Scaphisoma inexspectatum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1408-1409

5.2.18.3. Scaphisoma inexspectatum nov.sp. ( Figs 15 , 162, 163 )

Type specimens: Holotype 6: Burungkot, Upi, Cotabato Prov. Mindanao Elev. 1500 ft., I,1-9, 1947 / Lot 154 taken on polypores / CNHM-Philippine Zool. Exped. (1946-47) F.G.Werner leg. (FMNH). Paratypes: Mindanao, with the same data as the holotype, 11 66, 5♀♀, 4 ex. sex not examined (FMNH, MHNG, BZLA); Mindanao, Davao Prov., E. slope of Mt. McKinley, 3200 ft., 7-8.IX.1946, lot 54 beating, leg. F.G. Werner, 3 66, 2♀♀ (FMNH, MHNG); Mindanao, Davao Prov., 25 km NW of New Bataan, 1200 m, 20-22.V.1996, leg. Bolm 2 66, 4 ♀♀ (SMNS); Mindanao, Misamis occ., Don Victoriano, 1700 m, 1-3.V.1996, leg. Bolm, 8 66, 6♀♀ (SMNS, MHNG).

D e s c r i p t i o n: Length 1.10-1.23 mm, width 0.75-0.83 mm. Head and most of body dark reddish-brown to blackish, elytra near apical margins as in middle or darkened, elytral apices light brown, apical abdominal segments and appendices ochreous to yellowish. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 13/5: IV 24/5: V 46/5: VI 34/5: VII 50/8: VIII 40/6: IX 52/10: X 53/9: XI 60/10. Pronotum very finely and sparsely punctate with or without coarser and dense punctures on basomedian area, with lateral contours weakly rounded, lateral margin carinae hardly visible in dorsal view, lateral striae distinctly punctate. Exposed tip of scutellum minute. Elytra with lateral margin carinae entirely visible in dorsal view, apical margins truncate, inner apical angles not prominent, situated in level with outer angles; sutural margin not raised, sutural striae deep, starting somewhat posterior level of scutellum, parallel toward apical fourth of sutural length, converging toward apices, coarsely punctate; adsutural areas narrow, flat, densely and distinctly punctate, lateral and epipleural striae distinctly punctate. Elytral disc with punctation dense and coarse from bases to apical third, puncture rows fairly distinct, puncture intervals mostly about as large to twice as large as puncture diameters; apical fourth fairly finely and sparsely punctate. Hind wings fully developed. Hypomera and mesanepisterna smooth. Mesepimera about twice as long as intervals to mesocoxae, about five times as long as wide. Median part of metaventrite hardly convex, with shallow apicomesal impressions, lacking distinct microsculpture; with few fairly coarse and dense punctures in apicomedian impressions, punctation very fine and sparse and very fine on remaining surface and on sides; antecoxal puncture rows present, in impressed lines. Submesocoxal areas 0.03 mm, about as fifth of intervals to metacoxae, submesocoxal lines convex, coarsely punctate. Metanepisternum convex, narrowed anteriad, inner margin rounded, impressed below margin of metaventrite. Tibiae straight. Abdomen with strigulate microsculpture, very finely punctate; sternite 1 with submetacoxal areas 0.05 mm, about as half of intervals to apical margin of sternite, submetacoxal lines convex, coarsely punctate.

M a l e: Protarsomeres 1 to 3 weakly widened. Aedeagus ( Figs 162, 163 ) 0.40-0.42 mm long, symmetrical, weakly sclerotized. Apical process of median lobe narrow, not inflexed, with ventral side almost straight and in axis with basal bulb, abruptly inflexed and narrowed toward acute tip. Articular process robust, not prominent. Parameres narrow, gradually narrowed toward mid-length, curved in apical halves, each with narrow membranous lobe. Internal sac with lacking sclerotized sclerotized rods, with very fine denticulate structures.

E t y m o l o g y: The species epithet is a Latin adjective meaning unexpected.

C o m p a r a t i v e n o t e s: This species is very similar to S. surigaosum (PIC) and to S. furcigerum described below. It differs from both by its pronotal punctation and, more distinctly, by the aedeagus, in particular by the shape of the tip of the median lobe.

Figures

Figs 14 - 21: Habitus of Scaphisoma; (14), S. furcillatum nov. sp.; (15), S. inopportunum nov. sp.; (16), S. lunabianum nov. sp.; (17), S. inexspectatum nov. sp.; (18), S. luteomaculatum PIC; (19), S. luteopygidiale (PIC); (20), S. maculosum nov. sp.; (21), S. mindanaosum PIC. Scale: 1.0 mm.

Figs 162 - 168: Aedeagi of Scaphisoma; (162, 163), S. inexspectatum nov. sp., in dorsal and lateral views, scale = 0.1 mm; (164, 165), S. inflexum nov. sp., in dorsal and lateral views, scale = 0.1 mm; (166), ditto, parameres in ventral view, scale = 0.1 mm; (167), S. luteopygidiale (PIC), in lateral view, scale = 0.1 mm; (168), S. ochropenne nov. sp., in dorsal view, scale = 0.2 mm.