Pachybrachis stygicus, , MCZ-FALL, 1915

Barney, Robert J., 2018, Definition and Revision of the Atomarius Species-Group of North American Pachybrachis Chevrolat (Coleoptera: Chrysomelidae: Cryptocephalinae), Including Descriptions of Nine New Species, The Coleopterists Bulletin 72 (1), pp. 9-74 : 59-65

Pachybrachis stygicus

Specimens Examined. Lectotype + 1217 specimens. USA: ALABAMA: Bibb Co., Bibb Co. Glades Preserve, 24.viii.2010, J. G. Hill [16, MEM]; Chambers Co. , Langsdale, H. W. Wenzel [16, OSUC]; same data, except H. H. Smith [2♀♀, USNM; 1♀, Fall 1915 Revis stygicus, MCZ-FALL ]; DeKalb Co. , DeSoto State Park , 18.v.1963, E. U. Balsbaugh, Jr. [1♀, AUEM]; Escambia Co. , 10 mi N Brewton, 13.vi.2003, J. Huether [1♀, USNM]; Delchamps, 12.vi [2♀♀, CIMNH]; Lee Co. , Auburn , 4.v.1908, W. E. Hinds [266 2♀♀, AUEM]; Auburn , east side, 15.v.1983, D. A. Rider [1♀, LSAM]; 3 mi N of Auburn, fish ponds, 24.iv.1963, E. U. Balsbaugh, Jr. [1♀, AUEM]; Goat Rock Dam, 1.vi.1964, H. G. Barwood [2♀♀, AUEM]; Marion Co. , Marion Co. Public Lake , 2.vi.1964, E. U. Balsbaugh, Jr. [1♀, AUEM]; Mobile Co. , Delchamps, 12.vi [2♀♀, CIMNH]; Shelby Co. , Oak Mountain State Park , 18.ix.1963, E. U. Balsbaugh, Jr. [16, AUEM]; Winston Co. , Bankhead National Forest , 2.5 mi N US 278, 3.vi.1964, E. U. Balsbaugh, Jr. [16, AUEM]. ARKANSAS: Hempstead Co. , Hope, 13.v.1925 [366 4♀♀, vestigialis, MCZ-FALL ]; same data, except 13-15.v.1923, L. Knobel [666 7♀♀, CEUM]; same data, except 13.v.1924 [766 5♀♀, CEUM]; same data, except 1-5.viii.1925 [866 4♀♀, CEUM]; Johnson Co. , 8 mi N Clarksville, 1.vii.1989, R. L. Brown [16 2♀♀, MEM]; Yell Co. , 11 mi NW Danville, 27.viii.1975, W. L. Johnson [1♀, MEM]. FLORIDA: Alachua Co. , 21.iv.1955, R. A. Morse [16, FSCA]; Archer , 1.v.1978, R. Turnbow [1♀, UGCA]; Gainesville, 14.vii, C. J. Drake [16, CIMNH]; same data, except 25.iv.1952, O. Peck [366 1♀, CNC]; same data, except 24.viii.1957, H. V. Weems [1♀, FSCA]; same data, except 27.v.1962, F. W. Mead [16, FSCA]; same data, except 28.xi.1963 [16, FSCA]; same data, except 22.iv.1968, H. V. Weems [16 1♀, FSCA]; same data, except 21.x.1979, S. Byby [1♀, FSCA]; same data, except vii.1990, W. P. Ponce [1♀, FSCA]; Gainesville (airport), 11.iv.2010, S. M. Clark [16 1♀, BYUC]; Payne’ s Prairie, 9.vii.1921, M. Leonard [2♀♀, USNM]; Payne’ s Prairie Preserve, 16.v.1982, R. J. Barney [1♀, RJBC]; Brevard Co. , Rockledge, 20.ii.1992, L. White [16, FSCA]; Broward Co. , Deerfield Beach Nature Preserve , 28.iv.1981, H. Katz [2♀♀, CMNH]; same data, except 18.xii.1986 [266, CMNH]; Ft . Lauderdale , 15.iv.1979, R. J. Barney [16 2♀♀, RJBC]; same data, except 6.iii.1979, P. W. Kovarik [1♀, EGRC]; Hollywood, 15.xii.1985, D. Furth [1♀, PMNH]; Pompano Beach, 8.i-21.ii.1971, E. J. Kiteley [1266 17♀♀, CNC]; Rockledge, 20.ii.1992, L. White [16, FSCA]; 2 mi N of I-75 on Rt. 27, 19.xii.2005, R. J. Barney & S. M. Clark [16, RJBC; 1♀, BYUC]; 2 mi N Andytown, 19. xii.2005, S. M. Clark [1♀, BYUC]; Calhoun Co. , 5 mi S Blountstown, Hwy 71, 30.iv.1985, O’ Brien & Marshall [4♀♀, FSCA]; Charlotte Co. , 13.iv.1986, N. M. Downie [266 2♀♀, FSCA]; Englewood Beach, 13.ix.1987, K. E. Jenkins [1♀, FSCA]; Charlotte Harbor Area , Little Gasparilla Is., 4. i.1978, E. G. Riley [1♀, EGRC]; Harbour Heights, 30.vii.1996, S. M. Clark [1♀, BYUC]; same data, except 31.vii.2000 [466 1♀, BYUC]; Citrus Co. , US 41 S Dunellon, 9.xi.2002, C. L. Staines [266 2♀♀, USNM]; Collier Co. , Naples, 12. v.1984, R. A. Belmont [16, FSCA]; same data, except 21.vi.1984, P. Hornby [16, FSCA]; 3 mi N of Jerome, Rt. 29, 20.xii.2005, R. J. Barney & S. M. Clark [1♀, RJBC]; Columbia Co. , Rt. 131 at Falling Creek , 19.x.1960, F. W. Mead [16, FSCA]; 4 mi SE Fort White, 17.iv.1999, S. M. Clark & W. Kovarik [3♀♀, FSCA; 1♀, EGRC; 2♀♀, BYUC]; Dade Co. , Coopertown, 28.iii.1974, B. F. & J. L. Carr [1♀, CNC]; Hialeah, 