Edaphus longicollis, Bernhauer, 1929

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium IV. Staphylinine Group (Part 1) Euaesthetinae, Leptotyphlinae, Megalopsidiinae, Oxyporinae, Pseudopsinae, Solieriinae, Steninae, Bulletin of the American Museum of Natural History 2440 (265), pp. 1807-2440 : 1807-2440

longicollis Bernhauer, 1929 , see: heikertingeri Bernhauer, 1929 .

lorianus Puthz, 1990b: 278 ( Edaphus ; Type locality: N. Guinea mer.: Bujakori [nahe Kemp- Welch-River, etwa 147°46′E / 9°42′S]).

- DISTRIBUTION: New Guinea.

luberoensis Puthz, 1992b: 19 ( Edaphus ; Type locality: Zaire: Forêt de Kasuo SW de Lubero). - DISTRIBUTION: Congo.

lubukensis Kistner, 1962a: 627 ( Edaphus ; Type locality: Congo Republic: Kivu, Uvira territory, Lubuka , 2780 m).

— Puthz, 1974s: 760 ( Edaphus ; characters).

- DISTRIBUTION: Congo.

luculentus Casey, 1884 , see: nitidus Motschulsky, 1857 .

luridus Blattny´, 1925, see: dilutus Schaufuss, 1887 .

luzonicus Bernhauer, 1929c: 94 , 98 ( Edaphus ; Type locality: Luzon: Imugan).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

- DISTRIBUTION: Philippines.

lynceus Puthz, 1990a: 210 ( Edaphus ; Type locality: Côte d’Ivoire: Abidjan: Parc national du Banco).

- DISTRIBUTION: Ivory Coast.

mabaliensis Kistner, 1962a: 628 ( Edaphus ; Type locality: Congo Republic: Equateur, Tshuapa , Mabali near Lake Tumba, 350 m).

— Puthz, 1974s: 760 ( Edaphus ; characters).

- DISTRIBUTION: Congo.

magnipennis Bernhauer, 1929 , see: dilutus Schaufuss, 1887 .

mahnerti Puthz, 1990a: 214 ( Edaphus ; Type locality: Côte d’Ivoire: Forêt de Taï , env. de la Station).

- DISTRIBUTION: Ivory Coast.

major Fauvel, 1895: 205 ( Edaphus ; Type locality: Bírmanie, Bhamo ).

— Bernhauer and Schubert, 1911: 188 ( Edaphus ; catalog).

— Cameron, 1930: 448 ( Edaphus ; characters; Burma).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

— Coiffait, 1978b: 112 ( Edaphus ; Bhutan).

— Puthz, 1979: 158 ( Edaphus ; characters).

— Puthz, 1979b: 161 ( Edaphus ; characters; Burma).

- DISTRIBUTION: Bhutan, Myanmar.

mandibularis Cameron, 1930 , see: dentiventris Fauvel, 1905 .

marlieri Kistner, 1962 , see: africanus Eppelsheim, 1885 .

marshalli Bernhauer, 1934 , see: africanus Eppelsheim, 1885 .

martensi Puthz, 1987a: 449 ( Edaphus ; Type locality: Nepal: Parbat Distr. , Chitre, 2400 m).

- DISTRIBUTION: Nepal.

mediopunctus Puthz, 1974 s: 64 ( Edaphus ; Type locality: Congo-Brazzaville: Galerienwald bei Bampaka, westlich von Brazzaville, Talmulde).

— Puthz, 1984g: 202 ( Edaphus ; Ghana).

- DISTRIBUTION: Congo, Ghana.

melculus Oke, 1933: 110 ( Edaphellus ; Type locality: Australia: Victoria: Belgrave, Gembrook, Warburton , Preston , Macedon ).

— Puthz, 1978c: 124 ( Edaphus ; characters; Australia).

- DISTRIBUTION: Australia.

mendax Puthz, 1979: 137 ( Edaphus ; Type locality: Singapore. Also cited from Siam).

- DISTRIBUTION: Singapore, Thailand.

mendicus Puthz, 1979: 116 ( Edaphus ; Type locality: Zentral-Nepal : Pande-Sattel zwischen Biritanti und Naudanda westlich Pokhara, ca. 1800 m. Also cited from Thailand).

- DISTRIBUTION: Nepal, Thailand.

mendosus Puthz, 1998b: 52 ( Edaphus ; Type locality: China: Hongkong: Kadoorie Farm ).

- DISTRIBUTION: China.

meo Orousset, 1988c: 186 ( Edaphus ; Type locality: Thaïlande: env. de Chiang Maï, chiang Dao , alt. 500 m).

- DISTRIBUTION: Thailand.

mexicanus Puthz, 1973 k: 54 ( Edaphus ; Type locality: Mexico: Chiapas, 8 mi. S. Simojovel). - DISTRIBUTION: Mexico.

microphthalmus Puthz, 1981 g: 455 ( Edaphus ; Type locality: Nepal: Taoundja, massif du Lamjung Himal, 2860 m).

- DISTRIBUTION: Nepal.

microps Puthz, 1976f: 156 ( Edaphus ; Type locality: Ceylon: Sabaragamuwa, Ratuapura ).

- DISTRIBUTION: Sri Lanka.

micros Coiffait, 1981d: 332 ( Edaphus ; Type locality: Jagath, 1650 m).

- DISTRIBUTION: Nepal.

mindoroensis Bernhauer, 1929c: 96 , 106 ( Edaphus ; Type locality: Mindoro: S. Theodoro). — Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

- DISTRIBUTION: Philippines.

minutus Puthz, 1979: 130 ( Edaphus ; Type locality: Nepal: Phulchoki , Ost-Hang, ca. 2000 m, bei Dalikhel).

- DISTRIBUTION: Nepal.

mjobergi Bernhauer, 1916a: 3 ( Edaphus ; cited as mjöbergi ; Type locality: Queensland: Blackal Range).

— Scheerpeltz, 1933: 1204 ( Edaphus ; cited as mjöbergi ; catalog).

— Oke, 1933: 110 ( Edaphus ; cited as mjobergi ; notes on description).

— Puthz, 1978c: 126 ( Edaphus ; cited as mjöbergi ; characters; Australia).

— Herman, 2001: 12 ( Edaphus ; spelling corrected to mjobergi ).

- DISTRIBUTION: Australia.

moloensis Puthz, 1974 s: 756 ( Edaphus ; Type locality: Kenya: Molo , Mau Escarpment, 2150–2200 m).

- DISTRIBUTION: Kenya.

monopticus Puthz, 1992b: 20 ( Edaphus ; Type locality: Zaire: Kivu : Itombwe, 2300 m., Mulenge, Nyalengwe).

- DISTRIBUTION: Congo.

monticola Puthz, 1975c: 177 ( Edaphus ; Type locality: China: West-Szechuan : Wassuland , Bezirk San-kiang-kou, Strecke S-S bis 3000 m).

- DISTRIBUTION: China.

montisamator Comellini, 1977: 266 ( Edaphus ; Type locality: Sri Lanka: province Central , Pidurutalagala, 2200 m).

— Puthz, 1979: 118 ( Edaphus ; characters).

- DISTRIBUTION: Sri Lanka.

montius Comellini, 1977: 263 ( Edaphus ; Type locality: Sri Lanka: province Central , Horton Plains, 2100 m).

- DISTRIBUTION: Sri Lanka.

mussardi Comellini, 1977: 252 ( Edaphus ; Type locality: Sri Lanka: province North Western, Rajakadaluwa ).

— Puthz, 1979: 154 ( Edaphus ; characters).

- DISTRIBUTION: Sri Lanka.

najtae Orousset, 1986e: 75 ( Edaphus ; Type locality: Venezuela: Cordillera de Mérida: Sierra de la Culata, páramo de Piedras Blancas, alt. 4250 m).

- DISTRIBUTION: Venezuela.

nakayamaensis Puthz, 1980 h: 243 ( Edaphus ; Type locality: Japan: Hokkaido: Nakayama-Pass südlich Sapporo , über 1000 m).

- DISTRIBUTION: Japan.

napporoensis Puthz, 1980 h: 244 ( Edaphus ; Type locality: Japan: Hokkaido: Napporo National Park bei Sapporo , 60–70 m).

- DISTRIBUTION: Japan.

neboissi Puthz, 1978c: 123 ( Edaphus ; Type locality: S. Australia: Englebrook Nat-Trust Res. near Adelaide. Also cited from Victoria).

