Chlaenius (Lithochlaenius) propeagilis Liu & Kavanaugh, 2011

Valim, Ye, Kavanaugh, David H., Shi, Hongliang & Liang, Hongbin, 2011, A key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species, ZooKeys 128, pp. 15-52 : 22-26

Chlaenius (Lithochlaenius) propeagilis Liu & Kavanaugh sp. n. Figs 14-15 20-21 54 68 82 98-99 121-122 148 158

Types.

Holotype: male (IZCAS),"Southwest Yunnan, Gongshan, Dulongjiang, 0.5km N of Dizhengdang, 28.08442, 98.32652 "/ "1880 m, 2004.10.29, David Kavanaugh Coll."/ "Holotype Chlaenius (Lithochlaenius) propeagilis Liu & Kavanaugh sp. n." [red label]. Paratypes: Total 452 specimens (IZCAS, CASC): 1 male, "China, Yunnan Provin. Gongshan County, Cikai town, along street, N27°44'43", E98°39'53" "/ "1500 m, 2006.5.5, Liang H.B, Ba Weidong"; 2 males, "China, Yunnan, Gongshan County, Cikai Township, Nu Jiang at Dashaba, N27.73845, E098.67092"/ "1430 m, 8-9 October 2002, Stop #DHK 2002-40, D.H. Kavanaugh, P.E. Marek & H.B. Liang collectors"; 17 males and 11 females, "China, Yunnan Provin. Gongshan County, Cikai town, Pulahe, N27°46'08", E98°39'12" "/ "1510 m, 2002.9.21-24, Liang Hongbin, Ba Weidong"; 1 male and 1 female, "China, Yunnan, Gongshan, Cikai Township, 3.3 airkm NW of Gongshan above hydropower diversion dam, 1530 m"/ "N27.77175, E098.64924, 24 September 2002, Stop # DHK 2002-028, D.H. Kavanaugh collector"; 2 males, "China, Yunnan Prov. Gongshan, Cikai Town, Pulahe joint of Nujiang, 27.74843N, 98.66498E "/ "1530m, 2004.10.23, D. Kavanaugh, Dong D.Z."; 6 males and 6 females, "China, Yunnan Provin. Gongshan County, Cilou (Power Station), N27°46'14", E98°39'16" "/ "1510m, 2002.5.6, Liang H.B, Ba Weidong, Yang Guodong, Li X.Q"; 5 males and 3 females, "China, Yunnan Provin. Gongshan County, Cikai town, Gazu Station, N27°44'35", E98°36'17" "/ "1600-1750 m, 2002.5.5, Liang H.B, Ba Weidong"; 2 males, "China, Yunnan Provin. Gongshan County, Cikai town to Qiqi Station, N27.43086, E98.34150"/ "1700-2000m, 2002.4.29, Liang Hongbin, Ba W.D."; 7 males and 26 females, "China, Yunnan, Gongshan, Cikai Township, 3.0 airkm N of Gongshan on Pula He at hydropower diversion dam, 1500 m"/ "N27.77055°, E098.65446°, 24 September 2002, Stop # DHK 2002-027, D.H. Kavanaugh, P.E. Marek & D.Z. Dong collectors"; 3 males and 4 females (CASC), "CHINA, Yunnan Province, Gaoligong Shan, Nujiang Prefecture, Gongshan County, Qiqi He just above hydroelectric plant, 1500m"/ "N27.75748°, E98.66073°, 22 July 2000, Stop #00-269, D.H. Kavanaugh, Liang H.-B., & Dong D.-Z. collectors"; 2 males and 4 females, "China, Yunnan, Gongshan, Bingzhongluo Township, 34km N of Gongshan at junction of Shuangla He and Nu Jiang, 1550m"/ "E27.96918°, E098.66198°, 25 September 2002, stop # DHK 2002-039, D.H. Kavanaugh, P.E. Marek & D.Z. Dong collectors"; 4 males and 6 females, "China, Yunnan Prov. Gongshan, Bingzhongluo, Shuangla He, beach, 27.96817N, 98.66187E "/ "1520 m, 2004.10.22, D. Kavanaugh collector"; 12 males and 10 females, "China, Yunnan, Gongshan, Bingzhongluo, Shuanglahe, riverside, N27°58'59", E98°39'15" "/ "1588 m, 2002.9.22-26, Liang H.B. Li Xiangqian"; 2 males and 5 females, "China, Yunnan, Gongshan, Bingzhongluo, Shuanglahe, riverside, N27°58'59", E98°39'15" "/ "1588 m, 2002.7.20, Ba Weidong"; 5 males and 2 female, "China, Yunnan Prov. Gongshan, Dulongjiang, 0.5 km N of Dizhengdang, 28.08442N, 98.32652E "/ "1880 m, 