11.viii.1966, C. E. Stegmeier [16 2♀♀, FSCA]; same data, except 26.iv.1969 [2♀♀, FSCA]; Homestead, 4.iv.1952, J. R. Vockeroth [366 1♀, CNC]; same data, except 28.ii.1957, R. M. Baranowski [16, FSCA]; Miami, 2.vii.1956, D. R. Paulson [1♀, FSCA]; same data, except 17.vii.1965, C. E. Stegmeier [16, FSCA]; same data, except 15. vi.1969 [1♀, FSCA]; same data, except 15. vi.1972, W. H. Pierce [16, FSCA]; same data, except 7.vii.2008, O. Garcia [1♀, FSCA]; Opalocka, 22.vii.1962, F. W. Mead [16, FSCA]; DeSoto Co. , Brownsville, 1.vii.1963, D. G. Kissinger [1♀, FSCA]; Fort Ogden , 8.iv.1952, O. Peck [16 1♀, CNC]; Hwy . 671 at Peace, 15.iv.1990, R. Turnbow [2♀♀, RHTC; 1♀, FSCA]; Hwy . 72, 5 mi W Arcadia , 29.vi.1980, Wibmer [1♀, EGRC]; 3 mi E Arcadia, 13.viii.1986, Z. Smith [1♀, FSCA]; Duval Co. , Atlantic Beach [1♀, MCZ]; Jacksonville [266 1♀, Fall 1915 Revis stygicus, USNM ]; same data, except Mrs. A. T. Slosson [16, AMNH]; same data, except 23.iv [16 2♀♀, MCZ]; same data, except P. Laurent [2♀♀, USNM]; same data, except 1.v.1979, J. R. Watts [2♀♀, FSCA]; Flagler Co. , 7 mi SW Bunnell, 29.iii.1976, C. O’ Brien & G. Marshall [1♀, EGRC]; Washington- Oaks State Garden, 29.vi.1991, R. Turnbow [16 1♀, FSCA; 3♀♀, RHTC]; Gadsden Co. , Mount Pleasant , 1.v.1952, O. Peck [466 2♀♀, CNC]; Quincy, 1.v.1929, G. L. Garrison [1♀, USNM]; Gilchrist Co. , 3 mi W SR 47 on 340, 11.iv.2016, K. E. Schnepp [16, KESC]; Glades Co. , Venus, 20. vii.1989, J. Banta [16, FSCA]; Hamilton Co. , 3 mi W White Springs, 8.v.1974, R. Turnbow [1♀, UGCA]; Hendry Co. , Clewiston, 4.ii.1941, M. Wright [2♀♀, OSUC]; La Belle, 16.vii.1930, P. B. Lawson [2♀♀, SEMC]; 1 mi S Felda, 16.v.1977, R. Turnbow [16, UGCA; 16, EGRC]; Hernando Co. , Weeki Wachee Springs, 15.iii.1963, F. W. Mead [16, FSCA]; Withlacoochee State Forest , Apollo Beach, 31. iii.1990, R. Morris [16, UGCA]; same data, except Withlacoochee State Forest , Goat Road, 12.viii.1990 [266, UGCA]; same data, except 14. v.1994 [3♀♀, UGCA]; same data, except 20.iii.1994 [16, UGCA]; Highlands Co. , Archbold Biological Station , 26.ix.1976, L. L. Lampert [1♀, FSCA]; same data, except 18-23.ix.1978, N. M. Downie [1♀, ABS]; same data, except 24.ii.1979, Weems & Halkin [1♀, FSCA]; same data, except 1.v.1982, R. M. Brattain [1♀, BYUC]; same data, except 8. ix.1987, N. M. Downie [1♀, ABS]; same data, except 17-20.iii.2000, ex. Lepidium virginicum, M. Deyrup [2♀♀, ABS]; same data, except 12. xi.1997, V. Golia [1♀, USNM]; same data, except 24.vi.2001 [16 2♀♀, USNM]; same data, except 30. iii.2005, M. Deyrup [16, ABS]; same data, except 25.viii.2008, J. Huether [3♀♀, BYUC]; same data, except Lake Annie , 14.iv.2010, N 27.21092 W 81.34937, R. J. Barney & S. M. Clark [366 2♀♀, RJBC; 1♀, BYUC]; same data, except Lake Placid , 19.iv.1961, A. & H. Dietrich [16, CUIC]; same data, except 9-11.v.1961 [3♀♀, CUIC]; same data, except 27.vi.1986, J. E. Eger [1♀, EGRC]; same data, except 29.iii.1967, M. A. Deyrup [366 4♀♀, AMNH]; same data, except nr HQ RR, 15. iv.2010, N 27.18137 W 81.35206, R. J. Barney [1♀, RJBC]; Highlands Hammock State Park , 12. iv.1964, G. C. Eickwort [1♀, SEMC]; Lake Wales Ridge NWR, 25.viii.2008, R. Turnbow [1♀, RHTC]; Parker Island , 25.iii.1968, M. A. Deyrup [16, CUIC]; Hillsborough Co. , Lutz, 9.viii.2003, L. Stange [1♀, FSCA]; Tampa, 21.iv, Hubbard & Schwarz [266, Fall 1915 Revis carbonarius, USNM]; Tampa, 1927, C. D. Bare [3♀♀, SEMC]; same data, except 20.iii.1982, J. E. Eger [1♀, EGRC]; 1 mi S of Sun City, 19.v.1982, R. J. Barney [1♀, RJBC]; Holmes Co. , I-10, 5.5 mi E Hwy. 279, 20.v.1985, E. Riley & D. Rider [1♀, EGRC]; Indian River Co. , 2.5 mi SE Wabasso, 16.v.1986, R. Turnbow [1♀, RHTC]; Jackson Co. , 10.iv.1985, N. M. Downie [16, FSCA]; Spring Lake , 22.v.1979, W. H. Cross [1♀, MEM]; Jefferson Co. , Monticello, 19.iv.1928, P. W. Fattig [1♀, UGCA]; Lafayette Co. , Hwy 27 near Suwannee River , 17.iv.1999, S. M. Clark [2♀♀, BYUC]; Lake Co. , Eustis, 10.iv.1955, W. F. Chamberlin [1♀, EGRC]; Hwy . 27, Vic. Clermont, 24.viii.2008, J. Huether [1♀, BYUC]; Umatilla, 26.v.1989, B. Daigneau [16, FSCA]; 5 mi E Cassia Station, 26.iii.1976, O’ Brien & Marshall [1♀, BYUC]; Lee Co. , Lehigh, 7-10.iv.1977, N. M. Downie [2♀♀, PERC]; same data, except 12.iv.1978 [3♀♀, PERC]; same data, except 9.vi.1985 [3♀♀, UNH]; 1 mi N of San Carlos Park, 28.vi.1987, R. J. Barney [16 2♀♀, RJBC]; 3 mi S of Estero [16 1♀, RJBC]; Sanibel Island , 13.iii.1986, J. E. Eger [2♀♀, EGRC]; Leon Co. , Apalachicola Mat. For., 0.5 mi E Clear Lake on Rt. 305, 16.iv.1999, S. M. Clark [16, BYUC]; Tallahassee, 14.vii.1934, P. McKintrey [16, SEMC]; Tall Timbers Ecological Research Station , 27.iv.1977, M. A. Altieri [1♀, EGRC]; same data, except 13.iv.1999, S. M. Clark [2♀♀, BYUC]; Levy Co. , 7.vii.1958, H. V. Weems [266 3♀♀, FSCA]; Cedar Keys, 12. vii.1929, P. B. Lawson [1♀, SEMC]; Gulf Hammock, 23.iv.1952, O. Peck [16 1♀, CNC]; Hwy 24 at Alachua Co. line, 9.iv.1980, R. Turnbow [1♀, UGCA]; SR24 at Alachua Co. line, 3.v.1989, M. C. Thomas [1♀, FSCA]; Yankeetown, 17.vii.1934, P. McKintrey [16 1♀, SEMC]; 2 mi W Archer, 22.iii.1953, B. K. Dozier [266 3♀♀, FSCA]; 3 mi SW Archer, 28.iv.1971, R. Turnbow [1♀, FSCA; 16, RHTC]; same data, except 6.v.2016, K. E. Schnepp [1♀, KESC]; 3 mi W Archer, 30.vii.1994, Morris & Brattian [1♀, UGCA]; 3.8 mi SW Archer, 4.v.1988, P. Skelley [16 2♀♀, FSCA]; same data, except 29.v.1990, R. S. Anderson [16, EGRC]; Manatee Springs State Park , 28.iv.1988, E. Riley & F. Whitford [266 1♀, EGRC]; 5.5 mi N Bronson, 12.iv.2010, S. M. Clark [1♀, BYUC]; Williston Highlands, 20. iv.2016, K. E. Schnepp [1♀, KESC]; Liberty Co. , Torreya State Park , 19.v.1985, E. Riley & D. Rider [1♀, EGRC]; Manatee Co. , Oneco, 29.iii.1955, D. M. Anderson [266 5♀♀, CUIC]; same data, except J. C. Martin [566, CNC]; 1 mi NE Parrish, 29. v.1976, C. O’ Brien & G. Marshall [1♀, EGRC]; Parrish area , 24.iv.1996, R. Morris [1♀, UGCA]; Marion Co. , I-75 exit near Ocala, 10.xii.1997, M. J. Rothschild [16, USNM]; SR 88 Ocala State Forest , 27.viii.2008, J. Huether [2♀♀, BYUC]; 1 mi SW Orange Lake, 17.iv.1993, S. M. & B. S. Clark [1♀, BYUC]; Monroe Co. , Key Largo , 27. iii.1956, H. V. Weems [466 6♀♀, FSCA]; same data, except 9.iv.1959 [1♀, FSCA]; same data, except 17.vii.1963, H. A. Denmark [16, FSCA]; 15 mi NE Key Largo City, 16.vi.1974, J. B. Hepner [16, FSCA]; Key West, H. W. Wenzel [1♀, OSUC]; Marathon Key , 1.v.2000, R. Turnbow [1♀, RHTC]; Plantation Key , 27.xi.1955, H. V. Weems [1♀, FSCA]; Upper Key Largo , 23-24.vii.1984, E. G. & T. J. Riley [16, EGRC]; same data, except 30.iv.1988, E. Riley & F. Whitford [1♀, EGRC]; Nassau Co. , Fernandina, 22.iv, P. Laurent [16, ANSP]; Okaloosa Co. , 30.vii.1955, F. W. Mead [16 2♀♀, FSCA]; Okeechobee Co. , near Okeechobee, 5.iv.1994, S. M. Clark [366 6♀♀, BYUC]; Orange Co. , Geneva, 25.ix.1993, C. Scherer [1♀, UCFC]; nr. Orlando, 12.i.1938, H. H. Hungerford [1♀, SEMC]; Orlando, 29.iii.1953, W. R. M. Mason [266 1♀, CNC]; UCF, Orlando, 2. xi.1990, S. M. Fullerton [1♀, UCFC]; same data, except 28.iv.1991 [16 1♀, UCFC]; same data, except 4.x.1991 [1♀, UCFC]; same data, except 14.ix.1991 [266 1♀, UCFC]; Hwy . 532, 1.5 mi S jct. Hwy . 520, 8.iv.1994, R. Turnbow [1♀, RHTC]; Palm Beach Co. , Jupiter Island , 20.vi.1965, F. W. Mead [1♀, FSCA]; Riviera Beach, 3.ii.1988, K. Nicholson [16, FSCA]; Royal Palm Beach, N SR98 & 441, 8.v.1973, W. H. Pierce [2♀♀, FSCA]; Singer Island , 2.v.1946 [1♀, USNM]; 13 mi NW of PB Co. line, 19.xii.2005, R. J. Barney & S. M. Clark [16 5♀♀, RJBC; 16 1♀, BYUC]; Pasco Co. , 6 mi N Land O’ Lakes, 23.vii.1973, O’ Brien & Marshall [1♀, EGRC]; Pinellas Co. , Clearwater, 17. vii.1990, M. A. Goodrich [2♀♀, EIUC]; Dunedin, 28.i.1917 [1♀, AMNH]; same data, except 6.iv.1922 [1♀, MCZ-FALL]; same data, except 15.iii.1913, W. S. Blatchley [1♀, MCZ-FALL]; same data, except 23.i.1917 [1♀, PERC]; Honeymoon Island State Park , 4.v.1988, E. G. Riley [1♀, EGRC]; St. Petersburg , 29.iii.1925 [2♀♀, MCZ-FALL]; Tarpon Springs, 9.viii.1970, F. W. Mead [1♀, FSCA]; Polk Co. , Haines City, 14.iv.1971, C. Brivio [16, PMNH]; Lake Wales , 17.xi.1987, P. Lindsay [1♀, FSCA]; Putnam Co. , Welaka, 10-17. v.1961, A. & H. Dietrich [266 5♀♀, CUIC; 266, RJBC]; Sarasota Co. , Englewood, 4.iv.1983, N. M. Downie [16 2♀♀, UNH]; same data, except 15. ix.1988, K. E. Jenkins [266, FSCA]; Fruitville, 11.viii.1930, R. H. Beamer [1♀, SEMC]; Laurel, 27.xi.1991, S. M. Clark [466 2♀♀, BYUC]; Oscar Scherer State Park , 13.iv.1966, H. V. Weems [1♀, FSCA]; Siesta Key , 17.iii.1963, F. W. Mead [1♀, FSCA]; Venice , 3.x.1988, K. E. Jenkins [2♀♀, FSCA]; 2 mi E Venice, 8.iv.1994, S. M. Clark [1♀, BYUC]; Seminole Co. , Geneva, 5.vi.1963, G. W. Desin [16, FSCA]; Sanford, iv.1908, Van Duzee [2♀♀, Fall 1915 Revis stygicus, USNM ]; same data, except 4.iv.1913, W. S. Blatchley [1♀, PERC]; same data, except 2.iv.1928, F. M. Uhler [16 1♀, USNM]; same data, except 11.vii.1951, Beamer, Price & Weed [366, SEMC]; St. John’ s Co., Anastasia State Park , 5.vii.1990, S. M. Fullerton [1♀, UCFC]; Ft . Matanzas National Monument, 16.iii.1983, M. C. Thomas [16 1♀, FSCA]; St. Augustine , 12.iv.1919 [1♀, MCZ- FALL; 1♀, MCZ]; St. Lucie Co. , Fort Capron , 1. iv, O. Lugger Collection [1♀, UMSP]; 22 mi W Ft. Pierce, 16.vii.1973, O’ Brien & Marshall [1♀, FSCA]; Suwannee Co. , Suwannee Spring, 24.viii.1939, A. T. Hardy [16, SEMC]; Taylor Co. , Blue Spring Lake , 7.vi.1974, C. L. Smith [1♀, UGCA]; Rt. 98 & Ecolina River , 1.iv.1993 S. M. Fullerton [1♀, UCFC]; 7 mi S Perry, 8. ix.1978, E. R. Hobeke [1♀, CUIC]; 28 mi SW Perry, 5.ix.1970, W. T. Atyeo [1♀, UGCA]; S Perry , Hwy . 381, 29.vi.1973, G. B. Marshall [16, BYUC]; Union Co. , Hwy 241, 1 km N Santa Fe River , 28.viii.1986, C. W. Mills [16, FSCA]; Hwy 239A near Hwy 241, 5.iv.1989, C. W. Mills [366 9♀♀, FSCA]; Volusia Co. , Daytona, 7.iv.1919 [16, MCZ]; same data, except 29.x.1971, T. L. Kipp [16, FSCA]; DeLand, 27.iii.1967, M. A. Deyrup [1♀, AMNH]; Edgewater, 7.iii.1939, C. A. Frost [2♀♀, MCZ]; Enterprise [266 2♀♀, LECTOTYPE, MCZ- FALL]; same data, except 14.iv [266 2♀♀, MCZ]; same data, except 15.iv, G. M. Greene [16 1♀, USNM]; New Smyrna, vi, Hubbard & Schwarz [1♀, Fall 1915 Revis carbonarius, USNM]; Ormond [16, MCZ]; same data, except Mrs. A. T. Slosson [16 1♀, AMNH]; Ormond Beach , 10. iv.1983, E. G. Riley [16 3♀♀, EGRC]; 1 mi N Ormond Beach, 31.iii.1979, E. G. Riley & D. LeDoux [16 2♀♀, EGRC]; Tomoka State Park , 1. iv.1974, G. C. Eickwort [16, CUIC]; Walton Co. , 2. vii.1938, G. R. Swank [1♀, INHS]; Wakulla Co. , Wakulla, 11.vii.1939, P. B. Lawson [16, SEMC]; Jay Beach, 5.v.1944, Frison & Ross [16, INHS]; Unknown county , Fla. [1♀, CIMNH; 3♀♀, MCZ- HORN; 1♀, MCZ-LEC]; Florida, [16, CMNH]; same data, except W. H. Ashmead [16 1♀, INHS]. GEORGIA: Bibb Co. , Lizella, 18. v.1939, P. W. Fattig [1♀, UGCA]; Chatham Co. , Savannah, 23.v.1926 [16, MCZ-FALL]; same data, except 23.v.1979, R. S. Peigler [1♀, UGCA]; Tybee Island , 24.vi, H. A. Wenzel [16 2♀♀, PARATYPE, USNM]; same data, except 12.iv, P. Laurent [1♀, USNM]; Cook Co. , Adel, 11.viii.1939, J. D. Beamer [1♀, SEMC]; Crawford Co. , Roberta, 12. v.1939, P. W. Fattig [16 2♀♀, UGCA]; same data, except 2.viii.1939 [1♀, UGCA]; Decatur Co. , Experimental Forest , Hwy . 