- DISTRIBUTION: Australia.

nepalensis Puthz, 1979: 121 ( Edaphus ; Type locality: Nepal: Barahbise gegen Ting Sang La [Wald oberhalb Durumtali am Aufstieg zum Ting Sang La, 2200–2300 m]).

- DISTRIBUTION: Nepal.

nepalicus Coiffait, 1982: 104 ( Edaphus ; Type locality: Nepal central: Madi Khola , 2385 m).

- DISTRIBUTION: Nepal.

nequam Puthz, 1990a: 202 ( Edaphus ; Type locality: Côte d’Ivoire: Abidjan: Parc national du Banco).

- DISTRIBUTION: Ivory Coast.

nipponensis Puthz, 1975c: 181 ( Edaphus ; Type locality: Japan: Kanagawa).

- DISTRIBUTION: Japan.

nitidifrons Puthz, 1990c: 816 ( Edaphus ; Type locality: Sumatra: Aceh-Selatan, Bahahrot ,

100 m).

- DISTRIBUTION: Indonesia.

nitidus LeConte, 1861 , see: nitidus Motschulsky, 1857 .

nitidus Motschulsky, 1857: 7 ( Edaphus ; Type locality: Am. bor.; [Note: See lectotype designation by Puthz, 1974t]).

— Bernhauer and Schubert, 1911: 188 ( Edaphus ; catalog).

— Kistner, 1962a: 621 ( Edaphus ; characters; Pennsylvania; Illinois; Indiana; Washington D.C.; Louisiana; Mississippi).

— Puthz, 1974t: 915 ( Edaphus ; lectotype designation: Am. bor.; characters; notes; Pennsylvania, Indiana, Illinois, Iowa, District of Columbia, Virginia, West Virginia, North Carolina, South Carolina, Georgia, Florida, Kentucky, Louisiana, Texas) .

— Downie and Arnett, 1996: 461 ( Edaphus ; characters; USA).

- DISTRIBUTION: USA.

nitidus LeConte, 1861a: 68 ( Edaphus ; [preoccupied]; Type locality: Louisiana).

— LeConte, 1863: 25 ( Edaphus ; North America).

— LeConte, 1863a: 50 ( Edaphus ; characters; Alabama, Louisiana).

— Casey, 1884a: 29 ( Edaphus ; characters; Alabama; Louisiana).

— Horn, 1885b: 121 ( Edaphus ; characters).

— Herman, 2001: 00 ( Edaphus ; synonym of nitidus Motschulsky ).

carinatus Casey, 1884a: 30 ( Edaphus ; Type locality: Washington, D.C.).

— Horn, 1885b: 121 ( Edaphus ; synonym of nitidus ).

— Horn, 1885a: 109 ( Edaphus ; synonym of nitidus ).

— Bernhauer and Schubert, 1911: 188 ( Edaphus ; synonym of nitidus ).

— Kistner, 1962a: 621 ( Edaphus ; synonym of nitidus ).

— Puthz, 1974t: 916 ( Edaphus ; synonym of nitidus ).

luculentus Casey, 1884a: 31 ( Edaphus ; Type locality: District of Columbia; [Note: See lectotype designation by Puthz, 1974t]).

— Horn, 1885b: 121 ( Edaphus ; characters).

— Horn, 1885a: 109 ( Edaphus ; synonym of nitidus ).

— Bernhauer and Schubert, 1911: 188 ( Edaphus ; synonym of nitidus ).

— Kistner, 1962a: 621 ( Edaphus ; synonym of nitidus ).

— Puthz, 1974t: 916 ( Edaphus ; lectotype designation; synonym of nitidus ).

novaeguineae Fauvel, 1878f: 220 ( Edaphellus ; Type locality: Nouvelle-Guinée, Hatam).

— Bernhauer and Schubert, 1911: 188 ( Edaphellus ; catalog).

— Székessy, 1939: 98 ( Edaphus ; characters; New Guinea).

— Puthz, 1990: 275 ( Edaphus ; characters; New Guinea).

- DISTRIBUTION: New Guinea.

noviesimpressus Puthz, 1998b: 53 ( Edaphus ; Type locality: China: Hongkong: Kadoorie Farm ). - DISTRIBUTION: China.

octopunctatus Kistner, 1962a: 628 ( Edaphus ; Type locality: Congo Republic: Kivu, counterfort S. Kahuzi, 2300 m).

— Puthz, 1974s: 761 ( Edaphus ; characters).

- DISTRIBUTION: Congo.

oculatus Jarrige, 1963a: 122 ( Edaphus ; Type locality: Ceylan: I d’Alugtgama).

— Puthz, 1979: 156 ( Edaphus ; characters).

- DISTRIBUTION: Sri Lanka.

okei Puthz, 1979a: 75 ( Edaphus ; characters published under “ Edaphus D” by Puthz, 1978c: 123, 129; Type locality: Queensland: Dividing Range, highway to Warwick, 800 m). - DISTRIBUTION: Australia.

okinawaensis Puthz, 1980 h: 241 ( Edaphus ; Type locality: Okinawa: Nishime-dake ).

- DISTRIBUTION: Japan.

ophthalmicus Bernhauer, 1929c: 95 , 102 ( Edaphus ; Type locality: Mindanao: Surigao).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

- DISTRIBUTION: Philippines.

pahangensis Puthz, 1979: 119 ( Edaphus ; Type locality: Malaya: Pahang, Cameron Highlands, Ringlet , 3200 feet).

- DISTRIBUTION: Malaysia.

papuanus Székessy, 1939: 100 ( Edaphus ; Type locality: Neu-Guinea: Stephansort, Astrolab-Bay).

- DISTRIBUTION: New Guinea.

parvipennis Bernhauer, 1929c: 96 , 109 ( Edaphus ; Type locality: Luzon: Imugan).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

- DISTRIBUTION: Philippines.

peninsularis Puthz, 1979: 119 ( Edaphus ; Type locality: Malaya: Pahang, Temerloh District, Krau Game Reserve ).

— Puthz, 1990c: 823 ( Edaphus ; characters; Malaya; Sumatra).

- DISTRIBUTION: Indonesia, Malaysia.

perminimus Puthz, 1990c: 822 ( Edaphus ; Type locality: West-Sumatra: Indrapoera ).

- DISTRIBUTION: Indonesia.

permodestus Puthz, 1990a: 203 ( Edaphus ; Type locality: Côte d’Ivoire: Monogaga ).

— Puthz, 1992b: 17 ( Edaphus ; characters).

- DISTRIBUTION: Ivory Coast.

perpusillus Coiffait, 1981d: 333 ( Edaphus ; Type locality: Kali Gandaki, 1900 m).

- DISTRIBUTION: Nepal.

perreti Puthz, 1990a: 218 ( Edaphus ; Type locality: Côte d’Ivoire: Forêt de Taï ).

— Puthz, 1992b: 18 ( Edaphus ; characters).

- DISTRIBUTION: Ivory Coast.

persimilis Puthz, 1992b: 15 ( Edaphus ; Type locality: Zaire: Kwango, terr. de Feshi,

rive gauche Kwenge).

peruanus Puthz, 1973 l: 148 ( Edaphus ; Type locality: Peru: Umgebung Tarapoto , Strasse nach Yutimagua am Huallaga).

- DISTRIBUTION: Peru.

perversus Puthz, 1994e: 390 ( Edaphus ; replacement name for japonicus Bernhauer ).

- DISTRIBUTION: Japan.

japonicus Bernhauer, 1907: 380 ( Euaesthetus ; preoccupied; Type locality: Ocyama,

2000 feet).

— Bernhauer and Schubert, 1911: 189 ( Euaesthetus ; catalog).

— Puthz, 1994e: 390 ( Edaphus ; synonym of perversus ).

petzi Bernhauer, 1929c: 94 , 98 ( Edaphus ; Type locality: Nonoc; Luzon: Balbalan).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

- DISTRIBUTION: Philippines.

peyrierasi Orousset, 1988: 42 ( Edaphus ; Type locality: Madagascar Est: col de Madiorano,

à l’Est de Vondrozo, 700 m).

- DISTRIBUTION: Madagascar.

philippinus Bernhauer, 1929c: 96 , 109 ( Edaphus ; Type locality: Luzon: Balbalan, Manila; Siargao : Dapa ).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

- DISTRIBUTION: Philippines.

phloeocharis Puthz, 1986d: 619 ( Edaphus ; Type locality: Paraguay: Provinz Itapua, 10 km sud de Santa Maria).

