Rhyacobates zetteli Tran & Nguyen, 2016

Leng, Zhaoqi, Tran, Anh Duc & Ye, Zhen, 2023, Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae), European Journal of Taxonomy 893 (1), pp. 1-73 : 66-67

Rhyacobates zetteli Tran & Nguyen, 2016

Figs 3R , 5Q , 7H

Rhyacobates zetteli Tran & Nguyen, 2016: 505–508 , figs 11–23 (original description).

Diagnosis

Body length of apterous females 9.80–10.60, of apterous males 7.30–7.70. Color ( Fig. 3R ): dorsum of head mainly yellow with small black markings; pronotum mainly yellow with narrow black marking on anterior margin and sometimes laterally; mesonotum with broad median marking from anterior to posterior margin (median marking broader in females than in males); metanotum with laterally expanded yellow marking (more extensive in females). Female: abdomen elongate and straight; ventral length of pregenital abdomen about half of body length ( Fig. 5Q ); connexivum of segments I–VI narrow; connexival corners of segment VI with small processes; abdominal segment VII tapering towards apex; mediotergite VII with small median process on posterior margin; sternum VII with bilobate posterior margin, connexival corners with small blunt processes which bend mesad. Male: middle trochanter with 6–7 spines; middle femur with spines but not in distinct row; length of middle tibia about equal to length of hind tibia; pygophore simple, with straight apical margin; proctiger with subtrapezoidal lobes laterally ( Fig. 7H ); paramere relatively long and slender, curved at basal third, distal part tapering towards narrowly rounded apex.

Material examined

Type specimens

See Tran & Nguyen (2016).

Non-type specimens

VIETNAM • 3 ♀♀ (apterous); Lào Cai Province, Sa Pa, Núi X ẻ area, Vàng stream and its feeders; 22°20′54.4″ N, 103°46′12.0″ E; 25 Oct. 2020; A.D. Tran et al. leg.; TAD20-19; ZVNU • 1 ♀ (apterous); same collection data as for preceding; NKUM .

GPS data of previous records: see Tran & Nguyen (2016).

Distribution

Vietnam: Lào Cai ( Tran & Nguyen 2016).

Comparative notes

Rhyacobates zetteli is most similar to R. angustus , as discussed by Tran & Nguyen (2016: 503 , 507– 508). Also see comparative notes under R. bui sp. nov.

Figures

Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm.

Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm.

Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran & Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen & Chen, 1995. D. R. recurvus Andersen & Chen, 1995. E. R. scorpio Andersen & Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran & Nguyen, 2016. Scale bar = 0.2 mm.