Lasianthus griffithii var. scabridus, 2012

Zhu, H., Roos, M. C. & Ridsdale, C. E., 2012, A taxonomic revision of the Malesian species of Lasianthus (Rubiaceae), Blumea 57 (1), pp. 1-102 : 46

b. var. scabridus (King & Gamble) H.Zhu , stat. nov.

Lasianthus scabridus King & Gamble,J.Asiat.Soc.Bengal 73 (1904) 111;Ridl. (1923a) 152; K.M. Wong (1989) 370. —

Typus: Ridley 6463 ( lecto SING, designated by Wong 1988), Peninsular Malaysia, Johore .

Branchlets terete, robust, glabrous, dark-brown when dry. Leaves: blades oblong-lanceolate to obovate-lanceolate, with stiff hairs on both sides; nervules reticulate, conspicuously prominent beneath. Stipules big, partly covering the cymes, ovate to triangular-ovate or broadly triangular, 1–2 by 1– 2 cm, coriaceous with membranaceous margin, glabrous. Inflorescences sessile; bracts numerous, linear, hirsute, forming a dense head. Flowers sessile, sealed (hidden) by bracts.

Distribution — Malaysia (Peninsular, Borneo), Indonesia ( Sumatra, Kalimantan).

Note — Lasianthus scabridus differs from L. griffithii in having stiff hairs on both sides of leaves, reticulate nervules conspicuously prominent beneath. It is suitable to sink L. scabridus to a variety of L. griffithii as var. scabridus .