- DISTRIBUTION: Paraguay.

piceus Cameron, 1933 , see: beszedesi Reitter, 1914 .

pilosus Puthz, 1979: 124 ( Edaphus ; Type locality: Nepal: Barahbise gegen Ting Sang La [Wald oberhalb Durumtali am Aufstieg zum Ting Sang La], 2200–2300 m).

- DISTRIBUTION: Nepal.

planifrons Puthz, 1990f: 260 ( Edaphus ; Type locality: Brasilien: Floresta de Tijuca bei

Rio de Janeiro, N. P. Alto de Boavista).

- DISTRIBUTION: Brazil.

planus Puthz, 1979: 140 ( Edaphus ; Type locality: Vietnam: Cuc Phuong, Ninh binh. Also cited from Thailand).

— Puthz, 1998b: 53 ( Edaphus ; China).

- DISTRIBUTION: China, Vietnam, Thailand.

plaumanni Puthz, 1978e: 213 ( Edaphus ; Type locality: Brazil: Santa Catarina , Nova Teutonia).

- DISTRIBUTION: Brazil.

plicatulus Schaufuss, 1877: 25 ( Tetratarsus ; Type locality: Bangkok, Siam; [ Note : See lectotype designation by Puthz, 1974r]).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

— Puthz, 1974r: 83 ( Edaphus ; lectotype designation: Bangkok, Siam; characters) .

— Puthz, 1979: 156 ( Edaphus ; characters).

- DISTRIBUTION: Thailand.

plicifer Puthz, 1979: 136 ( Edaphus ; Type locality: Malaya: Perak, Cameron Highlands, Cascade Sungei Simei ).

- DISTRIBUTION: Malaysia.

plicipennis Puthz, 1978c: 42 ( Edaphus ; Type locality: Neuguinea: Papua: near Kokoda ,

500 m).

- DISTRIBUTION: Papua New Guinea.

politus Motschulsky, 1857: 7 ( Edaphus ; Type locality: Nouvelle-Orléans).

— Puthz, 1974t: 915, 918 ( Edaphus ; characters; notes; Kentucky; Louisiana; Texas).

poulaini Orousset, 1986e: 78 ( Edaphus ; Type locality: Pérou: Loreto, Genaro Herrera ,

Río Ucayali, alt. 200 m).

- DISTRIBUTION: Peru.

praefectus Puthz, 1990f: 259 ( Edaphus ; Type locality: Ecuador: prov. Imbabura, Mojanda nord, 3470 m).

- DISTRIBUTION: Ecuador.

princeps Bernhauer, 1929c: 95 , 100 ( Edaphus ; Type locality: Siargao: Dapa).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

- DISTRIBUTION: Philippines.

bakerianus Bernhauer, 1929c: 100 ( Edaphus ; subspecies of princeps Type locality: Luzon: Mt. Makiling).

— Scheerpeltz, 1933: 1204 ( Edaphus ; variety of princeps ).

pselaphoides Eichelbaum, 1913: 122 ( Edaphus ; Type locality: Amani, Berg Bomole ).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

— Kistner, 1960: 31 ( Edaphus ; Tanganyika).

— Kistner, 1962a: 632 ( Edaphus ; incertae sedis).

— Puthz, 1975d: 39 ( Edaphus ; characters; Tanzania).

— Puthz, 1979b: 172 ( Edaphus ; characters; Kenya).

- DISTRIBUTION: Kenya, Tanzania.

pubescens Bernhauer, 1929c: 96 , 105 ( Edaphus ; Type locality: Luzon: Mt. Polis, Balbalan). — Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

- DISTRIBUTION: Philippines.

pulliclavis Puthz, 1990c: 821 ( Edaphus ; Type locality: Sumatra: Aceh-Selatan, Babahrot , 100 m).

- DISTRIBUTION: Indonesia.

pumilus Puthz, 1979: 138 ( Edaphus ; Type locality: Malaya: Gombak near Kuala Lumpur, ca. 800 feet).

- DISTRIBUTION: Malaysia.

punctatus Fauvel, 1895: 206 ( Edaphus ; Type locality: Pulo-Penang; [Note: See lectotype designation by Puthz , 1979b]. Also cited from Bírmanie, Bhamo).

— Bernhauer and Schubert, 1911: 188 ( Edaphus ; catalog).

— Cameron, 1930: 449 ( Edaphus ; characters; Burma; Penang).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

— Puthz, 1979b: 162 ( Edaphus ; lectotype designation: Penang; characters).

— Coiffait, 1981d: 324 ( Edaphus ; Nepal).

- DISTRIBUTION: Myanmar, Malaysia, Nepal.

punctatus Székessy, 1939 , see: szekessyi Puthz, 1971 .

puncticeps Puthz, 1974 s: 765 ( Edaphus ; Type locality: Congo-Brazzaville: Galerienwald bei Bampaka, westlich Brazzaville, Talmulde).

— Puthz, 1984g: 202 ( Edaphus ; Zaire).

- DISTRIBUTION: Congo.

puncticollis Puthz, 1979: 134 ( Edaphus ; Type locality: Thailand: Sakaerat Experimental Station ).

- DISTRIBUTION: Thailand.

punctulatus Puthz, 1979: 131 ( Edaphus ; Type locality: India: Darjeeling District , Chim- Khona, Ghum, 2200 m).

- DISTRIBUTION: India.

punctuliceps Puthz, 1998b: 54 ( Edaphus ; Type locality: China: Hongkong: Lantau Island : Tei Tong Tsai).

- DISTRIBUTION: China.

punctulifrons Puthz, 1990a: 199 ( Edaphus ; Type locality: Côte d’Ivoire: Forêt de Taï , env. de la Station).

- DISTRIBUTION: Ivory Coast.

puriventris Puthz, 1992b: 14 ( Edaphus ; Type locality: Rwanda: Forêt de la Rugege, 2150 m). - DISTRIBUTION: Rwanda.

puthzi Comellini, 1977: 254 ( Edaphus ; Type locality: Sri Lanka: province Western, Colombo). — Puthz, 1979: 154 ( Edaphus ; characters).

- DISTRIBUTION: Sri Lanka.

pygmaeus Székessy, 1939: 100 ( Edaphus ; Type locality: Neu-Guinea: Friedrich-Wilhelms-Hafen, Astrolab-Bay).

- DISTRIBUTION: New Guinea.

quadripunctus Kistner, 1962: 104 ( Edaphus ; Type locality: Angola: Parque Carisso, Dundo). — Kistner, 1962a: 629 ( Edaphus ; characters).

— Kistner, 1963: 199 ( Edaphus ; Congo Republic).

— Puthz, 1974s: 761 ( Edaphus ; characters).

— Puthz, 1984g: 198 ( Edaphus ; characters; Angola; Zaire; Ghana).

- DISTRIBUTION: Angola, Congo, Ghana.

quisquilius Puthz, 1990f: 262 ( Edaphus ; Type locality: Ecuador: Pichin [.], Rio Palenque).

- DISTRIBUTION: Ecuador.

rastellus Comellini, 1977: 260 ( Edaphus ; Type locality: Sri Lanka: province Central , Hasalaka près Weragamtola).

— Puthz, 1979: 140 ( Edaphus ; characters).

- DISTRIBUTION: Sri Lanka.

remilleti Orousset, 1983b: 116 ( Edaphus ; Type locality: Mont Grand-Matoury, 7 km S. de Cayenne, alt. 200 m, Guyane française).

- DISTRIBUTION: French Guiana.

remyi Jarrige, 1963a: 120 ( Edaphus ; Type locality: Ceylan: Ratnapura ).

— Puthz, 1979: 132 ( Edaphus ; characters).

— Orousset, 1988: 47 ( Edaphus ; characters; Sri Lanka).

- DISTRIBUTION: Sri Lanka; Réunion.

batrisoides Comellini, 1977: 256 ( Edaphus ; Type locality: Sri Lanka: province Western , Colombo).

— Puthz, 1979: 132 ( Edaphus ; synonym of remyi ).

— Orousset, 1988: 47 ( Edaphus ; synonym of remyi ).

etiennei Orousset, 1988: 50 ( Edaphus ; subspecies of remyi ; Type locality: La Réunion:

forêt de Mare Longue).

reticulatus Puthz, 1987: 30 ( Edaphus ; Type locality: Äthiopien: Chillalo , ca. 9000 feet).

- DISTRIBUTION: Ethiopia.

riedeli Puthz, 1994d: 385 ( Edaphus ; Type locality: Indien: Kerala: Thekkady Peryar, W.L.S.). - DISTRIBUTION: India.

rishwani Makhan, 1995 , see: humeralis Puthz, 1973 .

riukiuensis Puthz, 1980 h: 242 ( Edaphus ; Type locality: Okinawa: Kanna ).

