Enoggerinae Burks, new subfamily, 2022

Burks, Roger, Mitroiu, Mircea-Dan, Fusu, Lucian, Heraty, John M., Jansta, Petr, Heydon, Steve, Papilloud, Natalie Dale-Skey, Peters, Ralph S., Tselikh, Ekaterina V., Woolley, James B., van Noort, Simon, Baur, Hannes, Cruaud, Astrid, Darling, Christopher, Haas, Michael, Hanson, Paul, Krogmann, Lars & Rasplus, Jean-Yves, 2022, From hell's heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera), Journal of Hymenoptera Research 94, pp. 13-88 : 13

Enoggerinae Burks, new subfamily incertae sedis

Type genus.

Enoggera Girault, 1926.

Diagnosis.

Antenna with 9-12 flagellomeres, including either an incompletely divided clava or up to 4 clavomeres, sometimes including a small 4th clavomere. Temple absent, thus posterior margin of eye coincident with the posterior margin of the head dorsally (Fig. 65 ). Clypeus with transverse subapical groove. Labrum exposed, well-sclerotized, subrectangular, with marginal setae in a row. Mandibles with 3 teeth. Subforaminal bridge with postgena separated by lower tentorial bridge. Mesoscutellum with frenum indicated at least laterally, with axillular sulcus. Mesopleural area without an expanded acropleuron, with pits; mesepimeron not extending over anterior margin of metapleuron; two mesofurcal pits present. All legs with 5 tarsomeres; protibial spur stout and curved; basitarsal comb oblique. Metasoma with syntergum, therefore without epipygium. Egg parasitoids of Chrysomelidae ( Coleoptera ).

Discussion.

Enoggerinae share a pair of mesofurcal pits with a number of other chalcidoid groups, including Herbertiidae , Moranilidae , and Asaphesinae . However, these groups are unstable in molecular analyses and do not form a clade (Cruaud et al., submitted), with Enoggerinae more often as the sister group of Coelocybidae . Therefore, Enoggerinae can be separated from all similar groups by the absence of the temple. With a different biology, Enoggerinae would also represent a discordant element if placed in any of the other groups.

Figures

Figures 61 - 66. 61 Austrosystasis atricorpus Girault (Austrosystasinae, incertae sedis): mesosoma lateral view 62 - 64 Ditropinotella sp. (Ditropinotellinae, incertae sedis) 62 antenna 63 mesosoma lateral view 64 metasoma 65 Enoggera reticulata Naumann (Enoggerinae, incertae sedis): mesosoma lateral view 66 Eopelma sp. (incertae sedis): apex of mesotibia and mesotarsus.