2004.10.29; D. Kavanaugh, Dong D.Z."; 7 males and 4 females, "China, Yunnan Provin. Gongshan County, Dulongjiang, Kongdang, headlamp, 27.87764N, 98.33618E "/ "1510m, 2006.8.27, David Kavanaugh"; 12 males and 17 females, "China, Yunnan Provin. Gongshan, Dulongjiang, Xianjiudang village, 27.94092N, 98.33340E "/ "1580m, 2004.11.4, D. Kavanaugh, Dong D.Z.";1 male, "China, Yunnan, Fugong County, Shangpa Town, west bank of Nu Jiang, 1185 m, N26.90668, E098.86339"/ "13 October 2002, Stop #DHK 2002-047, D.H. Kavanaugh, P.E. Marek & H.B. Liang, D.Z. Dong collectors"; 5 males and 3 females, "China, Yunnan Prov. Fugong, Shangpa Town, Nujiang, River, 26.90650N, 98.86397E "/ "1175 m, 2004.4.20, Liang H.B. coll."; 9 males and 4 females, "China, Yunnan Prov. Fugong, Shangpa Town, Beach of Nujiang, 26.90650N, 98.86397E "/ 1175 m, 2005.8.20, Liang H.B. Zhang J.F. Dong D.Z."; 7 males and 5 females, "China, Yunnan Prov. Fugong, Shangpa Town, Mugujia, riverside, 26.86203N, 98.87142E "/ 1177 m, 2005.8.22, Dong Dazhi collector"; 1 male and 2 females, "China, Yunnan Prov. Fugong, Shangpa, Mugujia, round waterfall, 26.86203N, 98.87142E "/ 1177 m, 2005.8.22, Liang H.B. Zhang J.F."; 1 female, "China, Yunnan, Fugong, Lumadeng Township, 2km airkm S of Aludi on Nu Jiang, 1245 m, N27.09037, E098.87359"/ "20 September 2002, Stop # DHK 2002-022, D.H. Kavanaugh & H.B. Liang collectors"; 1 female, "China, Yunnan Prov., Fugong, Pihe at junction of Nujiang River, 26.53177N, 98.89753E "/ "1060 m, 2004.4.20, D. Kavanaugh, C. Griswold"; 2 males and 6 females, "China, Yunnan, Lushui, Liuku, west bank of Nu Jiang, 960 m, N25.854, E098.852/ "15 October 2002, stop # DHK 2002-050, D.Z. Dong collector"; 1 male and 2 females, "China, Yunnan Provin. Lushui, Beach of Nujiang River, under stone, N25°51'20", E98°50'58" "/ 800 m, 2002.9.19, Liang Hongbin"; 2 males and 2 females (CASC), "CHINA, Yunnan Province, Gaoligongshan Mountains, Nujiang Prefecture, Lushui County, Salween River, 17 km N of Liuku, 970m"/ " 25°58.7'N, 98°50.4'E, 21 October 1998, Stop #98-120, D.H. Kavanaugh & C.-L. Long collectors"; 2 females (CASC), "CHINA, Yunnan Province, Nujiang Prefecture, Lushui County, Liuku Township, Liuku, 800m"/ " 25.86010°N, 98.85155°E, 25-26 June 2000, Stop #00-7, D.H. Kavanaugh & Liang H.-B. collectors"; 4 males and 5 females (CASC), "CHINA, Yunnan Province, Nujiang Prefecture, Lushui County, San jiang Township, Nu Jiang, 790m"/ " 25.72964°N, 98.87180°E, 26 June 2000, Stop #00-9, D.H. Kavanaugh & Liang H.-B. collectors"; 7 male and 7 females, "China, Yunnan Prov. Baoshan, Longyang, Bawan, Dongfengqiao, 24.98742N, 98.87047E "/ "670 m, 2005.6.1, D. Kavanaugh, Dong D.Z."; 15 male and 16 females, "China, Yunnan Prov. Baoshan, Longyang, Bawan, Dongfengqiao, 24.98535N, 98.87382E "/ "670 m, 2005.5.29-6.1, Liang H.B. Dong D.Z."; 1 male, "China, Yunnan Prov. Tengchong, Mangbang, Longwenqiao, 25.02329N, 98.67710E "/ "1290 m, 2006.6.5, Liang H.B. Hu P."; 6 males and 7 females, "China, Yunnan Prov. Tengchong, Mangbang, Longwenqiao, beach, 25.02396N, 98.67675E "/ "1285 m, 2006.6.5, David Kavanaugh"; 3 males and 1 female, "China, Yunnan Province, Tengchong, Shangying, N25°02'29.7", E98°40'22.9" " / "1335 m, 2003.10.19, Liang H.B, Shi X.C."; 2 males and 8 females, " China, Yunnan Prov. Tengchong Co., Wuhe Town, Longjiangqiao, 24.89499N, 98.67510E "/ "1205 m, 2005.V.24, Kavanaugh D. Dong