- DISTRIBUTION: Japan.

rivalis Puthz, 1979: 133 ( Edaphus ; Type locality: Malaya: Gombak , near Kuala Lumpur,

ca. 800 feet).

- DISTRIBUTION: Malaysia.

rosskotheni Wüsthoff, 1935 , see: beszedesi Reitter, 1913 .

rougemonti Puthz, 1998b: 54 ( Edaphus ; Type locality: China: Hongkong: Tai Po ).

- DISTRIBUTION: China.

ruandae Puthz, 1992b: 24 ( Edaphus ; Type locality: Ruanda: Forêt de la Rugege, 2150 m).

- DISTRIBUTION: Rwanda.

ruwenzorianus Puthz, 1983f: 175 ( Edaphus ; Type locality: Zaire: Face N. Ruwenzori, Kilindera, 2750–2950 m).

- DISTRIBUTION: Congo.

scheerpeltzi Bernhauer, 1929 , see: sulciceps Bernhauer, 1929 .

scherpa Puthz, 1980 i: 246 ( Edaphus ; Type locality: Nepal: Umgebung Pokhara : Phewa-See , Wald am Abhang gegenüber dem Tempelhügel bei Piota).

— Puthz, 1987a: 452 ( Edaphus ; Nepal).

- DISTRIBUTION: Nepal.

sechellarum Bernhauer, 1922b: 170 ( Edaphus ; Type locality: Mahé; [Note: See lectotype designation by Orousset , 1985b]. Originally cited from Seychelles: Silhouette : high forest above Mare aux Cochons and Mahé: near Morne Blanc).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

— Kistner, 1962a: 629 ( Edaphus ; characters).

— Orousset, 1985b: 250 ( Edaphus ; lectotype designation: Mahé, Seychelles) .

— Orousset, 1988: 52 ( Edaphus ; characters; Seychelles).

- DISTRIBUTION: Seychelles.

setosus Puthz, 1979: 135 ( Edaphus ; Type locality: Nepal: Sanghu Base , Camp area,

ca. 6500 feet, 27°21′N, 87°33′E).

- DISTRIBUTION: Nepal.

sexangulicollis Puthz, 1990b: 282 ( Edaphus ; Type locality: D’Entrecasteaux Islands: Fergusson Island: Deidei , Gomwa Bay, 0–20 m, Camp 6).

- DISTRIBUTION: Papua New Guinea.

sexpunctus Kistner, 1962 , see: africanus Eppelsheim, 1885 .

siamensis Puthz, 1979: 137 ( Edaphus ; Type locality: Thailand: Sakaerat Experimental Station ).

- DISTRIBUTION: Thailand.

similigenus Puthz, 1992b: 16 ( Edaphus ; Type locality: Zaire: Tshuapa , terr. Bikoro, Mabali). - DISTRIBUTION: Congo.

similior Puthz, 1979: 115 ( Edaphus ; Type locality: India: W. Bengal: Darjeeling District , North Point, 1500 m. Also cited from Nepal).

- DISTRIBUTION: India, Nepal.

similis Puthz, 1979a: 75 ( Edaphus ; characters published under “ Edaphus E” by Puthz, 1978c: 127, 130; Type locality: Queensland: Dividing Range, highway to Warwick, 800 m). - DISTRIBUTION: Australia.

simillimus Puthz, 1992b: 16 ( Edaphus ; Type locality: Zaire: Thysville ).

— Puthz, 1983f: 173 ( Edaphus ; [Note: Misidentified as franzi according to Puthz, 1992b: 16]; characters; Zaire).

- DISTRIBUTION: Congo.

singaporensis Puthz, 1979: 126 ( Edaphus ; Type locality: Singapore: Bukit Timah ).

- DISTRIBUTION: Singapore.

sparsus Puthz, 1979: 117 ( Edaphus ; Type locality: Malaya: Gombak near Kuala Lumpur,

ca. 900 feet. Also cited from Singapore).

- DISTRIBUTION: Malaysia, Singapore.

spectabilis Bernhauer, 1922b: 169 ( Edaphus ; Type locality: Praslin , Seychelles; [Note: See lectotype designation by Orousset, 1986]. Originally cited from Seychellen: Praslin: Côtes d’Or Estate; Silhouette: Mare aux Cochons).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

— Kistner, 1962a: 631 ( Edaphus ; characters).

— Orousset, 1986: 402 ( Edaphus ; lectotype designation: Praslin , Seychelles) .

— Orousset, 1988: 50 ( Edaphus ; characters; Seychelles).

- DISTRIBUTION: Seychelles.

spiniphallus Orousset, 1988: 43 ( Edaphus ; Type locality: Madagascar Centre: forêt d’Ambohiboatavo, à l’Est du lac de Mantasoa, 1340 m).

- DISTRIBUTION: Madagascar.

strangulatus Cameron, 1929b: 452 ( Edaphus ; Type locality: Penang).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

— Puthz, 1979: 158 ( Edaphus ; characters).

— Puthz, 1979b: 165 ( Edaphus ; characters).

- DISTRIBUTION: Malaysia.

subbiformis Puthz, 1992b: 26 ( Edaphus ; Type locality: Zaire: Kivu , terr. Mwenga,

riv. Lubushwa, 650 m).

- DISTRIBUTION: Congo.

subcarinatus Puthz, 1979: 118 ( Edaphus ; Type locality: Süd-Thailand: Kachong Experimental Station, 80 m).

- DISTRIBUTION: Thailand.

sulciceps Bernhauer, 1929c: 95 , 104 ( Edaphus ; Type locality: Leyte).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

— Puthz, 1979: 142 ( Edaphus ; characters).

- DISTRIBUTION: Philippines.

scheerpeltzi Bernhauer, 1929c: 95 , 101 ( Edaphus ; Type locality: Biliran).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

— Puthz, 1979: 142 ( Edaphus ; synonym of sulciceps ).

sumatrensis Schaufuss, 1887: 110 ( Edaphus ; Type locality: Sumatra).

— Bernhauer and Schubert, 1911: 188 ( Edaphus ; catalog).

— Cameron, 1927c: 261 ( Edaphus ; Fiji; Sumatra; Timor).

— Cameron, 1930d: 330 ( Edaphus ; Sumatra).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

— Puthz, 1974r: 88 ( Edaphus ; lectotype designation; characters).

— Puthz, 1979: 156 ( Edaphus ; characters).

— Puthz, 1990c: 819 ( Edaphus ; characters; Sumatra).

- DISTRIBUTION: Indonesia, Fiji.

sybaris Puthz, 1979: 120 ( Edaphus ; Type locality: India: W. Bengal: Darjeeling, North Point , 1300 m).

- DISTRIBUTION: India.

szechuanus Puthz, 1975c: 182 ( Edaphus ; Type locality: China: West-Szechuan , Wassuland, Bezirk San-kiang-kou, 1000 m).

— Puthz, 1979: 155 ( Edaphus ; characters).

- DISTRIBUTION: China.

szekessyi Puthz, 1971 p: 203 ( Edaphus ; replacement name for punctatus Székessy ).

- DISTRIBUTION: New Guinea.

punctatus Székessy, 1939: 104 ( Edaphus ; [preoccupied]; Type locality: Neu-Guinea: Marega).

— Puthz, 1971p: 203 ( Edaphus ; synonym of szekessyi ).

taiwanensis Puthz, 1975c: 182 ( Edaphus ; Type locality: Formosa: Takao , 300 m).

- DISTRIBUTION: Taiwan.

tamoul Comellini, 1977: 263 ( Edaphus ; Type locality: Sri Lanka: province Sabaragamuwa, entre Ambepussa et Polganawela).

— Puthz, 1979: 139 ( Edaphus ; characters).

- DISTRIBUTION: Sri Lanka.

tasmani Puthz, 1991: 284 ( Edaphus ; Type locality: Fiji: Vanua Levu : Ndelaikoro, 500 m).

- DISTRIBUTION: Fiji.

tavakiliani Orousset, 1983b: 117 ( Edaphus ; Type locality: Mont Grand-Matoury, 7 km S. de Cayenne, alt. 200 m, Guyane française).

- DISTRIBUTION: French Guiana.

taylori Puthz, 1979: 120 ( Edaphus ; Type locality: Malaya: Upper Gombak Valley near

Kuala Lumpur, 1500 feet. Also cited from Singapore).

- DISTRIBUTION: Malaysia, Singapore.

termitophilus Bernhauer, 1916a: 2 ( Edaphus ; Type locality: Queensland: Blackal Range ).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

— Seevers, 1957: 319 ( Edaphus ; doubtfully termitophilous).

