Lichtwardtia nodulata Grootaert & Tang

Tang, Chufei, Yang, Ding & Grootaert, Patrick, 2018, Revision of the genus Lichtwardtia Enderlein in Southeast Asia, a tale of highly diverse male terminalia (Diptera, Dolichopodidae), ZooKeys 798, pp. 63-107 : 63

Lichtwardtia nodulata Grootaert & Tang sp. n. Figs 6, 7

Material examined.

Holotype male (coll. LKCNHM): SINGAPORE: Semakau, 30 August 2012, sweep netting (leg. Jayanthi Puniamoorthy & P Grootaert). Paratypes (all coll. LKCNHM): SINGAPORE: 1 male, Pulau Ubin, 14 July 2012, sweep netting in mangrove (leg. Jayanthi Puniamoorthy & P Grootaert); 9 males, 14 females, Kranji nature trail, 27 July 2005, sweep netting in park (leg. P Grootaert); 1 male, Semakau, 13 December 2012, sweep netting in halophilous vegetation on sandy beach (leg. P Grootaert).

Diagnosis.

Basal 7/8 of mid coxa and basal half of hind coxa brown, mid coxa with one black band at middle. Vein R1 with one oval thickness at where connect to C. Phallus with two rows of black denticles on ventral half, anterior row only with four small sparse denticles, posterior row with eight dense denticles.

Description.

Male.Body length 3.6-3.8 mm, wing 3.2-3.3 × 1.0 mm.

Head dark metallic green, with thick pale pollinosity; face slightly raised, frons and face both with thick pale pollinosity, gradually narrowed downward. Hairs and bristles on head black except posteroventral hairs pale. Antenna dark yellow; postpedicel nearly triangular, blunt apically, nearly as long as wide, with short black pubescence; arista-like stylus dorsal, inserted at basal half of postpedicel, nearly as long as head width, black, feather-like, with long black pubescence, basal segment 0.3 times as long as apical portion of arista-like stylus. Proboscis brown, with black hairs; palpus dark brown, with one short black apical bristle.

Thorax dark metallic green, with pale grey pollinosity. Hairs and bristles on thorax black; five strong dc, ten pairs of acr. Scutellum with two pairs sc, apical pair long strong, basal pair short and weak. Legs mainly dark yellow, but base of fore coxa, basal 7/8 of mid coxa and basal half of hind coxa brown, mid coxa with one black band at middle. Fore coxa anteriorly with three strong bristle at apical half, mid coxa with cluster of anterior hairs three marginal bristles anteriorly and one long strong outer bristle, hind coxa with one long strong outer bristle. Mid trochanter with two weak apical bristle dorsally. Fore femur without distinct bristle. Mid and hind femora each with one preapical pv. Fore tibia with one row of two ad, two pd, one pv and three ap. Mid tibia with three ad, four pd, two pv and four ap. Hind tibia with four ad and three ap. Mid tarsomere I with one short strong pd. Hind tarsomere I with one short strong ad at middle and one av at basal quarter. Relative lengths of tibia and five tarsomeres of legs LI: 6.6: 3.3: 1.6: 1.3: 1.0: 1.0; LII: 10.6: 5.0: 2.0:?:?:? (mid tarsus partly lost); LIII: 10.0: 4.0: 5.0: 3.3: 2.0: 1.3. Wing nearly hyaline, tinged brown; veins brown. R1 with one oval thickness at where connect to C. M with fading M2, M1 with one short subvein. Crossvein dm-cu somewhat arched. CuAx ratio 1.4. Lower calypter pale with black hairs. Haltere pale.

Abdomen metallic green, with pale pollinosity. Hairs and bristles on abdomen black.

Male genitalia: Epandrium 1.7 times longer than wide; epandrial lobe with three pale ap. Ventral surstylus and dorsal surstylus both with five short nail-like ap. Cercus nearly triangular, pale except the black ring, with weak digitations around outer margin, with long strong black marginal bristle on digitations, with one long strong bristles on digitation at apical half. Hypandrium simple. Phallus with two rows of black denticles on ventral half, anterior row only with four small sparse denticles, posterior row with eight dense denticles.

Female. Similar to male in size and morphology except for the male terminalia.

Etymology.

The name refers to the swelling of the costa.

Comments.

The new species should be compared to L. hirsutiseta since there is a node on the costa that is however distinctly before R1 joins the costa (Figure 22) in the latter.

Distribution.

Singapore.