Sweltsa mogollonica, Stark & Baumann, 2018

Stark, Bill P. & Baumann, Richard W., 2018, Two New Stonefly Species In The Sweltsa Coloradensis (Banks) Complex (Plecoptera: Chloroperlidae), Illiesia 14 (2), pp. 30-43 : 41-42

Sweltsa mogollonica , sp. n.

( Figs. 17 , 30-35 , 40-41 , 44, 47 ) http://lsid.speciesfile.org/urn:lsid: Plecoptera .speciesfile.org: TaxonName:502537

Material examined. Holotype ♂, 1♂, 10♀ paratypes, Arizona, Apache Co. , North Fork White

River, Hwy 473, 12 June 1974, B. Stark, (holotype United States National Museum, paratypes BPSC). Additional paratypes, Arizona: Graham Co., Grant Creek, Hospital Flat Campground , Graham Mountains , 5 June 1982, R . W. Baumann, S.M. Clark, 12♂, 16♀ ( BYU). Greenlee Co., Hannagan Creek, Hannagan Meadow , Sitgreaves National Forest , 29 May 1993, C. R . Nelson, J.K. Gelhaus, 29♂, 14♀ ( BYU). New Mexico: Catron Co., Willow Creek, Willow Creek Campground , 11 June 1974, B. Stark, 8♂, 7♀ ( BPSC) .

Adult habitus. Pigment pattern not distinct from that of S. coloradensis or S. lyrata ( Fig. 44 ). Male forewing length 6.8-7.5 mm, female forewing length 7.8-9.0 mm.

Male epiproct. Maximum width 320-340 μm, length 390-430 μm. Base and stem relatively wide and usually, more or less continuous with epiproct body ( Figs. 30-35 ). Apical lobes not projecting from shoulders ( Fig. 47 ).

Aedeagus (n=1). Length 357 μm, width 614 μm. Similar in shape and setation to S. coloradensis but conspicuously shorter in the only everted specimen available ( Fig. 17 ).

Female terminalia. Subgenital plate not always distinct from that of S. coloradensis or S. lyrata . The specimen illustrated in Fig. 39-40 has a more rounded subgenital plate than is typical for the other species.

Larva. Unknown.

Etymology. This species name is based on the Mogollon Rim of Arizona and New Mexico.

Diagnosis and distribution. Epiprocts of specimens of this species have, at most, slightly projecting shoulders near the hook base and are relatively wide across the lateral lobes ( Figs. 29 , 32 ). The new species is similar to S. lyrata in the former character but the two are distinguished on the basis of the relatively greater width of the epiproct for males of S. lyrata (compare Figs. 19 and 29 ). The aedeagus may also be distinct (see Fig. 17 ), but verification is needed from additional material. Specimens are known from three Arizona sites, and an adjacent site in New Mexico.

Figures

Figs. 14-17. Sweltsa sp. aedeagus, ventral. 14. S. coloradensis, Nevada, White Pine Co., Berry Creek. 15. S. coloradensis, Colorado, Gilpin Co., South Boulder Creek. 16. S. lyrata, Yukon, Ogilvie River. 17. S. mogollonica, Arizona, Graham Co., Grant Creek. (al = apical lobe; ll = lateral lobe; vl = ventral lobe).

Figs. 30-35. Sweltsa mogollonica male epiproct. 30-31. Arizona, Apache Co., North Fork White River. 32-33. New Mexico, Catron Co., Willow Creek. 34-35. Arizona, Greenlee Co. Hannagan Creek.

Figs. 36-41. Sweltsa sp. female terminalia, ventral. 36-37. Sweltsa coloradensis, Colorado, Gilpin Co., South Boulder Creek. 38-39. Sweltsa lyrata, Montana, Mineral Co., St. Regis River. 40-41. Sweltsa mogollonica, Arizona, Apache Co., North Fork White River (SgP = subgenital plate).

Figs. 42-47. Sweltsa sp. head and pronotal pigment and epiproct variation. 42. Sweltsa coloradensis, Colorado, Eagle Co., Deep Creek. 43. Sweltsa lyrata, Montana, Mineral Co., St. Regis River. 44. Sweltsa mogollonica, New Mexico, Catron Co., Willow Creek. Pigment patterns shown here are variable within and between populations of the three species. 45. Sweltsa coloradensis epiproct, Colorado, Gilpin Co., South Boulder Creek. 46. Sweltsa lyrata epiproct, Washington, Pend Oreille Co., Sullivan Creek. 47. Sweltsa mogollonica epiproct, New Mexico, Catron Co., Willow Creek.

Figs. 24-29. Sweltsa lyrata male epiproct, dorsal. 24. Montana, Mineral Co., St. Regis River. 25. Montana, Gallatin Co., Duck Creek. 26. Montana. Mineral Co. Trout Creek. 27. Washington, Pend Oreille Co., Sullivan Creek. 28. British Columbia, Duhamel Creek. 29. Yukon, Klondike River.

Figs. 18-23. Sweltsa lyrata male epiproct, dorsal. 18-19. Idaho, Bonner Co., Pack River. 20. Washington, Skagit Co., Friday Creek. 21. Washington, Whatcom Co., Padden Creek. 22-23. Alberta, Cameron Creek.