D.Z."; 1 female, "China, Yunnan Province, Tengchong, Wuhe Township, Longjiang Bridge on Longchuanjiang"/ " N24.89889, E098.67667, 1215 m, 28 October 2003, under rocks, Dong D.Z. collector"; 17 males and 9 females, "China, Yunnan Prov. Tengchong, Wuhe, Longjiangqiao, beach, 24.89176N, 98.67551E "/ "1230 m, 2006.6.3, Kavanaugh D. Brett R."; 4 males and 4 females, "China, Yunnan Prov. Tengchong Co., Wuhe Town, Longjiangqiao, 24.89284N, 98.67439E "/ "1210 m, 2005.V.24, Liang H.B. Yang J.L"; 1 male, "China, Yunnan Prov. Tengchong, Wuhe, Longjiangqiao, beach, 24.89293N, 98.67489E "/ "1220 m, 2006.6.3, Liang H.B. Hu P."; 3 males, "China, Yunnan Province, Tengchong Co. Qushi Township, Shuang He Cun, N25.32555, E098.60861 "/ "1464 m, 21 October 2003, under rocks, Dong D.Z. collector"; 2 males, "China, Yunnan Prov. Tengchong, Qushi, Xiangyangqiao, beach, 25.21221N, 98.57836E "/ "1500 m, 2006.V.24, David Kavanaugh"; 2 females, "China, Yunnan Prov. Tengchong, Qushi, Xiangyangqiao, beach, 25.23939N, 98.62723E "/ "1440 m, 2006.V.24, David Kavanaugh"; 11 males and 14 females (CASC), "CHINA, Yunnan Province, Gaoligongshan Mountains, Baoshan Prefecture, Tengchong County, Longchuan River at Longkou village"/ " 25°16.9'N, 98°35.5'E, 1500 m, 25 October 1998, Stop #98-126, D.H. Kavanaugh, C.E. Griswold, & C.-L. Long collectors"; 6 males and 1 female, "China, Yunnan Prov. Tengchong, Qushi, Longkou, beach, 25.28175N, 98.59246E "/ "1500 m, 2006.6.6, David Kavanaugh"; 5 males and 1 female (CASC), "CHINA, Yunnan Province, Gaoligongshan Mountains, Baoshan Prefecture, Tengchong County, Longchuan River at Longkou village"/ " 25°16.9'N, 98°35.5'E, 1500 m, 2 November 1998, Stop #98-128, D.H. Kavanaugh, C.E. Griswold, C.-L. Long, R. Li, & H.-X. He collectors"; 3 males and 5 females, "China, Yunnan Prov. Tengchong, Qushi, Qinqiao, 25.27236N, 98.60093E "/ "1460 m, 2006.6.2-6, David Kavanaugh, Brett R."; 4 males and 4 females, "China, Yunnan Province, Tengchong, Zhoujiapo Village, N25.33222, E098.67611 "/ "1740 m, 24 October 2003, under rocks Dong D.Z. collector"; 3 males, "China, Yunnan Prov. Tengchong, Jietou, Yonganqiao, beach, 25.32502N, 98.70459E "/ "1470 m, 2006.V.24, Liang H.B."; 2 males and 1 female, "China, Yunnan Prov. Tengchong, Jietou, Yonganqiao, beach, 25.32504N, 98.60959E "/ "1470 m, 2006.5.24, Kavanaugh D. Brett R."; 1 male, "China, Yunnan Prov. Tengchong, Hehua, Dengma, on beach, 24.92346N, 98.38612E "/ "1105 m, 2006.6.2, Kavanaugh D. Brett R."; 3 males and 1 female, "China, Yunnan Province, Jingping Co., Mengla Town, Mengla-daqiao, River side 22˚39'45.7",103˚04'44.7" "/ "312m, 2003.12.15, day, Liang H B, Boris Kataev Colls."; 12 males and 10 females (IZCAS), "Yunnan, Jingdong, 1100m"/ "1982.IV.29-V.2, Yu P.Y.& Liao S.B."; 1 female, "Yunnan, Honghe Prefecture, Lvchun, Huanglianshan, 2009.V. 16, Bai X. X. coll." 2 males and 1 female, "Yunnan, Menglun, Xishuanbanna Botanical Garden", "2005.5.22, light trap, Zheng Guo leg."; 3 males and 5 females, "China, Sichuan, Yajiang, Hekou Town, Shabeihou, Yalongjiang, N30.00020, E101.01526 "/ "2583m, 2009.5.27-28, Liang H.B."; 1 female (IZCAS), "China, Tibet, Bomi Yi"ong, Tangmai bridge, Beach of Yi"ong Zangbo, 30.09633N, 95.06577E "/ "2035m, 2006.8.30 N, Liang H.B., Song Z.S.". Each paratype with an additional yellow label: "Paratype Chlaenius (Lithochlaenius) propeagilis Liu & Kavanaugh sp. n."