97, near Faceville, 1. vi.2000, C. W. O’ Brien [366, BYUC]; Dougherty Co. , Albany, 26.iv.1938, P. W. Fattig [16, UGCA]; same data, except 8.vi.1939 [1♀, NDSRIC]; Elbert Co. , 1.4 mi SW Bowman, 1.viii.2004, R. Turnbow [266, RHTC]; Emanuel Co. , George L. Smith State Park , 10.vi.2013, S. M. Clark & E. G. Riley [16 1♀, BYUC]; 1.3 mi E Twin City, roadside, 10.vi.2013, R. J. Barney [1♀, RJBC]; Glynn Co. , St. Simmons Island , 13.vii.1931, C. A. Frost [266 2♀♀, MCZ]; same data, except 1-6.viii.1988, A. J. Gilbert [2066 5♀♀, AJGC]; Habersham Co. , Yonah Mt. , 20. vi.1937, P. W. Fattig [1♀, UGCA]; Harris Co. , Hamilton, 16.v.1936, P. W. Fattig [1♀, UGCA]; Hart Co. , 0.5 mi S Lake Hartwell, 11.vi.2000, W. B. Sikora [266, UGCA]; Houston Co. , Perry , 13. v.1939, P. W. Fattig [1♀, NDSRIC]; Jefferson Co. , Louisville, 8.v.1937, P. W. Fattig [1♀, UGCA]; Johnson Co. , 1 mi E Kite, 18.viii.1976, on Quercus laevis, R. Turnbow [16, AMNH]; same data, except 1.ix.1976 [16 1♀, UGCA]; Macon Co. , Macon, 13.v.1939, P. W. Fattig [1♀, NDSRIC]; McIntosh Co. , Sapelo Island , 28.iv.1962, H. O. Lund [266 1♀, UGCA]; Wolf Island , 7.v.1955, R. T. Franklin [16, UGCA]; Meriwether Co. , Greenville, 26.v.2004, J. H. Chandler [2♀♀, UGCA]; same data, except 27.vi.2004, on foliage of Eupatorium capillifolium [16, UGCA]; Rabun Co. , Pine Mountain 1400’, 25.v.1957, W. J. Brown [2♀♀, CNC]; Schley Co. , Ellaville, 28.iv.1938, P. W. Fattig [266 2♀♀, UGCA]; Spalding Co. , Experiment, 17.ix.1937, T. L. Bissell [1♀, UGCA]; Griffin, 12.viii.1939, A. T. Hardy [1♀, SEMC]; Tattnall Co. , 3 mi E 147 along Ohoopee Road, 8-10.v.1998, R. Morris [16 1♀, UGCA]; Thomas Co. , Thomasville, 16.v.1937, P. W. Fattig [266 1♀, UGCA]; Tift Co. , Tifton, 1.viii.1948, P. McKintrey [16, SEMC]; same data, except 2. viii.1936, P. A. Glick [1♀, USNM]; Upson Co. , Mt. Pleasant rd. , S of Yatesville, 16.v.1992, R. Morris [266 4♀♀, UGCA]; Ware Co. , Okefenokee Swamp, 27.vii.1939, R. H. Beamer [2♀♀, SEMC]; Washington Co. , Sandersville, 8. v.1937, P. W. Fattig [1♀, UGCA]; Worth Co. , Hwy . 300, 2.7 mi N Dougherty Co. line, 14. v.1988, R. Turnbow [16, RHTC]; Unknown county , Geo., Morrison [16 1♀, stygicus HCF, SEMC ]. KANSAS: Barber Co. , Hardtner, 28. viii.2002, B. & J. Smith [16 1♀, EGRC]; same data, except 24.ix.2003 [1♀, EGRC]; same data, except 9.viii.2004 [266 1♀, EGRC]; 16 mi W Medicine Lodge, Hwy. 60, 5.vi.2007, R. J. Barney [16, RJBC]; Comanche Co. , 5 mi E Coldwater, Hwy. 160, 6.vi.2007, S. M. Clark [1♀, BYUC]; Douglas Co. , Lawrence, 24.viii.1948, R. H. Beamer [16, SEMC]; Edwards Co. , 6.5 mi S Kinsley, 5.vi.2007, S. M. Clark & R. J. Barney [16 1♀, BYUC; 16 1♀, RJBC]; Hamilton Co. , 3350 ft., F. H. Snow [16, vestigialis HCF, SEMC ]; Harvey Co. , 3 mi N Burton, 4. vi.2007, S. M. Clark & R. J. Barney [266 2♀♀, BYUC; 766 2♀♀, RJBC]; Montgomery Co. , Caney , vi.1931, Beck [16, BYUC]; Elk City , 24.v.1940, M. Sanderson [266 3♀♀, SEMC]; Pratt Co. , Pilot Knob , 4.vi.2007, R. J. Barney [1♀, RJBC]; Reno Co. , 4.ix.1923, W. E. Hoffmann [16 3♀♀, SEMC]; Medora , 25.vi.1940, D. R. Lindsay [266 2♀♀, SEMC]; Rice Co. , Sterling , 25.vi.1940, D. R. Lindsay [16, SEMC]; 10.5 mi NW Medora, 4.vi.2007, R. J. Barney [1♀, RJBC]; Wabaunsee Co. , 17.vi.1963, J. R. McCoy [1♀, MEM]. LOUISIANA: Bossier Pr., Barksdale AFB, 19. v.1996, T. L. Schiefer [16, MEM]; same data, except 17.v.1996, E. G. Riley [16 1♀, EGRC]; Calcasieu Pr., Sicily Island Hills W.M.A., 29. iv.1988, C. B. Barr [16, LSAM]; Cameron Pr., Cameron , 14-28.viii.1903 [16 2♀♀, OSUC]; same data, except 20-30.vi.1905, J. S. Hine [16 2♀♀, OSUC]; Catahoula Pr., Sicily Island Hills W.M. A., 8.v.1991, C. B. Barr [1♀, LSAM]; East Baton Rouge Pr. , Baton Rouge , Siegan Lane at I-10, 10.x.1987, D. A. Rider [16, LSAM]; East Feliciana Pr. , Boy Scout Camp Avondale ; LA Hwy 10 E of Clinton, 9.v.1986, C. B. Barr [3♀♀, LSAM]; Evangeline Pr., 3 mi. E of Ville Platte, 26. viii.1992, D. A. Rider [1♀, LSAM]; Livingston Pr., Corbin , 18.vii.1971, L. Klenk [2♀♀, LSAM]; Denham , 20.vi.1948, H. W. Crowder [16 1♀, SEMC]; Natchitoches Pr., 16.viii.1928, R. H. Beamer [1♀, SEMC]; Kisatchie National Forest Red Dirt W.M.A., 30.iv.1989, D. A. Rider [266 2♀♀, LSAM]; same data, except 17.v.1984, E. G. Riley [266 1♀, EGRC]; Winn