Scaphisoma jankodadai, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1426-1427

5.2.19.12. Scaphisoma jankodadai nov.sp. ( Figs 212, 213 )

Holotype 6: Philippines, Palawan centr. above San Rafael, ca 300 m, 4.XII.1995, J. Kodada leg. (MHNG).

Description: Length 1.58 mm, width 1.06 mm. Head, body and femora ochreous, apical abdominal segments, tibiae, tarsi and antennae lighter, yellowish. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 13/8: IV 34/7: V 44/7: VI 41/7: VII 47/14: VIII 37/8: IX 45/12: X 41/12: XI 58/14. Pronotum very finely and fairly densely punctate; lateral contours evenly rounded, lateral margin carinae hardly visible in dorsal view, lateral striae appearing impunctate. Exposed tip of scutellum small. Elytra with lateral margin carinae visible in dorsal view, apical margins truncate, inner apical angles not prominent, situated posterior level of outer angles; sutural margin not raised, sutural striae fairly deep, hardly curved at base, very weakly converging apically, punctate; adsutural areas narrow, flat, densely and very finely punctate, lateral striae appearing impunctate; discal punctation coarser than pronotal punctation, dense and fine, with puncture intervals slightly larger to four times as large as puncture diameters. Hind wings fully developed. Hypomera and mesanepisterna not microsculptured, smooth. Mesepimera about 1.5 times as long as intervals to mesocoxae, about four times as long as wide. Metaventrite in middle part hardly convex, with strigulate microsculpture and very finely, sparsely punctate, with single shallow mesal impression; microsculpture present between mesocoxae and metacoxae, lateral parts of metaventrite not microsculptured, sparsely and very finely punctate; antecoxal puncture rows absent. Submesocoxal areas 0.03 mm, about as fifth of intervals to metacoxae, submesocoxal lines parallel, distinctly punctate. Metanepisternum flat, narrowed anteriad, inner margin straight in middle part, rounded near angles, impressed below margin of metaventrite. Tibiae straight. Abdomen with strigulate microsculpture, very finely punctate, submetacoxal areas 0.05 mm, almost as fourth of intervals to apical margin of sternite 1, submetacoxal lines convex, distinctly punctate.

M a l e: Protarsomeres 1 to 3 and mesotarsomeres 1 and 2 distinctly widened. Aedeagus ( Figs 212, 213 ) 0.90 mm long, symmetrical, fairly sclerotized. Median lobe with dorsal branches of apical process short, ventral branch shorter than basal bulb, weakly inflexed, tapering (dorsal view), with somewhat sinuate ventral side and two subapical denticles on dorsal side (lateral view). Articular process robust, not prominent. Parameres sinuate and widened posterior mid-length in lateral view, almost evenly wide and strongly bent in dorsal view. Internal sac complex, with very dense, elongate bunch of proximal denticles, followed by two robust admesal teeth joint in middle, pair of hornlike teeth, several small sclerites in-between, and several elongate subapical spines; membranes posterior robust teeth with scale-like structures.

E t y m o l o g y: The species is dedicated to a friend of the senior author, Jan Kodada, Bratislava, Slovakia, who collected this species.

C o m p a r a t i v e n o t e s: The aedeagal characters suggest close relationships of S. jankodadai with S. luzonicum PIC. The internal sacs differ conspicuously, in particular by the proximal bunch of denticles and the shape of the large admesal teeth present in S. jankodadai. The new species differs from S. luzonicum in external characters by the uniformly ochreous body and the parallel submesocoxal lines.

Figures

Figs 212 - 218: Aedeagi of Scaphisoma; (212, 213); S. jankodadai nov. sp., in dorsal and lateral views, scale 0.2 mm; (214), S. luzonicum PIC, in dorsal view, scale = 0.2 mm; (215), ditto, internal sac, scale = 0.1 mm; (216, 217), S. bilobum nov. sp., in dorsal and lateral views, scale 0.2 mm; (218), ditto, internal sac, scale = 0.1 mm.