Diagnosis.

Antennomere 1 elongate ovoid ( Fig. 54 ); color of antennomeres 1-3 lighter than antennomeres 4-11; intervals 1-5 almost glabrous medially, with one or two rows of pubescence at each lateral side, more evenly pubescent near apex ( Fig. 82 ); apical lamella of aedeagus moderately triangular ( Fig. 121 ), thick and reflexed in lateral view ( Fig. 122 ).

Description.

Total length = 15.50-17.00 mm (mean = 15.71), width = 5.60-6.13 mm (mean = 5.81); HW = 2.75-2.95 mm (mean = 2.82), EYL = 1.00-1.15 mm (mean = 1.09), ratio Ant3/Ant1 = 1.73-1.93 (mean = 1.81), PL/PW = 0.78-0.85 (mean = 0.83), EL/EW = 1.62-1.79 (mean = 1.73), EL/ PL = 1.29-1.45 (mean = 1.36).

Head and pronotum black with green or coppery metallic luster; elytra black; ventral surface black, with some slight metallic reflection; coxae almost black; mandibles dark brown; antennomeres 4-11, palpomere, trochanters, tibiae at both ends, and tarsomeres reddish or brown; antennomeres 1-3, femora and middle portions of tibiae yellow or yellowish-brown.

Head rugose near eyes and occiput, vertex sparsely rugose; eyes prominent; antennomere 1 elongate-ovoid ( Fig. 54 ); labrum concave at apex, with distinct microsculpture; mentum tooth emarginate apically.

Pronotum cordate, moderately convex; lateral margins bordered; disk almost smooth, sparsely rugose and setose near base and lateral margins; basal foveae moderately deep.

Elytra widest at mid-length; humeral angle obtuse ( Fig. 68 ); striae deep, punctate; intervals moderately convex, intervals 1-5 almost glabrous medially, with a row of pubescence laterally, intervals more evenly pubescent near apex ( Fig. 82 ); intervals 6-9 pubescent throughout and with intervals 7-9 more densely pubescent than 6.

Abdominal sterna bordered, densely pubescent laterally; sterna IV-VI sparsely pubescent medially; sternum VII rugose-pubescent, with apex narrowly rounded or subobtuse in males ( Fig. 98 ), more broadly rounded in female ( Fig. 99 ).

Apical lamella of aedeagus moderately triangular ( Fig. 121 ), thick and bent ventrally ( Fig. 122 ). Gonostylus ( Figs 14-15 ).

Etymology.

The Latinized name propeagilis refers to the similarity of members of this species to those of Chlaenius agilis .

Geographical distribution.

Fig. 158 . Known only from southeastern Tibet and Yunnan Province, China.

Remarks.

Mensural data cited in the description were based on measurements obtained from 5 males and 5 females selected for maximum variation.

Specimens of Chlaenius (Lithochlaenius) collected from western Yunnan were initially determined as Chlaenius agilis Chaudoir. However, after comparison of the ma le genitalia with those of type specimens of Chlaenius agilis , we are convinced that they represent a distinct new species. Males differ from those of Chlaenius agilis in having the median lobe of aedeagus gradually bent near the base in lateral view ( Fig. 122 ), whereas Chlaenius agilis males have the median lobe abruptly bent with a depression near the base ( Fig. 124 ).