— Puthz, 1978c: 124 ( Edaphus ; characters; Australia).

- DISTRIBUTION: Australia.

camponoti Oke, 1933: 111 ( Edaphellus ; variety? or species? of melculus ; Type locality: Australia: Victoria: Gern Tree Gully, Evelyn, Lilydale).

— Puthz, 1978c: 124 ( Edaphus ; synonym of termitophilus ).

thai Orousset, 1988c: 184 ( Edaphus ; Type locality: Thaïlande: env. de Chiang Maï,

Doï Inthanon, alt. 2500 m).

- DISTRIBUTION: Thailand.

tongkingensis Puthz, 1979: 130 ( Edaphus ; Type locality: Vietnam: Luc yen).

- DISTRIBUTION: Vietnam.

topali Puthz, 1979: 133 ( Edaphus ; Type locality: India: W. Bengal, Darjeeling District , Ghum, 2200 m).

- DISTRIBUTION: India.

torresbolloi Puthz, 1985c: 355 ( Edaphus ; Type locality: Paraguay: alto Paraná, Pto Pte Stroessner ).

- DISTRIBUTION: Paraguay.

transvaalensis Puthz, 1989a: 71 ( Edaphus ; Type locality: South Africa: E. Transvaal: Mariepskop).

- DISTRIBUTION: South Africa.

tricornis Székessy, 1939: 102 ( Edaphus ; Type locality: Neu-Guinea: Friedrich-Wilhelms-Hafen).

- DISTRIBUTION: New Guinea.

tshuapaensis Kistner, 1962a: 631 ( Edaphus ; Type locality: Congo Republic: Tshuapa , Mabali, near Lake Tumba, 350 m).

— Kistner, 1963: 199 ( Edaphus ; Congo Republic).

— Puthz, 1974s: 762 ( Edaphus ; characters).

— Puthz, 1984g: 202 ( Edaphus ; Zaire).

- DISTRIBUTION: Congo.

tuberculicollis Puthz, 1990b: 280 ( Edaphus ; Type locality: Neu Guinea: Huon Golf: Sattelberg).

- DISTRIBUTION: New Guinea.

tuberculifer Puthz, 1990a: 197 ( Edaphus ; Type locality: Côte d’Ivoire: env. Bingerville ).

- DISTRIBUTION: Ivory Coast.

ulcilabris Puthz, 1992b: 28 ( Edaphus ; Type locality: Zaire: Kivu, Mwenga, 1250 m. Also cited from Rwanda).

- DISTRIBUTION: Congo, Rwanda.

unicornis Székessy, 1939: 105 ( Edaphus ; Type locality: Neu-Guinea: Marega).

- DISTRIBUTION: New Guinea.

uvaensis Comellini, 1977: 266 ( Edaphus ; Type locality: Sri Lanka: province Uva, Monaragala ). - DISTRIBUTION: Sri Lanka.

uvirensis Kistner, 1962a: 632 ( Edaphus ; Type locality: Congo Republic: Kivu, near Uvira , 800 m).

— Puthz, 1974s: 763 ( Edaphus ; characters).

- DISTRIBUTION: Congo.

variabilis Puthz, 1974 s: 761 ( Edaphus ; Type locality: Angola: Lunda: Alto Cuílo Cacola, Brüke über den Cuvembo).

— Puthz, 1984g: 201 ( Edaphus ; characters; Ghana; Tanzania; Uganda).

— Puthz, 1990a: 211 ( Edaphus ; Ivory Coast; Gabon; Cameroon).

- DISTRIBUTION: Angola, Gabon, Ghana, Ivory Coast, Cameroon, Tanzania, Uganda. discipunctus Puthz, 1974 s: 766 ( Edaphus ; Type locality: Uganda: zwischen Kampala und Jinja).

— Puthz, 1983f: 173 ( Edaphus ; Angola; Rhodesia; Kenya).

— Puthz, 1984g: 201 ( Edaphus ; synonym of variabilis ).

— Puthz, 1992b: 31 ( Edaphus ; synonym of variabilis ).

variatus Puthz, 1990a: 215 ( Edaphus ; Type locality: Côte d’Ivoire: Abidjan: Parc national du Banco).

- DISTRIBUTION: Ivory Coast.

variicollis Puthz, 1986c: 311 ( Edaphus ; Type locality: Rwanda: Cyangugu , Nyakabuye. Also cited from Zaire, Angola, and Cameroon).

- DISTRIBUTION: Angola, Cameroon, Congo, Rwanda.

varius Puthz, 1990a: 211 ( Edaphus ; Type locality: Côte d’Ivoire: Agboville: forêt de Yapo, près Yapo-Gare).

- DISTRIBUTION: Ivory Coast.

vedda Comellini, 1977: 259 ( Edaphus ; Type locality: Sri Lanka: province Central, Hasalaka près de Weragamtola).

— Puthz, 1979: 139 ( Edaphus ; characters).

- DISTRIBUTION: Sri Lanka.

ventriosus Puthz, 1989a: 73 ( Edaphus ; Type locality: South Africa: Anthrax Camp, Pafuri, Kruger National Park, 15 km W junction Limpopo River and Levuuhu River).

— Puthz, 1992b: 18 ( Edaphus ; characters).

- DISTRIBUTION: South Africa.

vietnamensis Puthz, 1979: 139 ( Edaphus ; Type locality: Vietnam: Cuc phuong, Ninh binh). - DISTRIBUTION: Vietnam.

vietus Puthz, 1974 r: 85 ( Edaphus ; Type locality: Vietnam: Provinz Quang Tri, 7 mi. W. of Dong Ha, Cam Lo ).

- DISTRIBUTION: South Vietnam.

vulcanicola Puthz, 1979b: 167 ( Edaphus ; Type locality: Sumatra: Sibajak vulkan).

— Puthz, 1990c: 823 ( Edaphus ; characters; Sumatra).

- DISTRIBUTION: Sumatra.

vulneratus Bernhauer, 1929c: 94 , 99 ( Edaphus ; Type locality: Leyte).

— Scheerpeltz, 1933: 1204 ( Edaphus ; catalog).

- DISTRIBUTION: Philippines.

werneri Puthz, 1990d: 33 ( Edaphus ; Type locality: Rwanda: Rangiro ).

— Puthz, 1992b: 19 ( Edaphus ; characters).

- DISTRIBUTION: Rwanda.

wernerianus Puthz, 1990d: 34 ( Edaphus ; Type locality: Rwanda: Rangiro ).

— Puthz, 1992b: 19 ( Edaphus ; characters).

- DISTRIBUTION: Rwanda.

wittmeri Puthz, 1979: 124 ( Edaphus ; Type locality: India: Assam, Megalaya, Garo Hills, Darugiri , 450 m).

xiao Puthz, 1983 g: 163 ( Edaphus ; Type locality: China: Guanxi [ Kwangsi ]: Guilin, Yaoshan).

- DISTRIBUTION: China. zukwalanus Puthz, 1990d: 39 ( Edaphus ; Type locality: Ethiopia: Shoa Province: L. Helene S.

Mt. Zughala).

- DISTRIBUTION: Ethiopia. zwicki Puthz, 1978c: 121 ( Edaphus ; Type locality: Victoria: Dandenong Range, Belgrave.

Also cited from Queensland).

- DISTRIBUTION: Australia. zwickianus Puthz, 1978c: 125 ( Edaphus ; Type locality: Victoria: Tolmie near Whitfield).

- DISTRIBUTION: Australia.

Euaesthetotyphlus

[1 species; Palaearctic Region]

Euaesthetotyphlus Coiffait and Decou, 1970: 377 (species included: almajensis ). TYPE SPECIES: Euaesthetotyphlus almajensis Coiffait and Decou , fixed by original designation.

— Tikhomirova, 1973a: 171 (checklist of species of USSR).

— Coiffait, 1984f: 366 (characters).

almajensis Coiffait and Decou, 1970: 379 ( Euaesthetotyphlus ; Type locality: Roumaine: Ogasul Porcariului, Vallée de la Nera).

— Coiffait, 1984f: 368 ( Euaesthetotyphlus ; characters).

- DISTRIBUTION: Romania.

Euaesthetus

[41 species; Neotropical, Nearctic, Palaearctic, and Oriental Regions] Euaesthetus Gravenhorst, 1806: 201 (species included: scaber ). TYPE SPECIES: Euaesthetus scaber Gravenhorst , fixed by monotypy.

