Scaphosoma tricolor HELLER, 1917

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1434-1435

5.2.21.3. Scaphosoma tricolor HELLER, 1917 (Figs 38, 39, 234-237)

Scaphosoma tricolor HELLER, 1917 : 46. Lectotype 6, SMTD; type locality: Luzon, Mt. Makiling. Lectotype designation: LÖBL 1970a: 126; apical margin of elytra: LÖBL 1971a: 249. Additional published records. Palawan: Pinigisan; Balabac: Dalawan Bay ( LÖBL 1971a: 250)

. Scaphosoma latum PIC, 1920b: 3 . Lectotype 6, MNHN; type locality: Luzon, Los Banos. Lectotype designation and synonymy: LÖBL 1970a: 126.

A d d i t i o n a l m a t e r i a l e x a m i n e d: Luzon, Lagunas Prov., Mt. Makiling, 4 0 0- 600 m, summit road, 19., 21-22., 26. and 28.XI.1995, leg. I. Löbl, 59 ex. (MHNG, BZLA); same but above Mad Springs, 400-700 m, 19-22.XI.1 995, leg. J. Kodada 14 ex. (MHNG); Luzon, Lagunas Prov., Los Banos, small river, 28.XI.1 995, J. Kodada & B. Rigová, 1 ex. (MHNG); Luzon, Lagunas Prov., Mt. Banahaw above Kinabuhayan, 600-700 m, trail to Crystalino, 24- 25.XI.1995, leg. J. Kodada & B. Rigová, 5 ex. (MHNG); Luzon, Laguna Prov., Mt. Makiling 4 km SE Los Banos, 4-12.V.197 7, leg. L.E. Watrous, 5 ex. (MHNG); Luzon, Los Banos, 1 ex. (SMTS); Luzon, Ifugao Prov., Mt. Polis, 1900 m, 4-5.VI.1977, leg. M. Satô, 1 ex. (MHNG); Palawan: Central, Cabayugan nr. Lion’s Cave, sea level, 1.XII.1995 I. Löbl, 1 ex. (MHNG); same but leg. J. Kodada, 2 ex. (MHNG); Palawan, Sabang, trail to Underground River, sea level, 30.XI.1995, leg. I. Löbl, 4 ex. (MHNG); Palawan, Sabang, 50-100 m, degraded forest on slope, leg. J. Kodada, 8 ex. (MHNG); Palawan, Sabang, N. of Mt. St. Paul, 11-13.VII.1977, leg. M. Satô, 3 ex. (MHNG); Palawan, Conception, sea level, 6-7.XII.1995, leg. I. Löbl, 6b ex. (MHNG); Palawan Central, Conception, large logs across Conception River, NE San Rafael, ca 20 m, 8.XII.1995, leg. J. Kodada & B. Rigová, 4 ex. (MHNG); Palawan, Olangoan, 18 km NE San Rafael, sea level, 5- 6.XII.1995, leg. I. Löbl, 6 ex. (MHNG); same but 1.IX.1985, leg. M. Sakai, 5 ex. (EUMJ); Palawan, Binaluan, 1 ex. (SMTD); Palawan, Matalangao, 150 m, nr. Roxas, 29.VIII.1985, leg. M. Sakai, 1 ex. (EUMJ); Palawan, Roxas, region of Matalangao, 80 m, 5.IV.1983, leg. S. Nagai & C. Lienhard, 1 ex. (MHNG); Palawan, Mantalingajan Range, S. slope Mt. Balabag, 2800 ft., 4- 17.V.47, leg. F.G. Werner, 3 ex. (FMNH); Palawan, Trident Mina , foot of Victoria Peak, 100 m, nr. Narra, 5.IX.1985, leg. M. Sakai, 2 ex. (EUMJ, MHNG); Leyte, Visca N Baybay, primary forest, 200-500 m, 22.II., 27.II. and 8.III.1991, leg. W. Schawaller et al., 6 ex. (SMNS, MHNG); same but secondary forest, 100-200 m, 27.II. and 1.III.1991, 2 ex. (SMNS, MHNG); Leyte, Tarragona, 30.VI.1945, leg. C.L. Remington, 1 ex. (FMNH); Masbate, Aroroy, 14.VIII.1918, leg. G. Böttcher, 2 ex. (ZMUB); Mindoro, 28 km S Calapan, Balete, 27-29.XI.2002, leg. H. Schillhammer, 1 ex. (NHMW); Bucas, Sorocco, X.1916, leg. G. Böttcher, 1 ex. (ZMUB); Mindanao, Surigao, 30-X.1915, leg. G. Böttcher, 3 ex. (ZMUB); Mindanao, Cotabato Prov., 50 km N. of Parang, 500 ft., 6.XII.1946, leg. F.G. Werner, 19 ex. (FMNH, MHNG); Mindanao, Cotabato Prov., Burungkot, Upi, 1500 ft., 6.I.1947, leg. F.G. Werner, 2 ex. (FMNH); Mindanao, Davao Prov., Sitio Taglawig, Maco, Tagum, nr. sea level, X.1946, leg. H. Hoogstraal & D. Heyneman, 5 ex. (FMNH, MHNG); Mindanao, 25 km NW of Zamboanga, 800 m, Camp Susana, 28-30.IV.1996, leg. Bolm, 3 ex. (SMNS, MHNG); Mindanao, Bukidnon Prov., 5 km N of Malaybalay, 900 m, 12.V.1996, leg. Bolm, 1 ex (SMNS); Mindanao, 30 km E of Malaybalay, Busdi, 1000 m, 5-9.V.1996, leg. Bolm, 18 ex. (SMNS, MHNG); Mindanao, 15 km NW of Bislig, 6-7.VII.1977, leg. M. Satô, 4 ex. (MHNG); Mindanao, Todaya, 29.VII.1970, leg. M. Satô, 2 ex. (MHNG); Mindanao, Baracatan, 1500 m, 27-29.VI.1977, leg. M. Satô, 10 ex. (MHNG).

D i s t r i b u t i o n: Philippines: Luzon, Palawan, Balabac, Leyte, Masbate, Mindoro, Dinagat, Bucas, Mindanao.

C o m p a r a t i v e n o t e s: The almost entirely ochreous male pronotum with very narrowly darkened along basal margin is diagnostic for this species. The elytral colora- tion is somewhat more variable, and specimens from Palawan have the apical light area narrower than that exhibited by Luzon specimens. The aedeagal characters are for the first time illustrated in the present paper ( Figs 234-237 ). Though the number, position and shape of the sclerites of the internal sac are diagnostic, they may appear highly vari- able if internal sac not completely in repos. The shape of the long and robust ventral branch of the apical process of the median lobe and the internal sac with a small tuber- culate lobe are also diagnostic. The inner apical angle of female elytra is weakly crenu- lated, not dentate.

Figures

Figs 234 - 237: Aedeagus of Scaphisoma tricolor HELLER; (234), in dorsal view; (235), in lateral view, without proximal part of basal bulb, scale = 0.2 mm; (236, 237), internal sac in dorsal and lateral views, scale = 0.1 mm.