This species is clearly very closely related to Chlaenius agilis . At present, the known ranges of these two species are broadly disjunct ( Fig. 158 ). We know of no locality records for any Chlaenius (Lithochlaenius) species from the intervening area (i.e., between northcentral India and southeastern Tibet and western Yunnan Province. Whether this distributional gap represents a real disjunction or is only an artifact of inadequate collecting in the area to date can only be determined from additional sampling efforts in the region. It would be particularly informative to determine whether or not any populations representing the propeagilis/agilis lineage occur in the region and, if so, whether or not males display intermediate genitalic traits.

Figures

Figure 158. Map showing the known geographical distributions of Chlaenius (Lithochlaenius) spp.

Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view.

Figures 16 - 21. Habitus of Chlaenius (Lithochlaenius) spp 16 Chlaenius chuanqianensis sp. n., holotype, male, dorsal view 17 Chlaenius chuanqianensis sp. n., holotype, male, ventral view 18 Chlaenius linwensini sp. n., holotype, male, dorsal view 19 Chlaenius linwensini sp. n., holotype, male, ventral view 20 Chlaenius propeagilis sp. n., holotype, male, dorsal view 21 Chlaenius propeagilis sp. n., holotype, male, ventral view. Scale line = 5.0 mm.

Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm.

Figures 66 - 79. Humeral region of elytra of Chlaenius (Lithochlaenius) spp 66 Chlaenius chuanqianensis sp. n., holotype 67 Chlaenius linwensini sp. n., holotype 68 Chlaenius propeagilis sp. n., holotype 69 Chlaenius agilis Chaudoir, holotype 70 Chlaenius formosensis Lorenz, CCCC 71 Chlaenius leishanensis Kirschenhofer, IZCAS 72 Chlaenius noguchii Bates, IZCAS 73 Chlaenius wrasei Kirschenhofer, IZCAS 74 Chlaenius agiloides Jedlicka, IZCAS 75 Chlaenius rambouseki Lutshnik from Ussuri region, ZCAS 76 Chlaenius anchomenoides Bates, paratype 77 Chlaenius nuristanus Jedlicka, paratype 78 Chlaenius nuristanus Jedlicka, paratype 79 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm.

Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm.

Figures 94 - 116. Abdominal sternum VII (ventral aspect) of Chlaenius (Lithochlaenius) species 94 Chlaenius chuanqianensis sp. n., male, holotype 95 Chlaenius chuanqianensis sp. n., female, paratype 96 Chlaenius linwensini sp. n., male, holotype 97 Chlaenius linwensini sp. n., female, paratype 98 Chlaenius propeagilis sp. n., male, holotype 99 Chlaenius propeagilis sp. n., female, paratype 100 Chlaenius agilis Chaudoir, male, holotype 101 Chlaenius formosensis Lorenz, male 102 Chlaenius formosensis Lorenz, female 103 Chlaenius leishanensis Kirschenhofer, male 104 Chlaenius leishanensis Kirschenhofer, female 105 Chlaenius noguchii Bates, male 106 Chlaenius noguchii Bates, female 107 Chlaenius wrasei Kirschenhofer, male 108 Chlaenius wrasei Kirschenhofer, female 109 Chlaenius agiloides Jedlicka, male 110 Chlaenius agiloides Jedlicka, female 111 Chlaenius rambouseki Lutshnik, from Ussuri region, male 112 Chlaenius rambouseki Lutshnik from Ussuri region, female 113 Chlaenius anchomenoides Bates, male, paratype 114 Chlaenius nuristanus Jedlicka, male, paratype 115 Chlaenius nuristanus Jedlicka, female, paratype 116 Chlaenius formosanus Jedlicka from Taiwan, male. Scale line = 1.0 mm.

Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130).

Figures 146 - 151. Photographs of habitats for Chlaenius (Lithochlaenius) spp. 146 Chlaenius chuanqianensis sp. n., Dabaitang, Xishui county, North Guizhou, China 147 Chlaenius anchomenoides Bates, Goorais valley, Pakistan (Provided by Dr. Muhammad Abbas in Pakistan Museum of Natural History) 148 Chlaenius propeagilis sp. n., Gaoligongshan, Yunnan, China 149 Chlaenius leishanensis Kirschenhofer, Xiaodanjiang, Leigongshan, Leishan county, Southeast Guizhou, China 150 Chlaenius chuanqianensis sp. n. and Chlaenius agiloides Jedlicka, Jinshajiang, Sichuan 151 Chlaenius rambouseki Lutshnik, Ussri river, Heilongjiang, China.