— Gyllenhal, 1810: 461 (characters).

— Dejean, 1821: 25 (list of species in collection).

— Latreille, 1825: 245 (cited as Euaesthète; characters).

— Berthold, 1827: 331 (characters).

— Le Peletier and Serville, 1828: 483 (characters; notes).

— Zetterstedt, 1828: 91 (characters).

— Curtis, 1829: 30 (catalog; Britain).

— Stephens, 1829a: 24 (catalog; Britain).

— Stephens, 1829: 294 (catalog; Britain).

— Latreille, 1829: 437 (characters).

— C. Sahlberg, 1832: 420 (characters).

— Griffith and Pidgeon, 1832: 294 (characters).

— Dejean, 1833: 66 (list of species).

— Stephens, 1834: 328 (characters; habitat).

— Gistel, 1834: 9 (list of species).

— Westwood, 1838a: 18 (characters; type species).

— Zetterstedt, 1838: 72 (characters).

— Erichson, 1839a: 574 (characters).

— Heer, 1839: 228 (characters).

— Stephens, 1839: 407 (characters).

— Erichson, 1840: 746 (characters).

— Agassiz, 1847: 145 (cited as Evaesthetus ).

— Redtenbacher, 1849: 731 (characters).

— Schaum, 1852: 28 (list of species; Europe).

— Lacordaire, 1854: 109 (characters; notes; list of species).

— Fairmaire and Laboulbène, 1856: 597 (characters).

— Jacquelin du Val, 1857: 53 (characters).

— Kraatz, 1857d: 734 (characters).

— Redtenbacher, 1857: 226 (characters; Austrian species).

— Thomson, 1858: 39 (characters).

— G. Waterhouse, 1858: 27 (catalog of British species).

— Thomson, 1859: 42 (characters; type species: ruficapillus ).

— Schaum, 1859: 29 (catalog; European species).

— LeConte, 1861a: 67 (characters).

— Thomson, 1861: 115 (characters).

— Gredler, 1863: 112 (list of species from Tirol).

— LeConte, 1863: 25 (list of North American species).

— Wencker and Silbermann, 1866: 32 (list of species; collecting notes; France).

— Gemminger and Harold, 1868: 631 (catalog).

— Jacquelin du Val, 1868: 76 (catalog; European species).

— Sharp, 1871d: 12 (list of British species).

— Fauvel, 1873: 63 [= 1873a: 221] (characters; key to species).

— Redtenbacher, 1874: 236 (characters; key to Austrian species).

— Fauvel, 1875a: XIV [= 1875b: 216] (catalog).

— Mulsant and Rey, 1878a: 303 (characters; key to French species).

— Heyden, 1880: 79 (list of species of Siberian region).

— Duvivier, 1883: 178 (catalog).

— Casey, 1884a: 19 (key to North American species).

— Fowler, 1888: 324 (notes; key to British species).

— Seidlitz, 1891: 359 (key to some European species).

— Seidlitz, 1891a: 383 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 113 (list of species of Europe and Caucasus).

— Heyden, 1893: 46 (list of species of Siberian region).

— Ganglbauer, 1895: 544 (characters; key to middle European species).

— Heyden, 1896: 35 (list of species of Siberian region).

— Heyden, Reitter, and Weise, 1906: 151 (list of species of Europe and Caucasus).

— Reitter, 1909: 153 (characters).

— Blatchley, 1910: 458 (characters; key to Indiana species).

— Sainte-Claire Deville, 1910a: 127 (notes; key to and annotated list of species of Seine Basin, France).

— Bernhauer and Schubert, 1911: 188 (world catalog; 21 species).

— Petri, 1912: 63 (list of species of Siebenbürgen).

— Johansen, 1914: 475 (characters; key to Danish species).

— Leng, 1920: 100 (catalog of North American species).

— Winkler, 1925: 357 (catalog for Palaearctic region).

— Porta, 1926: 65 (characters; key to species of Italy).

— Bernhauer, 1928c: 38 (key to North American species).

— Portevin, 1929: 390 (characters; key to French species).

— Roubal, 1930: 348 (catalog; Slovakia).

— Joy, 1932: 14 (key to British species).

— Scheerpeltz, 1933: 1204 (world catalog supplement).

— Blackwelder, 1944: 116 (checklist of species from Mexico, Central and South America, and the West Indies).

— Kloet and Hincks, 1945: 164 (list of British species).

— Tottenham, 1949: 366, 413 (type species: bipunctatus Ljungh = scaber Gravenhorst ; notes; checklist of British species).

— Blackwelder, 1952: 152 (type species: scaber ).

— Tottenham, 1954: 74 (characters; collecting notes; key to British species).

— Hatch, 1957: 240 (characters; key to species of Pacific Northwest).

— Palm, 1961a: 117 (characters; key to species of Sweden).

— Horion, 1963: 402 (list of central European species).

— Lohse, 1964: 131 (characters; key to central European species). — Kasule, 1966: 277 (larval characters). — Scheerpeltz, 1968a: 46 (catalog of Austrian species). — Scheerpeltz, 1974: 106 (characters). — Tikhomirova, 1973a: 171 (checklist of species of USSR). — Moore and Legner, 1975: 262 (catalog; North America north of Mexico). — Shibata, 1976: 211 (checklist of species of Japan). — Pope, 1977: 26 (list of British species). — Topp, 1978: 307 (larval characters). — Moore and Legner, 1979: 244 (characters; notes). — Uhlig and Vogler, 1981: 102 (list of some species collected in Germany; habitat and collecting notes). — Coiffait, 1984f: 358 (characters; key to species of western Palaearctic region). — Ádám, 1985: 250 (list of species from southeast Hungary). — Frank, 1986a: 367 (checklist of species of Florida). — Muona and Viramo, 1986: 22 (list of species of northeastern Finland). — Segers, 1986: 34 (checklist of species; Belgium). — Ádám, 1987: 143 (list of four species with collecting notes; Hungary). — Lohse and Lucht, 1989: 149 (supplement for key to genera of Lohse, 1964). — Newton, 1990b: 1164, 1170 (larval characters). — Frank, 1991: 345 (larval characters). — Silfverberg, 1992: 20 (list of species of Fennoscandia, Denmark, and the Baltic States). — Siitonen, 1993: 230 (list of Finnish species). — Boháč, 1993: 45 (list of species of Czech Republic and Slovakia). — Puthz, 1994e: 392 (key to species of Old World). — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 28 (additional records to Danish list of species). — Terlutter, 1995: 55 (list of some species of Germany). — Zanetti, 1995: 19 (list of species of Italy). — M. Hansen, 1996: 102 (list of Danish species). — Downie and Arnett, 1996: 460 (characters; key to species of Northeastern North America). — Lundgren, 1998: 48 (list of species from Florida; records for other States listed for each species). — M. Hansen, Palm, Pedersen, and Runge, 1998: 71 (list of species collected in 1997; Denmark). — Newton, Thayer, Ashe, and Chandler, 2000: 382 (27 Nearctic species; notes; characters in key).

Eristhetus Leach, 1819: 174 (unjustified emendation; cited by Blackwelder, 1952: 152 as an emendation of Euaesthetus because it also had been treated as such by Mannerheim, 1830: 41). TYPE SPECIES: Euaesthetus scaber Gravenhorst , fixed by objective synonymy. — Mannerheim, 1831: 422, 455 (emendation of Euaesthetus ; [Note: Unjustified emendation]). — Lacordaire, 1835: 438 (characters). — Laporte, 1840: 184 (characters). — Lacordaire, 1854: 109 (synonym of Euaesthetus ). — Fauvel, 1873: 63 [= 1873a: 221] (synonym of Euaesthetus ). — Bernhauer and Schubert, 1911: 188 (synonym of Euaesthetus ; [Note: Mannerheim, 1830 cited as the author of genus]). — Coiffait, 1984f: 358 (synonym of Euaesthetus ).

albertae Puthz, 1998a: 229 ( Euaesthetus ; Type locality: Canada: Alberta: 2 mi. S. Ponaka, Battle River marsh). - DISTRIBUTION: Canada.

americanus Erichson, 1840: 747 ( Euaesthetus ; Type locality: Pensylvania). — LeConte, 1850: 220 ( Euaesthetus ; Lake Superior). — Casey, 1884a: 27 ( Euaesthetus ; characters; Michigan; Massachusetts; Washington, D.C.; Florida; New York; Delaware).

— Bernhauer and Schubert, 1911: 188 ( Euaesthetus ; catalog).

— Notman, 1919a: 95 ( Euaesthetus ; Ontario).

