Scaphisoma trilobum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1440-1441

5.2.21.9. Scaphisoma trilobum nov.sp. ( Figs 43, 44, 251-254 )

T y p e m a t e r i a l: Holotype 6: Philippines: Luzon env. Sagada Latan cave 15.-19.XII.79 Deharveng-Orousset (MNHN) [6 37, terre et herbe sèches à extérieur en bordure de cultures]. Paratypes: with the same data as the holotype, 1♀ (MHNG); Luzon, Mountain Prov., Sagada 15.- 19.XII.79 leg. L. Deharveng & J. Orousset, 1♀ (MHNG) [6 39, en aval du village, humus pied de parois calcaire]; Luzon, Mountain Prov., N Sagada nr. Ambasing 15.-19.XII.79, leg. L. Deharveng & J. Orousset, 2 66, ♀ f (MHNG) [6 41, litière d’arbustes feuillus dans une doline]; Luzon, Mountain Prov., N Sagada 15.-19.XII.79, leg. L. Deharveng & J. Orousset, 3♀♀ (MHNG) [6 51, mousses sur tronc de caféier. Doline sombre au nord de Sagada]; Luzon, Mountain Prov., Sagada, Latipan Cave, 15.XII.79, leg. L. Deharveng & J. Orousset, 1 6 (MHNG) [6 221, litière de feuillus près de l’entrée]; Luzon, Mountain Prov., doline N Sagada 15.-19.XII.79, leg. L. Deharveng & J. Orousset, 5 66, 10♀♀ (MHNG) [6 52, 53, litter]; Luzon, Mountain Prov., doline N Sagada 15.- 19.XII.79, leg. L. Deharveng & J. Orousset, 1 6 (MHNG) [6 71, litière très himude sous caféier]; 1 6, 3♀♀, same locality (MHNG) [6 72, litière humide]; 1 6, same locality (MHNG) [6 74, bois pourri contre paroi au fond de la doline]; Luzon, Mountain Prov., Sagada, 4.-5.I.80, leg. L. Deharveng & J. Orousset, 1♀ (MHNG) [6 143, vallon à l’est du cimetière, litière épaisse humide]; Luzon, Mountain Prov., Mt. Data, 8.I.1980, leg. J. Orousset 154, 1 6 (MHNG) [6 154, litière près de la lodge]; 1♀, same data but [6 164, mousses et feuilles au sol sur la crête]; Mt. Data, 2300 m, 7-9.VI.1977, leg. M. Satô, 1 6 (MHNG); Mt. Data, 2250 m, 13., 14. and 25.VII.1985, leg. M. Sakai, 17 66, 9♀♀ (EUMJ, MHNG); same but Ifugao Prov., 2355 m, 14.VII.1985, 1 6 (EUMJ); Luzon, Banguio, Mt. Santo Thomas, 2150 m, 14.I.1980, leg. L. Deharveng & J. Orousset, 1♀ (MHNG) [6 188, litière, surtout de bambous]; Luzon, Banguio, Mt. Santo Thomas, 8.VII.1970, leg. M. Satô, 3♀♀ (MHNG); same but 2250 m, 12.VI.1977, 1 6 (MHNG); Luzon, Mountain Prov., Mt. Puguis, 1900-2000 m, 18.VII.1985, leg. Y. Nishikawa, 2 66, 4♀♀ (EUMJ, MHNG); Luzon, Mountain Prov., Mt. Puguis, 2000 m, 5-6.VI.1977, leg. M. Satô, 1 6, 2♀♀ (MHNG); Luzon, Mountain Prov., Mt. Puguis, 1800-1900 m, 17.VII.1985, leg. M. Sakai, 3♀♀ (EUMJ); same data but 1900-2000 m, 18.VII.1985, leg. Y. Nishikawa, 2 6, 1♀ (EUMJ, MHNG); Luzon, Mountain Prov., Mt. Pangao, 2400 m, 30.V.1977, leg. M. Satô, 4 66, 1 ♀ (MHNG); same but 2350 m, 14.VII.1985, 1 6 (MHNG); Luzon, Benguet Prov., Sayangan, Paoay, 2400 m, 11.VII.1985, leg. M. Sakai, 4 6, 4♀♀ (EUMJ, MHNG, BZLA); Luzon, Ifugao Prov., Mt. Polis, 1900 m, 2.VI.1977, leg. M. Satô, 1♀ (MHNG); same data but 5.VI.1977, leg. Y. Kurosawa, 1 6 (EUMJ); Luzon, Benguet Prov., Mt. St. Tomas, 6.VII.1970, leg. M. Satô, 2♀♀ (EUMJ).

D e s c r i p t i o n: Length 1.90-2.80 mm, width 1.40-1.72 mm. Head, thorax, most of elytra and abdomen uniformly reddish-brown blackish, pronotum and hypomera often somewhat lighter than elytra, or basolateral areas of pronotum clearly lighter than remaining pronotal surface. Hypomera as pronotum or lighter, ochreous. Apical fourth to third of elytra lighter brown or yellowish, and well delimited. Appendages yellowish. Length/width ratio of antennomeres as: III 16/10: IV 48/7: V 70/8: VI 60/8: VII 64/15: VIII 52/8: IX 65/12: X 58/12: XI 75/15. Most other external characters as in S. tridens. Elytra with sutural striae weakly converging apically. Elytral punctation less dense and somewhat coarser, with most puncture interval about 1.5 to three times as large as puncture diameters. Metaventrite with two shallow, distinctly punctate apicomedian impressions. Intercoxal process of sternite 1 with several fairly distinct punctures, remaining abdominal punctation very fine, well visible at 100 times magnification, that along submetacoxal lines excepted.

M a l e: Mesotarsomere 1 about as wide as apex of mesotibiae. Sternite 6 flat. Aedeagus ( Figs 251-254 ) 1.20-1.54 mm long. Apical process of median lobe with ventral branch long, obliquely inflexed, tapering apically, truncate at tip. Dorsal branch of median lobe fairly long, triangular and reaching beyond mid-length of ventral branch in dorsal view. Internal sac with two hook-like proximal sclerites, two long, curved admesal teeth, mesal elongate vesicle bearing membranous denticular structure followed by dense spine-like structures.

F e m a l e: Apical margins of elytra oblique, uneven, notched near inner angles, inner angles prominent, tooth-like, each about 0.03 to 0.05 mm long.

E t y m o l o g y: The species epithet is Latin and means three-lobed.

C o m p a r a t i v e n o t e s: This species is unusually variable in its body-size and colour. The females may be easily distinguished from other Philippine congeners by the shape of the apical margins of the elytra. In S. dentipenne the inner apical angles of elytra are also denticulate but not prominent, and the margins laterad notches are crenulate. The males of S. trilobum may be readily distinguished from the similar S. tridens by the shape of the ventral branch of the apical lobe that is curved and much shorter in the latter species.

Figures

Figs 251 - 255: Aedeagi of Scaphisoma; (251, 252), S. trilobum nov. sp., in dorsal and lateral views, scale = 0.3 mm; (253, 254), ditto, internal sacs, in dorsal and lateral views, scale = 0.15 mm; (255), S. trimaculatum nov. sp., aedeagus in dorsal view, scale = 0.3 mm.