Achanius Erichson, 1847

Flores, Gustavo E. & Aballay, Fernando H., 2015, Two Evaniosomini Species (Coleoptera: Tenebrionidae) Associated with Decaying Carcasses in Argentina, with Remarks on the Tribal Assignment of Achanius Erichson, The Coleopterists Bulletin (mo 14) 69, pp. 167-179 : 173-174

Genus Achanius Erichson, 1847

( Figs. 13–20 , 22 )

Achanius Erichson 1847: 118 . Lacordaire 1859: 82; Gemminger and Harold 1870: 1842 (catalog); Gebien 1910: 30 (catalog), 1937: 579 (catalog); Blackwelder 1945: 511 (catalog); Kulzer 1950: 30 (revision), 1956: 898 (revision); Peña 1966: 404 (catalog); Doyen 1994: 500.

Type species: Achanius anthicoides Erichson, 1847 , by monotypy.

Description. In addition to the characters stated above: Mandibles finer, acute, base 1.5X wider than apex, with dorsal cusp only on right mandible on basal half, left mandible with sharp, raised dorsal edge ( Fig. 17 ), both mandibles with ventral fossa ( Fig. 18 ); labrum not exposed, entirely membranous; clypeus oval, produced anteriorad of antennal insertions, covering labrum, anterior margin triangular or concave and armed with row of teeth directed downwards, apically with a large tooth prolonged anteriorad, genoclypeal suture wellmarked ( Fig. 17 ), epicanthus separated from clypeus ( Fig. 19 ); prementum entirely membranous, nearly concealed beneath mentum; anterior margin of mentum straight or slightly concave ( Fig. 18 ); eyes not protruding, upper margin with slightly or wellmarked carina always present, starting posterior to epicanthus and ending in posterior margin of eye; length of epicanthus equal to length of eye, head widest at eyes, postgenal margin inconspicuous ( Fig. 19 ); antennae surpassing posterior margin of pronotum, all antennomeres longer than wide, antennomere 1 longer than 3, emerging from distal ¾ of antennal insertion ( Fig. 19 ), antennomere 2 equal to or longer than 3 or shorter than 3 but almost equal (proportion 2/3 = 17/19) ( Fig. 19 ); antennomeres 9 and 10 triangular, expanded outwards, longer than wide ( Figs. 13, 14 ). Pronotum with anterior angles rounded, sloping towards venter. Metacoxae transverse, separated by less than third of metacoxal width ( Fig. 20 ). First tarsomere of hind legs longer than 2 + 3 combined ( Figs. 14, 16 ). Female genitalia as in Fig. 22 . Spiculum without arms. Paraprocts long (2.0 <P/C ≤ 3.0), basal lobe of coxite not extended over paraproct. Vagina saccate. Spermathecal accesory gland longer than vagina, duct not annulate. Spermatheca single, compact, locular, emptying at end of vagina, spermathecal accessory gland emptying inside spermatheca.

Material Examined. Syntype of A. (Achanius) anthicoides (MNHUB) ( Fig. 13 ); three specimens of Achanius (Achanius) bembidioides Fairmaire (1 NHMB, 2 IADIZA); six specimens of Achanius (Achanius) minutus Kulzer (IADIZA) ; holotype ( MNHN) and 113 specimens ( IADIZA) of Achanius (Ambigatus) rufonitens Fairmaire ; holotype ( NHMB) and three specimens ( IADIZA) of A. (Ambigatus) stricticollis ; holotype ( NHMB) and 39 specimens ( IADIZA) of Achanius (Ambigatus) bicolor Kulzer ; holotype of Achanius (Ambigatus) rhinosomoides Kulzer ( NHMB) ; paratype of Achanius (Ambigatus) piceus Kulzer (IADIZA) ; holotype of Achanius (Ambigatus) wittmeri Kulzer ( NHMB) ( Fig. 16 ); holotype, allotype, and nine paratypes of A. (Ambigatus) antofagastensis Flores and Aballay, new species.

Figures

Figs. 13–16. Dorsal habitus. 13) Achanius (Achanius) anthicoides, syntype (Image courtesy of Bernd Jaeger, MNHUB); 14) Achanius (Ambigatus) antofagastensis, holotype; 15) A. antofagastensis, paratype showing reduced wings (white); 16) Achanius (Ambigatus) wittmeri, holotype (Image courtesy of Eva Sprecher, NHMB. Scale bars = 1 mm.

Figs. 17–20. Achanius (Ambigatus) bicolor. 17) Mouthparts, dorsal view; 18) Mouthparts, ventral view, white arrow indicates ventral fossa on mandible; 19) Head, frontal view; 20) Meso- and metasterna.

Figs. 21–22. Ovipositor in ventral view, spiculum and internal female reproductive tract. 21) Vaniosus profana; 22) Achanius (Ambigatus) rufonitens. Abbreviations: c = coxite; o = oviduct; p = paraproct; s = spermatheca; sag = spermathecal accessory gland; sp = spiculum; v = vagina. Scale bar = 1 mm.