliberately used the rank convar. to indicate ‘groups of varietatis’, thus positioning it between subspecies and variety. We are treating these epithets as validly published despite the rank being nonstandard, as no confusion is apparent in Lekhnovich’s formulation of them.

Figures

Figure 4. Lectotype specimen of Solanum tuberosum L. (Chilotanum group) held in LINN (LINN 248.12). Reproduced with permission of the Linnean Society of London.

Figure 5. Lectotype specimen of Solanum andigenum Juz. & Bukasov (Solanum tuberosum L. Andigenum group) held in LE. Reproduced with permission of the V. L. Komarov Botanical Institute.