Geholaspis (Longicheles), Valle, 1953

Bolger, Thomas, Arroyo, Julio & Piotrowska, Katarzyna, 2018, A catalogue of the species of Mesostigmata (Arachnida, Acari, Parasitiformes) recorded from Ireland including information on their geographical distribution and habitats, Zootaxa 4519 (1), pp. 1-220 : 124-135

Subgenus Longicheles Valle, 1953

• Geholaspis (Longicheles) mandibularis (Berlese, 1904)

• Original name: Holostaspis mandibularis Berlese, 1904 .

• In: Redia , 1: 263.

• Combinations/variations used in Irish literature: Holostaspis longulus Berlese ( Halbert, 1915) , Macroche-

les longulus Berlese ( Hull, 1918) .

Irish Records: Moss, rotten wood ( Halbert, 1915); lake shore and seashore ( Halbert, 1915); pasture ( Valarasan, 1971); pasture, bracken best and Salix ditch at seashore ( Purvis, 1982); moss, flood debris, shallow creek, roadside ditch, between rocks, base of sycamore, spruce litter, rotten grass ( Hyatt & Emberson, 1988; Bolger, 1990); under stones, broadleaved forest soil ( Arroyo et al., 2009); nest of Lasius flavus and pasture ( Arroyo et al., 2015); soil in winter wheta field (unpublished data); on cliff, in woodland, moss and rough grassland (Bolger, in press); fungal sporocarp ( O’Connell, 1994, unpublished data); ash woodland (unpublished data); moss on soil under Scots pine and Sitka spruce, canopy of ash and, bark of Sitka spruce ( Arroyo et al., 2012, 2013a); soil under birch and Lawson cypress (unpublished data); fen and raised bog ( Wisdom, 2010; Wisdom et al., 2011).

Counties: Carlow, Clare, Dublin, Galway, Kildare, Leitrim, Limerick, Mayo, Offaly, Roscommon, Sligo, Waterford, Westmeath, Wexford, Wicklow.

Glyptholaspis Filipponi & Pegazzo, 1960

• Glyptholaspis americana (Berlese, 1888)

• Original name: Holostaspis marginatus var. americanus Berlese, 1888 .

• In: Bollettino della Societa Entomologica Italiana. Rome, 20: 195.

• Combinations/variations used in Irish literature: Macrocheles vagabundus (Berlese) ( Halbert, 1923) . Records: Under wood on seashore ( Halbert, 1923); straw ( Hyatt & Emberson, 1988; Hyatt, 1990b). Counties: Clare, Dublin.

Holostaspella Berlese, 1904

• Holostaspella ornata (Berlese, 1904)

• Original name: Holostaspis ornatus Berlese, 1904 .

• In: Redia , 1: 421.

Irish Records: Pasture ( Ganley, 1976).

Counties: Dublin.

• Holostaspella subornata Bregetova & Koroleva, 1960

• Original name: Holostaspella subornata Bregetova & Koroleva, 1960 .

• In: Parasitologicheskii Sbornik. Leningrad, 19: 60.

Irish Records: Grass roots ( Hyatt & Emberson, 1988; Bolger, 1990).

Counties: Clare.

Macrocheles Latreille 1829

• Macrocheles carinatus (Koch, 1839)

• Original name: Gamasus carinatus Koch, 1839 .

• In: Deutschlands Crustaceen, Myriopoden und Arachniden. 1835–1844. Regensburg, fasc. 24 (16).

Irish Records: Stored foodstuffs ( O’Farrell & Butler, 1948); manure on soil surface pasture ( Curry, 1969a, 1979; Cunningham, 1975; Ganley, 1976; Purvis, 1978; Tuohy, 1979; Bolger, 1980; O’Neill, 1981; Arroyo et al., 2015); winter wheat crop ( Kelly & Curry, 1985; unpublished data); in liverworts, mosses and grasses, rotten wood, flood debris ( Hyatt & Emberson, 1988; Bolger, 1990); ash woodland and pasture (unpublished data); soil under Norway spruce ( Heneghan, 1994); soil under oak (unpublished data).

