Scaphisoma inopportunum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1392-1393

5.2.13.7. Scaphisoma inopportunum nov.sp. ( Figs 16 , 127, 128 )

T y p e m a t e r i a l: Holotype 6: Luzon: Lagunas Mt. Banahaw, ca 1 km from Kinabuhayan, 500m, 25.XI.1995, I.Löbl, fungi on logs, under bark, degr. rainforest (MHNG). Paratypes: with the same data as the holotype, 4 66, 5 ♀♀ (MHNG, BZLA); same but above Kinabuhayan, 600-700 m, 24.XI.1995, 1 6, 2♀♀ (MHNG); Luzon, Lagunas, Mt. Makiling, 4 km S Los Banos, 7.V.1977, leg. L.E. Watrous, 3 ex., and with the same data but 11.IV.1977, 1 6, 1♀ (MHNG); Luzon, Los Banos / 1924, 1 6 (SMTD); Luzon, Lagunas Mt. Makiling, summit road 600 m, 21-22.XI.1995, leg. I. Löbl, 1♀ (MHNG); Luzon, Lagunas Mt. Makiling, 430 m, 17- 18.VI.1977, leg. M. Satô, 3♀♀ (EUMJ).

Description: Length 1.45-1.70 mm, width 1.05-1.23 mm. Head and body reddish-brown to blackish, mesoventrite and metaventrite often darkened, femora and tibiae reddish-brown, lighter than body, tarsi and antennae yellowish. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 12/8: IV 16/6: V 23/8: VI 38/9: VII 48/15: VIII 32/10: IX 45/14: X 39/14: XI 48/16. Pronotum with punctation very fine, lateral margins evenly rounded, lateral margin carinae not visible in dorsal view, lateral striae very finely punctate. Exposed tip of scutellum minute. Elytra convex, with lateral margin carinae usually exposed in dorsal view, apical margins truncate, inner apical angles not prominent, situated posterior level of outer angles; sutural margin not raised, sutural striae fairly deep, shortly bent at base, not extended along basal margins, parallel posteriad, very finely punctate; adsutural areas narrow, flat, each with single puncture row, lateral striae impunctate. Elytral disc with punctation fine, well visible, punctures usually somewhat larger than those on pronotum. Hind wings fully developed. Hypomera and mesanepisterna smooth. Mesepimera shorter than intervals to mesocoxae, about three times as long as wide. Metaventrite not microsculptured, very finely and sparsely punctate. Median part of metaventrite convex, without stria or impressions; antecoxal puncture rows absent. Submesocoxal areas 0.04 mm, about as thirds of intervals to metacoxae, submesocoxal lines convex, coarsely punctate, punctures not extended by striae. Metanepisternum hardly convex, slightly narrowed anteriad, inner margin rounded at angles, almost straight in-between, impressed, suture deep. Tibiae straight. Abdomen with sternite 1 lacking microsculpture, very finely punctate, submetacoxal areas 0.02 mm, about as eighth of intervals to apical margin of sternite, submetacoxal lines parallel, coarsely punctate. Following sternites with punctulate microsculpture.

M a l e: Protarsomeres and mesotarsomeres 1 to 3 slightly widened. Aedeagus ( Figs 127, 128 ) 0.32-0.37 mm long, asymmetrical, weakly sclerotized. Basal bulb oval, with inconspicuous articular process. Apical process inflexed, arcuate, gradually narrowed, with acute tip. Parameres strongly asymmetrical. Left parameres wide in basal part, abruptly in middle and irregularly curved, very narrow in apical section. Right paramere shorter than left parameres, evenly wide, bent and with truncate apex in dorsal view, irregularly curved and widened posterior mid-length in lateral view. Internal sac simple, with thin flagellum.

E t y m o l o g y: The species epithet is a Latin adjective meaning inopportune. Comparative notes. This species shares most diagnostic characters with S. palawanum and S. oviforme described below. It may be distinguished by its aedeagal features, in particular by the shape of the parameres.

Figures

Figs 14 - 21: Habitus of Scaphisoma; (14), S. furcillatum nov. sp.; (15), S. inopportunum nov. sp.; (16), S. lunabianum nov. sp.; (17), S. inexspectatum nov. sp.; (18), S. luteomaculatum PIC; (19), S. luteopygidiale (PIC); (20), S. maculosum nov. sp.; (21), S. mindanaosum PIC. Scale: 1.0 mm.

Figs 126 - 130: Aedeagi of Scaphisoma; (126), S. inconventum nov. sp., in lateral view; (127, 128), S. inopportunum nov. sp., in dorsal and lateral views; (129, 130) S. oviforme nov. sp., in dorsal and lateral views. Scale = 0.1 mm.