Scaphisoma cuyunon, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1400-1401

5.2.14.5. Scaphisoma cuyunon nov.sp. (Figs 9, 146, 147)

T y p e m a t e r i a l: Holotype 6: Palawan (central) Conception, sea level I. Löbl, 6- 7.XII.1995, fungi on logs, bark (MHNG). Paratypes: Palawan, Central, with the same data as the holotype, 3 66, 4 ♀♀, 10 ex. sex not examined (MHNG, BZLA); Palawan Central, Conception, large logs across Conception River, NE San Rafael, ca 20 m 8.XII.1995, leg. J. Kodada & B. Rigová, 9 ex., sex not examined (MHNG); Palawan Central, nr. Cabayugan, degraded rainforest, 150 m, 2.XII.1995, leg. J. Kodada & B. Rigová, 10 ex., sex not examined (MHNG); Olangoan, sea level, 18 km N San Rafael, 5-6-XII.1995, leg. I. Löbl, 1 6, 8♀♀, 9 ex. (MHNG); Palawan, Olanguan [sic], 0-50 m, between Puerto Princesa and Roxas, 29.VIII.1985, leg. M. Sakai, 1 6, 1♀, 3 ex. sex not examined (EUMJ, MHNG); with the same data but 1.IX.1985, 1♀ (EUMJ); Palawan Central, along Tarabanan River, NE San Rafael, ca 30 m, 7.XII.1995, leg. J. Kodada & B. Rigová, 1 ex., (MHNG); Palawan Central, Sabang, 50-100 m, degraded rainforest on slope, 30.XI.1995, leg. J. Kodada, 1 6 (MHNG); Palawan Central, Sabang, trail to Underground River, sea level, 30.XI.1995, fungi on logs, leg. I. Löbl, 1 6, 3♀♀ (MHNG); Palawan, Binaluan, 3 ex. (SMTD); Palawan, Matalangao, 500 m, 25 km W from Roxas, 21.XI.1978, leg. S. Nagai, 1 ex. (EUMJ).

Description: Length 1.15-1.30 mm, width 0.80-0.90 mm. Head, thorax and most of elytra reddish-brown to blackish-brown. Apical fourth to fifth of elytra light brown to yellowish. Most of abdomen reddish-brown, lighter than thorax. Abdominal apex and appendages yellowish. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 15/7: IV 30/6: V 41/6: VI 35/6: VII 50/13: VIII 40/8: IX 47/11: X 47/11: XI 67/12. Pronotum finely and densely punctate, with lateral margins evenly rounded, lateral margin carinae not or hardly visible in dorsal view, lateral striae finely punctate. Exposed tip of scutellum minute. Elytra with lateral margin carinae entirely and well visible in dorsal view, apical margins weakly rounded, inner apical angles not prominent, situated about in level with outer angles; sutural margin raised, sutural striae deep, bent at base, not extending along basal margin, strongly converging toward apices, coarsely punctate in anterior halves or two thirds, impunctate in apical thirds; adsutural areas conspicuously broad anteriad, about twice as wide at level of scutellar tip as at mid-length, flat, densely and coarsely punctate anterior mid-length or in anterior two thirds, with punctures usually larger than those on elytral disc, appearing impunctate near apices, lateral striae conspicuously coarsely punctate. Elytral disc with punctation almost evanescent. Hind wings fully developed. Hypomera and mesanepisterna smooth. Mesepimera longer than intervals to mesocoxae, about five to six times as long as wide. Median part of metaventrite slightly convex, with two apicomesal impressions, strigulate microsculpture present posterior mesocoxae, punctation on prevailing surface of metaventrite extremely fine and sparse; antecoxal puncture rows absent. Submesocoxal areas 0.04-0.05 mm, about as fifth to fourth of intervals to metacoxae, submesocoxal lines convex, coarsely punctate, punctures extended by striae on section between mesocoxae. Metanepisternum somewhat convex, narrowed anteriad, inner margin strongly rounded posteriad, strongly narrowed anteriad, suture conspicuously deep. Tibiae straight. Abdomen with strigulate microsculpture, fairly coarsely and densely punctate on small basomedian area, very finely and sparsely punctate on remaining surface; submetacoxal areas 0.04-0.05 mm, about as fourth to third of intervals to apical margin of sternite 1, submetacoxal lines convex, coarsely punctate.

M a l e: Protarsomeres 1 to 3 weakly widened. Aedeagus ( Figs 146, 147 ) 0.50-0.54 mm long, symmetrical, with basal bulb weakly sclerotized. Articular processes small, not prominent. Apical process of median lobe comparatively short, inflexed, bifid. Ventral branch of apical process robust, gradually narrowed, strongly sclerotized, covered by weakly sclerotized, broadly rounded dorsal lamina. Parameres in lateral view sinuate, almost evenly wide posterior bases up to apical tenth, in dorsal view almost evenly arcuate, with narrowly membranous inner margin in basal halves, evenly narrow in apical third. Internal sac complex, with large vesicle consisting of spines and scale-like structures, narrowed apically, in apical part narrowed and bearing longer spines.

E t y m o l o g y: The species epithet is a name of one of the Luzon’s languages.

Comparative notes: The species resembles the sympatric S. disparoides, though it has much longer antennae. It differs notably by the coarsely punctate lateral striae and adsutural areas of the elytra, the shorter and coarsely punctate submesocoxal lines, and the shape of the aedeagus. The aedeagal characters suggest relationships to S. lannaense LÖBL, 1990 from Thailand with which it shares parameres lacking lobes and the internal sac lacking sclerotized pieces.

Figures

Figs 146 - 150: Aedeagi of Scaphisoma; (146, 147), Scaphisoma cuyunon nov. sp., in dorsal and lateral views, scale = 0.1 mm; (148, 149), S. hexameroides nov. sp., in dorsal and lateral views, scale = 0.2 mm; (150), S. iridescens nov. sp., in lateral view, scale = 0.2 mm.