Pr., ca. 2.5 mi. SW Calvin along Parish Rd. 9, 9.v.1991, C. B. Barr [1♀, LSAM]; St. Landry Pr. , Opelousas , vi, Pilate [366 1♀, CMNH]. MISSISSIPPI: Chickasaw Co. , Tombigbee National Forest , 17.vi.2001, J. G. Hill [1♀, MEM]; same data, except 16.viii.2001 [16, MEM]; Jasper Co. , 6 mi N Montrose, 31.vi.2003, J. Huether [1♀, USNM]; Kemper Co. , 5 mi N of Scooba, 2.ix.2010, J. G. Hill [16 1♀, MEM]; Lauderdale Co. , ~ 7 mi E Lauderdale, 12.ix.2009, J. G. Hill [266 2♀♀, MEM]; Lincoln Co. , Brookhaven , 3.v.1927, ex. pecan, F. S. Fortenberry [11♀♀, MEM]; Lowndes Co. , Artesia , 1.v.1921, A. McIntosh [16, MEM]; Newton Co. , vic. Hickory , 1.vi.1987, R. Morris [1♀, UGCA]; I-20 & Hwy . 80, 29.iv.1987, R. Morris [1♀, UGCA]; Noxubee Co. , Noxubee National Wildlife Refuge , 4.vi.1997, T. L. Schiefer [1♀, MEM]; same data, except 11.v.2011 [266 2♀♀, MEM]; Oktibbeha Co. , Osborn , 28.viii.2009, J. G. Hill [266, MEM]; Scott Co. , 1-7 mi E Morton on Hwy. 80, 2.vi.1987, R. Morris [16 1♀, FSCA; 1♀, UGCA]; 2.4-5.5 mi W Forest, 11.vi.1989, R. Turnbow [2♀♀, RHTC]; same data, except 12.vi.1992 [1♀, RHTC]. NEBRASKA: Buffalo Co. , Kearney , 10.vii.2000, S. M. Clark [266, BYUC]; Dawson Co. , Willow Island Wildlife Area , 6.vii.1997, S. M. Clark [266 3♀♀, BYUC]; Hall Co. , Mormon Island Refuge , 5. vii.1980, B. C. Ratcliffe [1♀, UNSM]; Holt Co. , Spencer Dam , 15-24.vi.1974 [1♀, UNSM]; Merrick Co. , 2 mi W Chapman, 11.viii.1958, Aty & Andresen [16, UNSM]; Thomas Co. , Halsey , 4. viii.1958, R. Henzlik [16, UNSM]; same data, except vii.1969 [16, UNSM]; same data, except Forest Reserve , 4.vii.1908, R. H. Wolcott [266 3♀♀, UNSM]; Unknown county , Sand Hills , vii [266, UNSM]. NORTH CAROLINA: Avery Co. , Pineola , 11.vii.1942, D. L. Wray [4♀♀, NCSU]; Bertie Co. , Lewiston , 28.ix.1963, J. F. Cornell [16 1♀, NCSU]; Bladen Co. , Bladen Lakes State Forest , Jct. SR 1509 & 1510, 25.viii.1992, R. L. Blinn [16 4♀♀, NCSU]; White Lake , 14.v.1952, W. J. Gehweller [16 2♀♀, NCSU]; Brunswick Co. , Southport , 23.vii.1981, J. M. Stucky [16, NCSU]; Belville , John Long Park , 7.ix.1999, S. M. Clark [1♀, BYUC]; Buncombe Co. , Swannanoa , vi.1919, F. Sherman [16 1♀, NCSU]; Carteret Co. , Atlantic Beach , 3-4.ix.1986, G. F. & J. F. Hevel [16 3♀♀, USNM]; Duplin Co. , 25.vi.1964, H. D. Blocker [266 1♀, NCSU]; same data, except 20.vi.1963 [1♀, NCSU]; Moore Co. , Aberdeen , 25.v.1922, R. W. Lieby [1♀, NCSU]; Southern Pines , 20. v.1919, A. H. Manee [1♀, NCSU]; same data, except 13.v.1920 [16 1♀, NCSU]; New Hanover Co. , Carolina Beach , 4.vii.1927 [1♀, NCSU]; same data, except 15.vii.1951, R. H. Beamer [1♀, SEMC]; Fort Fisher State Park , 14.vi.2013, S. M. Clark [4♀♀, BYUC]; Onslow Co. , 4.viii.1959, W. D. Duckworth [16, NCSU]; same data, except 25. vi.1964, H. D. Blocker [2♀♀, NCSU]; Ashe Island , 19.viii.1975, on Eupatorium capillifolium, J. C. Dukes [16, NCSU]; Onslow Beach , 13.ix.1972, G. E. Bohart [566 3♀♀, BYUC]; Pender Co. , Willard , 11.v.1951, T. B. Mitchell [1♀, NCSU]; Scotland Co. , Lauringburg , 17.viii.1959, W. D. Duckworth [16, NCSU]; Wake Co., 17.v.1962, F. L. Roberts [16, NCSU]; Wilkes Co. , 16.vii.1964, J. F. Cornell [1♀, NCSU]. OKLAHOMA: Alfalfa Co. , Great Salt Plains Lake , Sandpiper , 36.8011 o N, 98.2516 o W, 5.vi.2007, R. J. Barney & S. M. Clark [366 2♀♀, RJBC; 266 1♀, BYUC]; 5.8 mi E Ingersoll, 36.7973 o N, 98.2862 o W, 5.vi.2007, R. J. Barney & S. M. Clark [266 3♀♀, RJBC; 566 1♀, BYUC]; Carter Co. , Ardmore vicinity, 3.vi.1961 [16, SEMC]; Ardmore , 8.vi.1939, Kaiser-Nailon [16, OSEC]; Choctaw Co. , 7.5 mi S Antlers, picnic area, 34.1214 o N, 95.5785 o W, 31. v.2007, R. J. Barney & S. M. Clark [366 2♀♀, RJBC; 2♀♀, BYUC]; Comanche Co. , 0.75 mi SE Medicine Park, 34.7234 o N, 98.4885 o W, 6.vi.2007, R. J. Barney & S. M. Clark [266 1♀, RJBC; 366 1♀, BYUC]; Ellis Co. , 6 mi SE Arnett, 1.viii-14. x.1967, D. C. Arnold [366, OSEC]; Harper Co. , Doby Springs Park , 16.v.2006, M. Gimmel [1♀, LSAM]; Haskell Co. , Whitefield , Canadian River , 5.vi.1983, K. Stephan [16 2♀♀, EGRC]; Jefferson Co. , Waurika , 4.vii.1937, Standish-Kaiser [16 1♀, SDSU]; Latimer Co. , Red Oak , 26.ix.1976, K. Stephan [1♀, EGRC]; 5 mi W Red Oak, viii.1980 [1♀, EGRC]; same data, except v-viii.1981 [4♀♀, EGRC]; same data, except v.1982 [2♀♀, EGRC]; same data, except vii-ix.1984 [6♀♀, EGRC]; same data, except ix.1986 [2♀♀, EGRC]; Le Flore Co. , Muse , 25.vi.1937, Standish-Kaiser [266 1♀, SDSU]; Page , 23.vi.1937, Standish-Kaiser [16 1♀, SDSU]; Mayes Co. , 2 mi N Chouteau, Hwy. 69, 36.215 o N, 95.341 o W, 1.vi.2007, S. M. Clark [1♀, BYUC]; Okmulgee Co. , 2 mi N Grayson, Hwy. 52, 35.5384 o N, 95.8605 o W, 1.vi.2007, R. J. Barney & S. M. Clark [16, RJBC; 1♀, BYUC]; Henryetta , 19.vii.1937, Standish-Kaiser [366 2♀♀, SDSU]; Ottawa Co. , Cardin , 36.975 o N, 94.844 o W, 1.vi.2007, S. M. Clark [1♀, BYUC]; Payne Co. , Stillwater , 10. vi.1971, V. M. Kirk [1♀, NDSRIC]; same data, except 15.vi.1917, O. Wade [1♀, UNSM]; Crosstimbers Exp. Range, 23.vi.1994, Bidwell-Engle [16, OSEC]; same data, except 15.vi.1995 [266, OSEC]; Pontotoc Co. , Pontotoc Ridge Preserve , National Conservation Property , 10.v.2014, M. Quinn [16 2♀♀, EGRC]; Woods Co. , 14 mi N, 7-8 mi W Alva, 30.vii-16. viii.2002, B & J Smith [3♀♀, EGRC]; same data, except 14.v.2004 [16, EGRC]; same data, except 28.ix.2003 [1♀, EGRC]; same data, except 12.ix.2004 [1♀, EGRC]; Unknown county , Wichita National Forest , 1928, A. C. Weese [266, SDSU; 1♀, NDSRIC]; same data, except 27.vi.1936, D. R. Lindsay [266 3♀♀, EGRC]. SOUTH CAROLINA: Aiken Co. , Aiken , 4.v.1939, J. G. Watts [16 1♀, CUAC]; same data, except 22.vi.1957, W. R. M. Mason [16 2♀♀, CNC]; SRP S-Area Bay , 15.vi.1979, A. Anderson [16, UGCA]; Barnwell Co. , Blackville , 6.viii.1937, J. G. Watts [266 2♀♀, CUAC]; same data, except 11.viii.1960, V. M.Kirk [1♀, NDSRIC]; Beaufort Co. , Beaufort , 15. viii.1935, J. G. Watts [266 2♀♀, CUAC]; same data, except 14.viii.1939, O. L. Cartwright [266 1♀, CUAC]; Bluffton , 19.v.1932, J. G. Watts [266, CUAC]; Charleston Co. , Charleston , 9.vii.1935, J. G. Watts [266 3♀♀, CUAC]; James Island , 21.vii.1950, W. F. Chamberlin [1♀, TAMU]; Mt. Pleasant , 13.viii.1935, O. L. Cartwright [2♀♀, CUAC]; Colleton Co. , Edisto Island , 14.viii.1935, J. G. Watts [266, CUAC]; Dorchester Co. , Givens Ferry , 3.vi.1950, W. F. Chamberlin [16, TAMU]; Summerville , 23.vii.1953, V. M. Kirk [1♀, NDSRIC]; Fairfield Co. , 2 mi W Winnsboro, 12.vi.2013, S. M. Clark [1♀, BYUC]; Florence Co. , Florence , 23.v.1935, J. G. Watts [966 9♀♀, CUAC]; Georgetown Co. , Georgetown , 13.viii.1936, O. L. Cartwright [16, CUAC]; Hampton Co. , Hampton , 15.viii.1935, O. L. Cartwright [266 4♀♀, CUAC]; Pickens Co. , Clemson , 8.v.1955, D. Dunavan [16, CUAC]; Richland Co. , Columbia , 18.ix.1925, O. L. Cartwright [266, CUAC]; Sumter Co. , Manchester State Forest , 5.5 mi N Rimini, 34 o 44’67”N, 80 o 30’80”W, 29.v.2006, E. G. Riley [366 6♀♀, EGRC]. SOUTH DAKOTA: Bon Homme Co. , Springfield , 13.vii.1966, V. M. Kirk [16 2♀♀, NDSIRC]; Custer Co. , Custer , 17.vii.1932, F. R. Bingham [16, SDSU]; same data, except 15.vii.1933 [566 3♀♀, SDSU]. TENNESSEE: Hardeman Co. , Hwy . 57 at Hatchie River , 4. vi.2007, R. Turnbow [16, RHTC]. TEXAS: Anderson Co. , Elkhart , 30.ix.1961, H. R. Burke [3♀♀, TAMU]; Salmon , 18.viii-20.ix.1974, Malaise trap, H. R. Burke [466, TAMU]; same data, except 17. v-9.vi.1975 Malaise trap, H. R. Burke [2♀♀, TAMU]; Gus Engeling WMA, 4.v.2002, S. M. Clark [16 2♀♀, BYUC]; Aransas Co. , Rockport , 7.viii.1928, J. G. Shaw [1166 7♀♀, TAMU]; Bastrop Co. , Bastrop State Park , 10.v.1989, A. J. Gilbert [16 2♀♀, AJGC]; Bexar Co. , Macdona , 29.vii, H. A. Wenzel [266 1♀, vestigialis, OSUC ]; Brazos Co. , 9.iv.1933, H. G. Johnston [16 2♀♀, TAMU]; same data, except iv.2006, E. G. Riley [266 1♀, BYUC]; College Station , 5-19.ix.1931, H. J. Reinhart [1166 13♀♀, TAMU]; same data, except 3.v.1931 [266, TAMU]; same data, except 21.iv-21.v.1932 [266 3♀♀, TAMU]; same data, except 15.v.1977, ex. Rudbeckia, S. J. Merritt [16, TAMU]; Riley Estate , 30 o 35’18”N 96 o 15’12”W, v.2006, E. G. Riley [866 8♀♀, TAMU]; Hwy . 