— Notman, 1920a: 180 ( Euaesthetus ; New York).

— Bernhauer, 1928c: 39 ( Euaesthetus ; characters in key).

— Scheerpeltz, 1933: 1205 ( Euaesthetus ; catalog).

— Frank, 1982: 16 ( Euaesthetus ; list of parasites).

— Campbell and Davies, 1991: 112 ( Euaesthetus ; USA; Canada).

— Downie and Arnett, 1996: 460 ( Euaesthetus ; characters; USA).

— Puthz, 1998a: 242 ( Euaesthetus ; lectotype designation; synonyms).

- DISTRIBUTION: Canada, USA.

subiridipennis Bernhauer, 1928c: 38 ( Euaesthetus ; Type locality: Massachusetts: Marion). — Scheerpeltz, 1933: 1206 ( Euaesthetus ; catalog).

— Downie and Arnett, 1996: 460 ( Euaesthetus ; cited as subviridipennis; [Note: Listed from Florida but perhaps erroneously]; USA).

— Puthz, 1998a: 242 ( Euaesthetus ; synonym of americanus ).

andersoni Puthz, 1998a: 233 ( Euaesthetus ; Type locality: Mexico: Guerrero, 10. 3 km SW Filo de Caballo , 2700 m).

- DISTRIBUTION: Mexico.

atomus Casey, 1884a: 28 ( Euaesthetus ; Type locality: Sand Point and Cedar Keys, Florida). — Bernhauer and Schubert, 1911: 189 ( Euaesthetus ; catalog).

— Bernhauer, 1928c: 40 ( Euaesthetus ; characters in key).

— Scheerpeltz, 1933: 1205 ( Euaesthetus ; catalog).

— Puthz, 1998a: 225 ( Euaesthetus ; character figured).

- DISTRIBUTION: USA.

bicoloratus Casey, 1924: 150 ( Euaesthetus ; Type locality: Florida: near Indian River).

— Scheerpeltz, 1933: 1205 ( Euaesthetus ; catalog).

- DISTRIBUTION: USA.

bierigi Puthz, 1998a: 225 ( Euaesthetus ; Type locality: Costa Rica: Vara Blanca , 2000 m).

- DISTRIBUTION: Costa Rica.

bipunctatus Ljungh, 1804: 68 ( Stenus ; Type locality: in glacie paludis Sjärsjöensis).

— Gyllenhal, 1810: 462 ( Euaesthetus ; synonym of scaber Gravenhorst ).

— C. Sahlberg, 1832: 420 ( Eristhetus ; synonym of scaber Gravenhorst ).

— Erichson, 1839a: 575 ( Euaesthetus ; synonym of scaber ).

— Erichson, 1840: 746 ( Euaesthetus ; synonym of scaber ).

— Kraatz, 1857d: 736 ( Euaesthetus ; synonym of scaber ).

— Fauvel, 1873: 64 [= 1873a: 222] ( Euaesthetus ; characters; France; Scandinavia; Britain; Germany; Austria; Transbaikal).

— Fauvel, 1873b: 129 [= 1873c: 22] ( Euaesthetus ; Russia).

— Mulsant and Rey, 1878a: 308 ( Euaesthetus ; characters; France).

— Ganglbauer, 1895: 545 ( Euaesthetus ; characters; north and middle Europe).

— Poppius, 1909: 20 ( Euaesthetus ; Lena Valley, Siberia ).

— Reitter, 1909: 153 ( Euaesthetus ; characters; Germany).

— Bernhauer and Schubert, 1911: 189 ( Euaesthetus ; catalog).

— Portevin, 1929: 391 ( Euaesthetus ; characters; France).

— Scheerpeltz, 1933: 1205 ( Euaesthetus ; catalog).

— Palm, 1961a: 119 ( Euaesthetus ; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 402 ( Euaesthetus ; France; Italy; Dalmatia; Romania; Germany; Austria).

— Lohse, 1964: 131 ( Euaesthetus ; characters; central Europe).

— Smetana, 1964d: 68 ( Euaesthetus ; Czechoslovakia).

— Szujecki, 1968a: 722 ( Euaesthetus ; Poland).

— Jarrige, 1968a: 141 ( Euaesthetus ; characters).

— Puthz, 1972q: 293 ( Euaesthetus ; characters).

— Ashworth, 1972: 215 ( Euaesthetus ; late-glacial subfossil; Britain).

— Outerelo, 1976: 265 ( Euaesthetus ; Spain).

— Pope, 1977: 26 ( Euaesthetus ; Britain).

— Coiffait, 1984f: 362 ( Euaesthetus ; characters).

— Lucht, 1987: 99 ( Euaesthetus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Lohse and Lucht, 1989: 149 ( Euaesthetus ; characters).

— Nowosad, 1990: 69 ( Euaesthetus ; collected from mole nest; notes; Poland).

— Sabella and Zanetti, 1991: 274 ( Edaphus ; notes).

— Boháč, 1993: 45 ( Euaesthetus ; Slovakia; Czech Republic).

— Puthz, 1994e: 393 ( Euaesthetus ; characters).

— Sparacio, 1995: 149 ( Euaesthetus ; characters; notes; Sicily).

— Zanetti, 1995: 19 ( Euaesthetus ; Italy).

— Ádám, 1996: 242 (Euaeathetus; Hungary).

- DISTRIBUTION: Europe, European and Asia Russia.

scaber Gravenhorst, 1806: 202 ( Euaesthetus ; Type locality: Brunsvigae).

— Gyllenhal, 1810: 462 ( Euaesthetus ; characters; Sweden).

— Germar, 1823a: 13 ( Euaesthetus ; characters; Germany; Sweden).

— Gyllenhal, 1827: 499 ( Staphylinus ; note).

— Zetterstedt, 1828: 92 ( Euaesthetus ; characters; Lapland).

— Latreille, 1829: 437 ( Euaesthetus ; France).

— Mannerheim, 1830: 41 ( Eristhetus ; Sweden).

— Mannerheim, 1831: 455 ( Eristhetus ; Sweden).

— C. Sahlberg, 1832: 420 ( Eristhetus ; characters; Finland).

— Stephens, 1834: 328 ( Euaesthetus ; characters; England).

— Lacordaire, 1835: 439 ( Eristhetus ; characters; France).

— Westwood, 1838a: 18 ( Euaesthetus ; type species).

— Zetterstedt, 1838: 72 ( Eristhetus ; characters; habitat; Lapland).

— Erichson, 1839a: 574 ( Euaesthetus ; characters; Germany).

— Heer, 1839: 228 ( Euaesthetus ; characters; Switzerland).

— Stephens, 1839: 415 ( Euaesthetus ; characters; England).

— Erichson, 1840: 746 ( Euaesthetus ; characters; Germany; Sweden).

— Laporte, 1840: 184 ( Eristhetus ; characters; France).

— Guérin-Méneville, 1844a: 31 ( Euaesthetus ; Europe).

— Redtenbacher, 1849: 731 ( Euaesthetus ; characters; Austria).

— Jacquelin Duval, 1850: 50 ( Euaesthetus ; characters; Germany; France).

— Fairmaire and Laboulbène, 1856: 597 ( Euaesthetus ; characters; France).

— Kraatz, 1857d: 736 ( Euaesthetus ; characters; Germany).

— Redtenbacher, 1857: 226 ( Euaesthetus ; characters; Austria).

— Thomson, 1861: 115 ( Euaesthetus ; characters; Scandinavia).

— Fauvel, 1873: 64 [= 1873a: 222] ( Euaesthetus ; synonym of bipunctatus ).

— Redtenbacher, 1874: 237 ( Euaesthetus ; characters; Austria).

— Seidlitz, 1875: 252 ( Euaesthetus ; characters; Baltic region).

— J. Sahlberg, 1876: 171 ( Euaesthetus ; Finland).

— Mulsant and Rey, 1878a: 306 ( Euaesthetus ; characters; France).

— Fowler, 1888: 324 ( Euaesthetus ; characters; Britain).

— Ganglbauer, 1895: 545 ( Euaesthetus ; synonym of bipunctatus ).

— Bernhauer and Schubert, 1911: 189 ( Euaesthetus ; synonym of bipunctatus ).

— Johansen, 1914: 476 ( Euaesthetus ; characters; Denmark).

— Scheerpeltz, 1933: 1205 ( Euaesthetus ; synonym of bipunctatus ).

— Pope, 1977: 26 ( Euaesthetus ; synonym of bipunctatus ).