Counties: Antrim, Carlow, Clare, Derry, Down, Dublin, Kildare, Kilkenny, Leitrim, Limerick, Meath, Westmeath, Wexford.

• Macrocheles decoloratus (Koch, 1839)

• Original name: Gamasus decoloratus Koch, 1893

• In: Deutschlands Crustaceen, Myriopoden und Arachniden. 1835–1844. Regensburg, fasc.25 (14).

Irish Records: Sand martin ( Riparia riparia ) nest ( Hyatt & Emberson, 1988; Bolger, 1990).

Counties: Cork.

• Macrocheles dentatus ( Evans & Browning, 1956)

• Original name: Macrholaspis dentatus Evans & Browning, 1956 .

• In: Bulletin of the British Museum (Natural History) Zoology, 4: 49.

Irish Records: Salix ditch at seashore ( Purvis, 1982).

Counties: Wexford.

• Macrocheles glaber (Müller, 1860)

• Original name: Holostaspis glabra Müller, 1860 .

• In: Jahresheft der Naturwissenschaftlieben Section der k. k. magr. schles. Gesellschaft fur Ackerbau, Nature - und Landeskunde, 1860: 178.

• Combinations/variations used in Irish literature: Holostaspis marginatus (Hermann) var littorali s Halbert (Halbert, 1915), Macrocheles marginatus Hermann ( Hull, 1918) .

Irish Records: Under stones and refuse on upper seashore ( Halbert, 1915, 1920); pasture ( Curry, 1969a, 1976a; Valarasan, 1971; Bolger, 1980); manure on soil surface (Curry, 1979); soil of sugar beet crop ( Breslin, 1979); winter wheat crop ( Kelly & Curry, 1985); seaweed, compost heap, rotting hay, on Geotrupes stercorarius , in meadow ( Hyatt & Emberson, 1988; Bolger, 1990); soil under Norway spruce ( Heneghan, 1994); woodland (Bolger, in press).

Counties: Carlow, Clare, Cork, Dublin, Galway, Kildare, Kilkenny, Mayo, Meath, Wexford.

• Macrocheles matrius (Hull, 1925)

• Original name: Nothrholaspis matrius Hull, 1925 .

• In: Annals and Magazine of Natural History London, 9/15: 212.

Irish Records: Farm grain stores ( Cusack, 1974; Cusack et al., 1975).

Counties: Not specified.

• Macrocheles montanus (Willmann, 1951)

• Original name: Nothrholaspis montana Willmann, 1951 .

• In: Bonner zoologische Beiträge, 2: 158.

Irish Records: Roadside ditch ( Hyatt & Emberson, 1988; Bolger, 1990); soil under Norway spruce ( Heneghan, 1994); nest of Lasius flavus ( Arroyo et al., 2015) .

Counties: Kilkenny, Limerick, Westmeath.

• Macrocheles muscaedomesticae (Scopoli, 1772)

• Original name: Acarus muscae domesticae Scopoli, 1772 .

• In: Annali V. Historico -Naturalis, 125: 127

Irish Records: Garden compost heap, mixture of bedding and faeces from dog houses ( Bolger, 1990).

Counties: Dublin.

• Macrocheles opacus (Koch, 1839)

• Original name: Gamasus opacus Koch, 1839 .

• In: Deutschlands Crustaceen, Myriopoden und Arachniden. 1835–1844. Regensburg, fasc. 25 (24).

• Combinations/variations used in Irish literature: Macrholaspis opacus (Koch) ( Curry, 1969a; Whelan, 1976).

Irish Records: Pasture ( Curry, 1969a; Valarasan, 1971; unpublished data); pasture on peat ( Whelan, 1976); pine shelter bed on bog ( Longworth, 1973); dune, bracken bed, gorse bank and Salix ditch at seashore ( Purvis, 1982); moss on wall and rotten stump, mossy turf and roadside ditch ( Hyatt & Emberson, 1988; Bolger, 1990); soil under Norway spruce ( Heneghan, 1994); broadleaved forest soil ( Arroyo et al., 2009); nest of Lasius flavus and pasture ( Arroyo et al., 2015); Sphagnum (Bolger, in press); fungal sporocarp ( O’Connell, 1994; unpublished data); ash and spruce woodland ( Little & Bolger, 1995; unpublished data); soil under birch (unpublished data).

