Galleruca humeralis Fabricius, 1801

Roie, Martijn Van, Kuhlmann, Michael, Mack, Anne & Konstantinov, Alexander, 2024, Fabrician types of new world Oedionychina Chapuis, 1875 (Coleoptera, Chrysomelidae, Alticini) deposited in the Zoological Museum of Kiel University collections with notes on Fabrician types of other collections and new combinations for species formerly placed in the subtribe, European Journal of Taxonomy 920 (1), pp. 1-60 : 27-32

Galleruca humeralis Fabricius, 1801

Figs 18–21

humeralis Fabricius, 1801: 494 (America meridionali).

Material examined

Lectotype (presently designated)

COUNTRY NOT INDICATED ON SPECIMEN • ♀; “ G. humeralis en Am. mer. Schmidt/ TYPE/ Lectotype Galleruca humeralis Fabricius 1801 des. Van Roie, Kuhlmann, Mack & Konstantinov, 2023”; ex-Copenhagen; ZMUC 00031095 .

Specimens probably misplaced

COUNTRY NOT INDICATED ON SPECIMEN • 2 specs; “[humeralis]/ [red squared label]”; Kiel . • 1 ♂; “ TYPE ”; ex-Copenhagen; ZMUC 00031094 .

Status in Zimsen (1964)

P. 111 No.1722. “in America meridionali Dom. Smidt Mus. D. de Sehestedt” Copenhagen 2 specimens, Kiel 2 specimens.

Original description

“G. saltatoria, pallide testacea, elytris puncto humerali nigro. Habitat in America meridionali. D. Smidt. Mus. D. de Sehestedt. Statura et magnitude omnino G. sellatae. Antennae pallidae. Caput et thorax pallidiora. Elytra laevia, pallide testacea, puncto humerali atro. Corpus testaceum, femoribus valde incrassatis. Variat elytris immaculatis. ”

Current status

Walterianella humeralis ( Fabricius, 1801) .

Measurements

Lectotype ( Fig. 18 ): LB = 5.97 mm; WB = 3.40 mm. Misplaced specimen 1 ( Fig. 19 ): LB = 4.39 mm; WB = 2.66 mm. Misplaced specimen 2 ( Fig. 20 ): LB = 4.58 mm; WB = 2.65 mm. Misplaced specimen 3 ( Fig. 21 ): LB = 3.06 mm; WB = 1.88 mm.

Remarks

The lectotype for Galleruca humeralis is designated here to have a unique bearer of this name and the standard for its application. We chose the female, and not the male, as lectotype because it conforms closely with the original description stating ‘elytra with a black humeral point’. The male specimen from the Copenhagen collection does show some differences from the female lectotype: humeral calli more developed, without dark spot, elytral edges more explanate, less rounded apically; basal part of pronotum with dark coloration, pronotal edges angulated. This could mean that there are two species included in this type series (Fabricius also did not mention a variety with dark pronotal base, only a variety without elytral spots). Although sexual dimorphism is not uncommon in Walterianella (e.g., Bechyné & Bechyné 1977), we chose to be conservative and not designate the male specimen as paralectotype. We confirm the correct placement in the genus Walterianella based on the following characters: eyes big, interocular space about as wide as diameter of eye, pronotal borders widely explanate, flattened, prosternal process keeled. Two specimens from Kiel collection ( Figs 20–21 ) do not conform at all with the description of Fabricius (1801) and are very likely misplaced. They belong to Altica .

Figures

Fig. 18. Galleruca humeralis Fabricius, 1801, lectotype, ♀ (ZMUC 00031095). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E. Labels. Scale bars = 1 mm.

Fig. 19. Galleruca humeralis Fabricius, 1801, misplaced specimen, ♂ (ZMUC 00031094). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E. Labels. Scale bars = 1 mm.

Fig. 20. Galleruca humeralis Fabricius, 1801, misplaced specimen, sex unknown (Kiel). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E. Labels. Scale bars = 1 mm.

Fig. 21. Galleruca humeralis Fabricius, 1801, misplaced specimen, sex unknown (Kiel). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E. Labels. Scale bars = 1 mm.