Brachineura polessica Fedotova et Perkovsky, 2014

Fedotova, Z. A. & Perkovsky, E. E., 2014, New Gall Midges (Diptera, Cecidomyiidae, Brachineurini, Ledomyiini) From The Late Eocene Rovno Amber, Vestnik Zoologii 48 (1), pp. 35-50 : 40

Brachineura polessica Fedotova et Perkovsky , sp. n.

M a t e r i a l. Holotype. SIZK N K–1236 , poorly preserved inclusion of female with deformed wing and partly missing tarsi, Klesov , Rovno amber, Late Eocene ( fig. 2 , 1 –11; 7, 4, 5). Syninclusions: K–1233: Phoridae ; K–1234: Dolichopodidae ; K–1235: Dolichopodidae ; K–1236: spider web; K–1237: Collembola Entomobryomorpha.

D e s c r i p t i o n. F e m a l e ( fig. 2 , 1 –11; 7, 4, 5). Body is very wide, as long as wings, 1.7 times longer than antennae. Wings are 1.7 times as long as antennae. Head with wide eye bridge. Head length 1.7 times as long as width. Scape is enlarged distally, longer than transversal pedicel. First flagellomere is 1.8 times as long as wide; 2nd flagellomere is as long as 1st one, with very short neck. Middle flagellomeres are almost parallelsided, 5th flagellomere is 2.6 times as long as wide, almost as long as 1st flagellomere one. Lengths of 9th and 10th flagellomeres are equal, 9th is narrowed distally, apically rounded. Flagellomeres with basal and medial whorls of very long setae. Basal enlargement with rings of sensorial filae, sometimes with little wide loops around lateral sides of flagellomeres. Palpi are 2-segmented, very narrow, almost parallelsided laterally, 2nd 1.4 times as long as 1st. Wing is deformed. Vein Cu is simple, without fork, curved. Fore and middle femur and tibia are shorter than hind femur and tibia. All legs with femur that longer than tibia or have almost equal length with it. First tarsal segment with narrow apical projection laterally ( fig. 2 , 9). Second middle tarsomere 4.8 times as long as 1st. Tarsal claw is simple, hook-form. Ovipositor is directed dorsocaudally, apical lobes are wide-oval, covered by long setae.

M e a s u r e m e n t s, mm: body length, 1.54; antennal length, 0.91; head length, 0.36; width of lateral side of head, 0.21; palpus length, 0.14;1st segment, 0.06; 2nd segment, 0.08 mm; wing length, 1.54; wing width 0.44 (unclear); length of fore: femur, 0.50; tibia, 0.52; 1st tarsal segment 0.10; length of middle: coxa, 0.18; trochanter, 0.06; femur, 0.55; tibia, 0.53; tarsus, 1.06: 1st segment, 0.10, 2nd, 0.48; 3rd, 0.19; 4th, 0.15; 5th, 0.13; length of hind: femur, 0.55; tibia, 0.58; metatarsus, 0.10.

C o m p a r i s o n. By the shape and size of the body, 2+10-segmented antennae, simple vein Cu, short legs and form of large round plates of ovipositor the new species is closely related to Brachineura (s. str.) enigmatosa Fedotova et Sidorenko, 2006, differing from it by shorter 2-segmented palpi; short and wide 1st and 2nd flagellomeres; very little neck of middle flagellomeres and presence of numerous round loops of sensorial filae around middle flagellomeres; presence of 1–2 swollen peritremae on basal enlargement of flagellomeres.

E t y m o l o g y. From the toponym Polesje (adj.).

Figures

Fig. 2. Brachineura polessica, female, holotype (SIZK, N K–1236): 1 — general appearance; 2 — scape, pedicel, flagellomeres 1–2; 3 — flagellomeres 9–10; 4 — flagellomeres 6–7; 5 — flagellomeres 3–5; 6 — palpus; 7 — middle tarsomeres 3–5; 8 — middle tarsomeres 1–2; 9 — middle tarsomere 1; 10 — middle tarsomere 5; 11 — ovipositor laterally. Scale bars 0.1 mm.

Fig. 1. Rovnobrachineura kiryeyevi, male, holotype (SIZK, N K–3701): 1 — general appearance; 2 — scape, pedicel, flagellomeres 1–2; 3 — flagellomeres 8–10; 4 — flagellomeres 6–7; 5 — mouth parts and palpi; 6 — genitalia dorsally. Scale bars 0.1 mm.