Counties: Carlow, Clare, Dublin, Galway, Kildare, Kilkenny, Limerick, Mayo, Westmeath Wexford, Wicklow.

• Macrocheles penicilliger (Berlese, 1904)

• Original name: Holostaspis penicilliger Berlese, 1904 .

• In: Redia , 1: 264.

Irish Records: Pasture ( Cunningham, 1975; Curry, 1976a; Tuohy, 1979; Bolger, 1980; O’Neill, 1981); pasture on peat ( Whelan, 1976); manure on soil surface (Curry, 1979); Sphagnum , pine shelter belt and pasture on bog ( Longworth, 1973); Burren ( Hyatt & Emberson, 1988; Bolger, 1990); soil under Norway spruce ( Heneghan, 1994).

Counties: Clare, Dublin, Kildare, Kilkenny, Mayo, Meath, Wexford.

• Macrocheles scutatus (Berlese, 1904)

• Original name: Holostaspis subbadius var. scutatus Berlese, 1904 .

• In: Redia , 1: 264.

Irish Records: Soil and vergetation in saltmarsh ( Healy, 1975; & Healy, 1977).

Counties: Dublin.

• Macrocheles subbadius (Berlese, 1904)

• Original name: Holostaspis subbadius Berlese, 1904 .

• In: Redia , 1: 264.

Irish Records: Compost heap ( Hyatt & Emberson, 1988; Bolger, 1990).

Counties: Clare.

• Macrocheles submotus Falconer, 1924

• Original name: Macrocheles submotus Falconer, 1924 .

• In: Naturalist, London, 1923: 152.

• Combinations/variations used in Irish literature: Holostaspis tridentinus (G. & R. Canestrini ( Halbert,

1915), Macrocheles tridentinus Hermann ( Hull, 1918) .

Irish Records: Moss and under bark ( Halbert, 1915); manure on soil surface (Curry, 1979); bracken bed at seashore ( Purvis, 1982); pasture and broadleaved forest soils ( Arroyo et al., 2009); pasture ( Valarasan, 1971; Arroyo et al., 2015); pasture on peat ( Whelan, 1976); moss, rough grassland, saltmarsh, cliff (Bolger, in press); fungal sporocarp ( O’Connell, 1994; O’Connell & Bolger, 1997b; unpublished data); small mammal nests (unpublished data); ash woodland (unpublished data); base of sycamore tree, compost heap, moss and liverworts on soil and trees, flood debris, turf, rotten grass, roadside ditch ( Hyatt & Emberson, 1988; Bolger, 1990); soil under oak and Scots pine; moss on soil under oak and Sitka spruce; moss on ash, Scots pine; canopy of ash ( Arroyo et al., 2010, 2012, 2013a); riverbank (unpublished data); mountain blanket bog ( Wisdom, 2010; Wisdom et al., 2011).

Counties: Clare, Dublin, Galway, Kerry, Kildare, Kilkenny, Laois, Leitrim, Limerick, Mayo, Meath, Offaly, Sligo, Waterford, Westmeath, Wexford, Wicklow.

• Macrocheles tardus (Koch, 1841)

• Original name: Gamasus tardus Koch, 1841 .

• In: Deutschlands Crustaceen, Myriopoden und Arachniden. 1835–1844. Regensburg, fasc. 39 (14).

Irish Records: Flood debris, roadside ditch and rotten grass ( Hyatt & Emberson, 1988).

Counties: Leitrim, Westmeath.

• Macrocheles terreus (Canestrini & Fanzago, 1877)

• Original name: Gamasus terreus Canestrini & Fanzago, 1877 .

• In: Atti del Reale Istituto veneto di scienze, lettere ed arti, Series 5, 4: 48.

Irish Records: In fungus, under bark, and in moss ( Halbert, 1915).

Counties: Dublin, Mayo.