30 7.6 mi E jct. Hwy . 6, 7.v.1981, ex. Coreopsis, S. J. Merritt [266 3♀♀, TAMU]; TAMU west campus, 31.v.1981, ex. Rudbeckia, S. J. Merritt [266 1♀, TAMU]; Peach Creek Road , near Lick Creek , 27.v.2007, S. M. Clark [16 2♀♀, BYUC]; Brooks Co. , 11 mi N Encino, 24.ix.1996, S. M. Clark [16 3♀♀, BYUC]; 9-10 mi S Falfurrias, 8-9.v.1989, E. G. Riley [266 9♀♀, TAMU]; Cameron Co. , Boca Chica Beach , 14.x.1988, E. G. Riley [466 8♀♀, TAMU]; same data, except 19.x.1991 [466 9♀♀, TAMU]; same data, except 4.xi.2013, 25.99709 o N, 97.15198 o W, E. G. Riley [16 2♀♀, TAMU]; same data, except 12.x.1993, S. M. Clark [16 2♀♀, BYUC]; same data, except dune area, 8.iii.2008, D. Heffren [266, TAMU]; salt flat & dune, 25.9970 o N 97.1515 o W, 4.xi.2013, R. J. Barney [466 3♀♀, RJBC]; Erath Co. , 3 mi W Bluffdale, 22-29.vii.1976, R. L. Sams [16, TAMU]; Fayette Co. , Flatonia , 30.vii, H. A. Wenzel [4♀♀, OSUC]; Freestone Co. , Old Spring Seat Church , nr. Donie , 6. v.1995, E. G. Riley [466 4♀♀, TAMU]; Galveston Co. , 3.5 mi SW Jamaica, 29.iii.1997, E. G. Riley [5♀♀, TAMU]; Goliad Co. , 4 mi SE Goliad, 2. v.1989, A. J. Gilbert [266 1♀, AJGC]; 11.2 mi SE Goliad, 3.v.1989, E. G. Riley [266 3♀♀, TAMU]; Harris Co. , Seabrook , 6-9.viii, H. A. Wenzel [366 6♀♀, vestigialis PARATYPE, OSUC]; Hopkins Co. , Minter Springs , 20.v.1989, R. Anderson [16 2♀♀, EGRC]; Kenedy Co. , 6 mi S Sarita, 14.x.1993, S. M. Clark [16 4♀♀, BYUC]; 31.8 mi S Sarita, 19.x.1990, T. Carlow [16 3♀♀, TAMU]; 4 mi N Norris, 14.x.1993, S. M. Clark [16 1♀, BYUC]; Kleberg Co. , 3 mi S Kingsville, Hwy. 77, 16.iv.1986, D. E. & S. J. Hanselman [566 3♀♀, BYUC]; 4 mi E Riviera, 7.v.1962, B. McDaniel [16 1♀, TAMU]; Kingsville , 29.iii.1962, ex. Coreopsis cordeminefolia [sic], B. McDaniels [16, TAMU; 166 1♀, SDSU]; Lamar Co. , Camp Maxey , Training Area 3, 24.v.2003, E. G. Riley [16 2♀♀, TAMU]; Lipscomb Co. , Jct. Hwy 23 on Hwy . 83, 18.v.2002, E. G. Riley [1♀, TAMU]; Live Oak Co. , 10 mi S George West, 22.iv.1978, D. W. Plitt [366, TAMU]; Montgomery Co. , Magnolia , 30.iv.1953, ex. Verbena, L. D. Beamer [1♀, TAMU]; Nueces Co. , Newport Pass , 3 mi N Galleon Bay on Rt. 361, 12.iv.1995, L. Torres-Miller [266 2♀♀, BYUC]; Patricio Co. , Lake Corpus Christi State Park , 4.v.1987, E. G. Riley [1♀, LSAM]; Randall Co. , Palo Duro Canyon State Park , Hackberry Campground , 15.v.2002, 34 o 58’03”N, 101 o 40’20”W, E. G. Riley [566 1♀, TAMU]; Robertson Co. , 7 mi W jct. OSR & FM 1940, 28. v.1978, ex. Rudbeckia, S. J. Merritt [2♀♀, TAMU]; 5.1 mi W jct. OSR & FM 1940, 28.v.1978, ex. Englemannia, S. J. Merritt [1♀, TAMU]; 12.1 mi E jct. OSR & FM 1940, 4.vi.1977, ex. Rudbeckia, S. J. Merritt [16, TAMU]; Sabine Co. , Toledo Bend Reservoir , Isla, 21.ix.1996, S. M. Clark [16, BYUC]; San Jacinto Co. , Point Blank , 22.ix.1996, S. M. Clark [266, BYUC]; San Patricio Co. , Welder Wildlife Refuge , 12-13.x.1988, E. G. Riley [16 5♀♀, TAMU]; same data, except nr. Venado Mill , 1-2. xi.1988, R. Anderson [10♀♀, TAMU]; Starr Co. , 2 mi NE La Gloria, 10.iv.1994, E. G. Riley [366 1♀, TAMU]; Taylor Co. , 21 mi S Abilene on 83, 1. viii.1989, R. Morris [16, UGCA]; Victoria Co. , rest stop, Hwy . 59 N, 2.5 air mi SW Inez, 20.iv.2009, J. Jones [16, TAMU]; Washington Co. , 15.8 mi W jct. Hwy . 36 & FM 1948, 8.vi.1982, ex. Ptilimniumn, S. J. Merritt [16, TAMU]; Wichita Co. , Burkburnett , Red River plain, 6.vi.2007, R. J. Barney [16 1♀, RJBC]; Willacy Co., 7 mi N Raymondville, 18.iii.1970, ex. Elocharis, H. R. Burke [3♀♀, TAMU].

APPENDIX 6