Ampullaceana relicta relicta ( Polinski , 1929)

Vinarski, Maxim V., Aksenova, Olga V. & Bolotov, Ivan N., 2020, Taxonomic assessment of genetically-delineated species of radicine snails (Mollusca, Gastropoda, Lymnaeidae), Zoosystematics and Evolution 96 (2), pp. 577-608 : 577

16a. Ampullaceana relicta relicta ( Polinski, 1929) Figs 8M ; 9A.

Radix relicta Poliński 1929: 158.

Lymnaea peregra - Hubendick 1951: 146, figs 1, 9 (partim).

Lymnaea (Peregriana) relicta - Kruglov and Starobogatov 1983b: 1469, figs 2(12), 3(12); 1993b: 166, fig. 5C.

Radix relicta - Albrecht et al. 2008: 160, fig. 1; Welter-Schultes 2012: 54, textfig.; Glöer 2019: 244, fig. 305

Ampullaceana relicta relicta - Aksenova et al. 2018a: 4.

TL.

Lake Ohrid.

Types.

Whereabouts unknown.

This subspecies has been characterised both morphologically and genetically by Albrecht et al. (2008). We examined the copulatory apparatus of two specimens and found that its structure is typical for the genus. Both praeputium and penis sheath are light-coloured; praeuputium is substantially larger than the penis sheath (see Fig. 8M ), the ratios between their lengths are 1.42 and 1.46, respectively.

A. r. relicta inhabits Ohrid Lake (Albania and Macedonia), as well the Drin system, at least upstream to the dam Globočičko, approximately 15 km N of the lake ( Albrecht et al. 2008).

Figures

Figure 8. Copulatory apparatuses of the species in the genera Ampullaceana, Peregriana, Kamtschaticana, Tibetoradix, Racesina, Orientogalba and Bullastra. A. Ampullaceana ampla (11.07.2017, Ukraine, Transcarpathian Region, Irshava River). B. A. balthica (Germany, Mecklenburg-Vorpommern, lake Torgelower See). C. A. dipkunensis (15.09.2016, Ukraine, Transcarpathian Region, Tisa River). D. A. fontinalis (20.08.2017, Russia, Rostov Region, Severskiy Donets River). E. A. lagotis (29.06.2016, Tajikistan, a stream in the Panj River floodplain). F. Peregriana dolgini (13.07.2015, Russia, Tyumen' Region, a floodplain of the Tol'ka oxbow). G. Peregriana peregra (01.07.2006, Russia, Omsk Region, vicinity of Chernoluchye settlement). H. Kamtschaticana kamtschatica (12.04.2014, Russia, Kamchatka Peninsula, Valley of Geysers). I. Tibetoradix hookeri (09 - 10.2012, China, Tibet, a brook, 250 km N of Lhasa). J. T. kozlovi (after Aksyonova et al. 2018 a, modified). K. Racesina oxiana (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village). L. R. siamensis (24.11.2016, Myanmar, a water reservoir on the Yetho River). M. Ampullaceana relicta relicta (2005, Ohrid Lake). N. Orientogalba ollula (08.07.2010, China, Shaanxi Province, a roadside ditch near Yandi village). O. Bullastra cumingiana (Philippines, Luzon Island, Lake Sampaloc). Scale bars: 1 mm. B - after Schniebs et al. (2011), modified.

Figure 9. Shells of species of the genera Ampullaceana, Kamtschaticana, Myxas, Tibetoradix, Racesina, Orientogalba and Bullastra. A. Ampullaceana relicta relicta (05.1975, Macedonia, a spring near St. Naum monastery, Ohrid Lake region; NHMW). B. A. r. pinteri, a paratype (05.06.1973, Macedonia, Prespa Lake near Perovo; MHMW). C, D. Kamtschaticana kamtschatica (Russia, Kamchatka Peninsula, Valley of Geysers; RMBH). E. Myxas glutinosa (14.09.1963, Kazakhstan, floodplain of the Tobol River near Kustanay City; ZISP). F, G. Tibetoradix hookeri, two syntypes (NHMUK). H. T. kozlovi, the holotype (ZISP). I. Racesina luteola (India, Bengal, Paksay, Pabna; NHMUK). J. R. oxiana (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village; LMBI). K. R. siamensis (" Thailand "; ZMB). L. Orientogalba viridis, a syntype (Guam Island; NMNH). M. O. ollula (1883, Japan, Yokohama; NHMUK). N. Limnaea bowelli, a type (after Preston 1909). O. Orientogalba cf. bowelli (30.06.1924, China, Tibet, Lingka, 14,500 feet a. s. l.; NHMUK). P. Bullastra cumingiana (Philippine, Cebu Island; NHMUK). Scale bars: 2 mm (C, D, F-H, L, M, O), 5 mm (A, B, E, J-K, N, P).