Levizonus malewitschi Lokschina & Golovatch, 1977

Mikhaljova, Elena V., 2021, Review of the millipede genus Levizonus Attems, 1898, with description of a new species from the Far East of Russia (Diplopoda, Polydesmida, Xystodesmidae), European Journal of Taxonomy 751, pp. 159-184 : 169-171

Levizonus malewitschi Lokschina & Golovatch, 1977

Figs 6 , 13

Levizonus malewitschi Lokschina & Golovatch, 1977: 73 , figs 1, 2 (holotype ♂, from Derevyankina Ravine, Kavalerovskiy District, Primorskiy krai, Russia, in ZMUM).

Levizonus malewitschi – Lokšina & Golovatch 1979: 385. — Kurcheva & Mikhaljova 1980: 120. — Mikhaljova 1981a: 66, map (fig. 5); 1983: 81; 1990: 136; 1993: 33; 1998: 52, figs 196–199, map 12; 2004: 243, figs 608–611, map 32; 2009a: 5; 2017: 294, figs 656–659, map 43. — Mikhaljova & Petukhova 1983: 53. — Gromyko 1990: 63. — Ganin 1997: 124; 2011: 341. — Marek et al. 2014: 72.

Levizonus malevitschi [sic!] – Tanabe 1994: 108, fig. 4; 2002: 2178.

Diagnosis

The species differs from its congeners mainly by the configuration of the gonopod telopodite apex, like two plates placed perpendicular to each other, the central one with a seminal groove, the second one being serrate at its outer margin ( Fig. 6A–B ) (vs without two plates in all congeners).

Material re-examined (specimens published by Mikhaljova 1993)

RUSSIA • 1 ♀; Primorskiy krai, Sikhote-Alin State Nature Biosphere Reserve, Blagodatnoe ; 22 Sep. 1982; M.N. Gromyko leg.; Quercus mongolica Fisch. ex Ledeb. forest; FSCB 11982 • 1 ♂; same locality as for preceding; 26–27 Jun. 1986; M.N. Gromyko leg.; FSCB 11986 .

Distribution

Russia: Far East, Primorskiy krai.

Remarks

Originally described from Kavalerovskiy District, Primorskiy krai, Russia ( Lokschina & Golovatch 1977), this species appears to be distributed in eastern and central parts of the Primorskiy krai ( Mikhaljova 2017). The abundance ranges from 3.5 ind./m 2 in mountainous forest to 62 ind./m 2 in valley forests ( Kurcheva & Mikhaljova 1980). Vulva as in Fig. 6C .

Figures

Fig. 6. Levizonus malewitschi Lokschina & Golovatch, 1977, ♂ (FSCB 11986) and ♀ (FSCB 11982). A. Gonopod, lateral view. B. Gonopod apex, mesal view. C. Vulva, mesal view. D. Gonopod, mesal view. Scale bars: A = 200 μm; C–D = 100 μm; B = 10 μm.

Fig. 13. Distribution of species of the genus Levizonus Attems, 1898. Filled square: L. circularis Takakuwa, 1942. Filled circle: L. distinctus Mikhaljova, 1990. Open square: L. laqueatus Mikhaljova, 1981. Open triangle: L. nakhodka Mikhaljova sp. nov. Filled triangle: L. malewitschi Lokschina & Golovatch, 1977. Filled diamond: L. thaumasius Attems, 1898. Heart: L. takakuwai (Verhoeff, 1941). Oval: L. montanus (Takakuwa, 1941).