Passalus (Pertinax) rugosus Gravely, 1918

Jimenez-Ferbans, Larry, Maestre-Guerra, Ana, Villalba-Fuentes, Evelin, Barros-Barrios, Mayelis M. & Munoz-Montero, Jeison, 2023, Passalidae (Coleoptera, Scarabaeoidea) from the Caribbean coast of Colombia: synopsis, key, and new species description, ZooKeys 1179, pp. 243-297 : 243

11. Passalus (Pertinax) rugosus Gravely, 1918

Fig. 14

Diagnosis.

20.1-23.2 mm total length. Body subcylindrical. Anterior border of the frons with strong middle indentation, suggesting secondary mediofrontal tubercles. Mediofrontal and laterofrontal fused, large. Central tubercle with apex not free. Lateroposterior tubercles large, conspicuous, and conical. Eyes large. Antennal club tri-lamellate, with lamellae long. Lacinia with apex bidentate. Mediobasal area of mentum protruding and glabrous. Marginal groove narrow, occupying 2/3 of the anterior margin of the pronotum. Prosternellum rhomboidal, acute. Mesosternum glabrous; mesosternal scars distinct and elongated. Metasternum glabrous anterolaterally and in lateral groove; disc smooth and delimited by punctations posteriorly to laterally. Humeri pubescent and epipleura pubescent in basal third. Anterior ventral border of the profemur with conspicuous groove. Meso- and metatibiae with small spines or unarmed.

Comments.

Jiménez-Ferbans et al. (2018b) considered this species as endemic to the western Andes of Colombia. However, the present records extend its distribution to the eastern Andes.

Figures

Figure 14. Passalus (Pertinax) rugosus Gravely, 1918 A head and pronotum in dorsal view B habitus dorsal C habitus ventral D habitus lateral. Scale bars: 1.0 mm (A); 2.0 mm (B, C, D).