— Coiffait, 1984f: 362 ( Euaesthetus ; synonym of bipunctatus ).

elongatus Motschulsky, 1860: 557 ( Euaesthetus ; Type locality: Baical; [Note: See lectotype designation by Puthz, 1973m]. Originally cited from environs du lac Baical en Daourie).

— Puthz, 1973m: 138 ( Euaesthetus ; lectotype designation: Baikal; synonym of bipunctatus ).

— Coiffait, 1984f: 362 ( Euaesthetus ; synonym of bipunctatus ).

mariae Bethe, 1867: 308 ( Euaesthetus ; Type locality: Dürkheim).

— Fauvel, 1873: 64 [= 1873a: 222] ( Euaesthetus ; synonym of bipunctatus ).

— Redtenbacher, 1874: 237 ( Euaesthetus ; characters).

— Ganglbauer, 1895: 545 ( Euaesthetus ; synonym of bipunctatus ).

— Bernhauer and Schubert, 1911: 189 ( Euaesthetus ; synonym of bipunctatus ).

— Scheerpeltz, 1933: 1205 ( Euaesthetus ; synonym of bipunctatus ).

— Coiffait, 1984f: 362 ( Euaesthetus ; synonym of bipunctatus ).

brevipennis Mulsant and Rey, 1878a: 308 ( Euaesthetus ; Type locality: Bresse).

— Ganglbauer, 1895: 545 ( Euaesthetus ; synonym of bipunctatus ).

— Bernhauer and Schubert, 1911: 189 ( Euaesthetus ; synonym of bipunctatus ).

— Coiffait, 1984f: 362 ( Euaesthetus ; synonym of bipunctatus ).

brevelytratus Outerelo and Gamarra, 1986: 592 ( Euaesthetus ; Type locality: Puerto de Honduras, Cáceres , entre Hervás y Cabezuela del Valle).

— Puthz, 1994e: 394 ( Euaesthetus ; characters).

- DISTRIBUTION: Spain.

brevipennis Mulsant and Rey, 1878 , see: bipunctatus Ljungh, 1804 .

brevipennis Casey, 1884 , see: duplex Herman, 2001 .

chandleri Puthz, 1998a: 234 ( Euaesthetus ; Type locality: USA: New Hampshire: Strafford Co., Spruce Hole , 3 mi SW Durham).

- DISTRIBUTION: USA.

chantali Puthz, 1998a: 236 ( Euaesthetus ; Type locality: USA: Massachusetts: Tyngsboro. Also cited from New Hampshire, Iowa, Mississippi, Texas, Ontario , Quebec; Cuba).

- DISTRIBUTION: Canada, USA, Cuba.

columbus Puthz, 1998a: 226 ( Euaesthetus ; Type locality: Cuba: Caimito del Guayabal ).

- DISTRIBUTION: Cuba.

cubanus Puthz, 1998a: 224 ( Euaesthetus ; Type locality: Cuba: Aspiro , Sierra del Rosario).

- DISTRIBUTION: Cuba.

dampfi Bernhauer, 1929 h: 190 ( Euaesthetus ; Type locality: Mexiko: Ajusco Berg , in einer Seehöhe von 4000 m).

— Scheerpeltz, 1933: 1205 ( Euaesthetus ; catalog).

- DISTRIBUTION: Mexico.

duplex Herman, 2001: 36 ( Euaesthetus ; replacement name for brevipennis Casey ).

- DISTRIBUTION: USA, Canada.

brevipennis Casey, 1884a: 20 ( Euaesthetus ; [preoccupied]; Type locality: Detroit , Michigan).

— Bernhauer and Schubert, 1911: 189 ( Euaesthetus ; catalog).

— Bernhauer, 1928c: 39 ( Euaesthetus ; characters in key).

— Campbell and Davies, 1991: 112 ( Euaesthetus ; Canada).

— Downie and Arnett, 1996: 460 ( Euaesthetus ; characters; USA).

— Herman, 2001: 36 ( Euaesthetus ; synonym of duplex ).

dybasi Puthz, 1998a: 230 ( Euaesthetus ; Type locality: USA: New York: Hamilton Co., Hope, N. Hope Falls Road. Also cited from New Hampshire, Illinois, Michigan, Wisconsin, Ontario and Quebec).

- DISTRIBUTION: Canada, USA.

elongatus Motschulsky, 1860 , see: bipunctatus Ljungh, 1804 .

fauveli Puthz, 1979b: 172 ( Euaesthetus ; Type locality: Tonkin).

— J. Li, 1993: 27 ( Euaesthetus ; China).

— Puthz, 1994e: 392 ( Euaesthetus ; characters).

floridae Casey, 1884a: 21 ( Euaesthetus ; Type locality: Florida, Tampa, Enterprise ).

— Blatchley, 1910: 458 ( Euaesthetus ; characters; Indiana).

— Bernhauer and Schubert, 1911: 189 ( Euaesthetus ; catalog).

— Bernhauer, 1928c: 39 ( Euaesthetus ; characters in key).

— Scheerpeltz, 1933: 1205 ( Euaesthetus ; catalog).

— Campbell and Davies, 1991: 112 ( Euaesthetus ; Canada).

— Downie and Arnett, 1996: 460 ( Euaesthetus ; characters; USA).

— Puthz, 1998a: 240 ( Euaesthetus ; illustration of aedeagus).

- DISTRIBUTION: USA, Canada.

franki Puthz, 1998a: 241 ( Euaesthetus ; Type locality: USA: Florida : Highlands Co., Fisheating Creek , 4 mi. W. of Venus).

- DISTRIBUTION: USA.

frigidus Poppius, 1909 , see: laeviusculus Mannerheim, 1844 .

fungicola Motschulsky, 1860 , see: Gyrophaena .

fulvus Motschulsky, 1860: 558 ( Euaesthetus ; Type locality: montagnes Oulou Tau dans les parties méridionales des Steppes des Kirguises).

— Fauvel, 1873: 65 [= 1873a: 223] ( Euaesthetus ; synonym of ruficapillus ).

— Bernhauer and Schubert, 1911: 189 ( Euaesthetus ; catalog).

— Scheerpeltz, 1933: 1205 ( Euaesthetus ; catalog).

— Coiffait, 1984f: 361 ( Euaesthetus ; characters; Caucasus; southern Russia; Kirghistan).

— Puthz, 1994e: 393 ( Euaesthetus ; characters).

- DISTRIBUTION: Caucasus, Kyrgyzstan.

ganglbaueri Bernhauer, 1928c: 39 ( Euaesthetus ; Type locality: Michigan).

— Scheerpeltz, 1933: 1205 ( Euaesthetus ; catalog).

— Campbell and Davies, 1991: 112 ( Euaesthetus ; Canada).

— Downie and Arnett, 1996: 460 ( Euaesthetus ; cited as ganglbauri; USA).

- DISTRIBUTION: USA, Canada.

granulipennis Eppelsheim, 1887: 429 ( Euaesthetus ; Type locality: Wladiwostok).

— Bernhauer and Schubert, 1911: 189 ( Euaesthetus ; catalog).

— L. Benick, 1924: 257 ( Euaesthetus ; Siberia).

— Puthz, 1973m: 139 ( Euaesthetus ; characters; notes; Siberia).

— Puthz, 1994e: 394 ( Euaesthetus ; characters).

- DISTRIBUTION: Russia.

hammondi Puthz, 1976d: 46 ( Euaesthetus ; Type locality: Japan: Yodogawa, Osaka. Also cited from China).

— J. Li, 1993: 27 ( Euaesthetus ; cited as hammionidi; China).

— Puthz, 1994e: 392 ( Euaesthetus ; characters).

- DISTRIBUTION: Japan, China.

hispanicus Coiffait, 1984b: 256 ( Euaesthetus ; Type locality: Espagne Picos de Europa , Lhoso, 1900 m).

— Outerelo and Gamarra, 1986: 598 ( Euaesthetus ; characters).

— Puthz, 1994e: 394 ( Euaesthetus ; characters).

- DISTRIBUTION: Spain.

immarginatus Erichson, 1840 , see: Stenaesthetus .

iripennis Casey, 1884a: 25 ( Euaesthetus ; Type locality: Ross County, Ohio).

— Bernhauer and Schubert, 1911: 189 ( Euaesthetus ; catalog).

— Bernhauer, 1928c: 38 ( Euaesthetus ; characters in key).

— Scheerpeltz, 1933: 1205 ( Euaesthetus ; catalog).

— Downie and Arnett, 1996: 460 ( Euaesthetus ; characters; USA).

- DISTRIBUTION: USA.

japonicus Bernhauer, 1907 , see: Edaphus .