Rhizomyia parkalovi Fedotova et Perkovsky, 2014

Fedotova, Z. A. & Perkovsky, E. E., 2014, New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber, Vestnik Zoologii 48 (1), pp. 35-50 : 44-45

Rhizomyia parkalovi Fedotova et Perkovsky , sp. n.

M a t e r i a l. Holotype. SIZK N K–7385 , well preserved inclusion of female with tarsi missing, Klesov , Rovno amber, Late Eocene ( fig. 5, 1–8, 7, 6, 7 ). Syninclusions: N K–7383: Collembola Entomobryomorpha, 2 ex.; Cecidomyiidae : Rhizomyia sp. , K–7384: Sciaridae .

D e s c r i p t i o n. F e m a l e ( fig. 5, 1–8, 7, 6, 7 ). Body (without genitalia) is 1.8 times longer than antennae. Wings 2.2 times as long as antennae. Length of head 2.2 times as long as lateral side of head. Antennae are 2+10-segmented. Head completely covered by eyes laterally, eye bridge narrow. Scape enlarged distally, almost as long as round pedicel. Necks of flagellomeres very short. First flagellomere slightly longer than 2nd; 5th flagellomere 2.0 times as long as wide. Ninth flagellomere without neck; 10th flagellomere 2.9 times as long as wide; narrowed distally. Palpi are very narrow, 2-segmented, 2nd segment is 1.8 times longer than the first. Fore and middle femur have equal length, shorter than tibia ones. Lengths of fore and middle tibia also almost equal. First tarsomere with lateral long projection is covered by setae. Wing is 2.3 times as long as wide. Vein R 4+5 strongly curved near apex, joining wing margin at smaller distance from tip of wing than Cu 2. All tibiae subequal to, or longer than, respective femora. Fore tibia 1.1 times as long as femur. Ovipositor is not protrusible. 8th abdominal segment without ventral lobes or projections.

M e a s u r e m e n t s, mm: body length, 1.05; wing length, 1.27; wing width, 0.65; antennal length, 0.59; head length, 0.24 (without mouth parts); width of lateral side of head, 0.11; thorax length, 0.34; abdomen length, 0.61; length of fore: coxa, 0.10; trochanter, 0.07; femur 0.45; tibia 0.48; 1st segment, 0.07; length of middle: coxa, 0.11; trochanter, 0.07; femur 0.43; tibia 0.45; length of hind: coxa, 0.14; trochanter, 0.06; femur, 0.45; tibia, 0.47; 1st segment, 0.07.

C o m p a r i s o n. By the 2+10-segmented antennae, arrangement of wing veins, long flagellomeres new species is closely related to R. operculata Fedotova et Sidorenko, 2006 , differing from it by the elongated segments of 2-segmented palpi; very short neck and wide basal enlargement of middle flagellomeres; narrowing distally 10th flagellomere; in wing, that is enlarged distally (not proximally); in R 4+5, that joining C before wing apex, and smaller body size.

E t y m o l o g y. The species is named after Andrej Parkalov who permanently helps the amber collection of Schmalhausen Institute of Zoology, NAS of Ukraine.

Figures

Fig. 5. Rhizomyia parkalovi, female, holotype (SIZK, N K–7385): 1 — general appearance; 2 — head; 3 — palpus; 4 — flagellomeres 9–10; 5, 6 — tarsomere 1 (variability of shape); 7 — flagellomere 5; 8 — apex of abdomen. Scale bars 0.1 mm.

Fig. 1. Rovnobrachineura kiryeyevi, male, holotype (SIZK, N K–3701): 1 — general appearance; 2 — scape, pedicel, flagellomeres 1–2; 3 — flagellomeres 8–10; 4 — flagellomeres 6–7; 5 — mouth parts and palpi; 6 — genitalia dorsally. Scale bars 0.1 mm.

Fig. 2. Brachineura polessica, female, holotype (SIZK, N K–1236): 1 — general appearance; 2 — scape, pedicel, flagellomeres 1–2; 3 — flagellomeres 9–10; 4 — flagellomeres 6–7; 5 — flagellomeres 3–5; 6 — palpus; 7 — middle tarsomeres 3–5; 8 — middle tarsomeres 1–2; 9 — middle tarsomere 1; 10 — middle tarsomere 5; 11 — ovipositor laterally. Scale bars 0.1 mm.

Fig. 3. Popovineura nacta, male, holotype (SIZK, N K–666): 1 — general appearance; 2 — scape, pedicel, flagellomeres 1–2; 3 — flagellomeres 5–6; 4 — flagellomeres 8–11; 5 — palpus; 6 — fore tarsus; 7 — fore tarsomere 5; 8 — genitalia. Scale bars 0.1 mm.

Fig. 4. Nascensluna mellea, female, holotype (SIZK, N K–1601): 1 — general appearance; 2 — flagellomeres 8–10; 3 — flagellomere 5; 4 — ovipositor. Scale bars 0.1 mm.

Fig. 6. Ledomyia olgae, male, holotype (SIZK, N K–3697): 1 — general appearance; 2, 3 — palpi (variability of shape); 4 — flagellomeres 9–11; 5 — flagellomere 5; 6, 7 — genitalia (6 — ventral view,7 — dorsal view).Scale bars 0.1 mm.

Fig. 7. Rovnobrachineura kiryeyevi, male, holotype SIZK, N K–3701 (1–3); Brachineura polessica, female, holotype (SIZK, N K–1236) (4, 5); Rhizomyia parkalovi, female, holotype (SIZK, N K–7385) (6, 7): 1, 4, 6 — head and antenna (1 — ×129; 4 — ×62; 6 — ×117); 2 — genitalia (×283); 3, 5, 7 — general appearance (3 — ×46; 5 — ×29; 7 — ×39).

Fig. 8. Popovineura nacta, male, holotype (SIZK, N K–666) (1, 2); Ledomyia olgae, male, holotype (SIZK, N K–3697) (3, 4); Nascensluna mellea, female, holotype (SIZK, N K–1601) (5–7): 1–3, 7 — general appearance (1 — ×45; 2 — ×38; 3 — ×48; 7 — ×64); 4 — antennae (4 — ×129); 5 — hind leg (×82); 6 — head and antenna (×131).