Therates ziyardamensis Matalin et Wiesner, 2023

Matalin, Andrey V. & Wiesner, Jürgen, 2023, Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae), Zootaxa 5256 (5), pp. 401-433 : 409

Therates ziyardamensis Matalin et Wiesner , sp. n.

Figs 12 , 35 , 64 , 96 , 131 .

Therates fruhstorferi vitalisi W. Horn, 1913 — Wiesner 1999: 372; Cassola & Klícha 2002: 39.

Type material. HOLOTYPE, ♂— MYANMAR N (Burma), 65 km NW Putao , 1250 m, Zi Yar Dam vill., 18– 21.05.1998, leg. S. Murzin & V. Sinyaev ( JW).

Diagnosis. This new species is reliably distinguished from T. vitalisi , T. biserratus , T. hunanensis sp. n., T. sauteri and T. motoensis by apex of aedeagus without lateral carinae ( Fig. 131 vs. Figs 152–174 ). From T. fruhstorferi and T. pseudovitalisi sp. n. this new species is recognized by thin apex of aedeagus ( Fig. 131 vs. Figs 115–130 and Figs 132–135 , respectively) as well as by the rough punctuation of the elytra ( Figs 12 , 96 vs. Figs 1, 3–11 , 81–95 ).

Description. TL = 10.6 mm. Head black with distinct violet reflection; orbital plates finely but distinctly striated in posterior third; frons semi-circular, with distinct impression and transverse rugoses (see above), slightly downward anteriorly (see at side), frontal sulci very deep, distinctly convergent in anterior half and shallow and sharply divergent posteriorly; occiput flat with shallow impression anteriorly. Mandibles dark brown underside, yellow-pale topside, with brown teeth and apical molar. Labial palpi pale with dark-brown apical palpomeres; maxillary palpomeres 1 and 2 light brown, palpomeres 3 and 4 dark brown. Antennae not extend shoulders, black, antennomeres 1–5 with light violet tinge and brown apices, antennomeres 9–11 club-shaped, antennomeres 9 and 10 serrated, with downward anterior lower margins, antennomere 11 clearly dilated with abrupt apical margin ( Fig. 35 ). Labrum as wide as long (LL = LW— 1.4 mm), black-brown with large elongate apical yellow spot unseparated from central apical teeth ( Fig. 64 ).

Pronotum shining black with blue-violet reflection; as wide as long (PL = PW— 1.9 mm), apical lobe slightly wider in medial part; thorax black with distinct blue reflection.

Fore femora pale along anterior margin and black with violet tinge along posterior margin; middle femora pale in basal half and black with violet tinge in apical half; hind femora pale in basal third and black with violet tinge in apical 2/3; tibiae black with distinct violet lustre; tarsi black with light violet tinge except brown apices of all apical tarsomeres; HTbL/HTaL = 1.22.

Elytra black-blue with bright purple-violet reflection; indistinctly divergent in apical third, EL/EW = 1.94 (EL = 6.6 mm, EW = 3.4 mm); punctuation deep and regular in basal 3/4 except humeral area, in apical quarter shallow and tenuous; scutellum black with violet tinge, apical margin rounded but shortly top-cut. Elytral pattern presented by small pale basal portion of humeral lunule (see in front) and relatively short practically transverse central dot indistinctly dilated in lateral portion ( Figs 12 , 96 ).

Aedeagus (only apical half) with thin elongate slightly sinuate rounded apex, without lateral carinae ( Fig. 131 ).

Female unknown.

Etymology. This new species is named after the type locality—Zi Yar Dam village.

Distribution. MYANMAR: Kachin State ( Fig. 184 ).

Figures

FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png.

FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels.

FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales.

FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes.

FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype.

FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm.

FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm.