Lanceobostra guatemalensis ( Redtenbacher, 1908 ) Hennemann & Conle, 2024
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publication ID |
https://doi.org/ 10.11646/zootaxa.5444.1.1 |
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publication LSID |
lsid:zoobank.org:pub:5DE4A9DD-99F7-4E23-AD50-58DC491BB75E |
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persistent identifier |
https://treatment.plazi.org/id/03FD87D9-FFC9-D80E-FF55-F5D32A38E213 |
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treatment provided by |
Plazi |
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scientific name |
Lanceobostra guatemalensis ( Redtenbacher, 1908 ) |
| status |
comb. nov. |
Lanceobostra guatemalensis ( Redtenbacher, 1908) comb. n.
( Figs. 36G View FIGURE 36 , 39K View FIGURE 39 , 40Q–S View FIGURE 40 , 95F View FIGURE 95 )
Clonistria guatemalensis Redtenbacher, 1908: 405 View in CoL .
Shelford, 1909: 359.
Zompro & Brock, 2003: 13. [Lectotype designation]
Otte & Brock, 2005: 107.
Zompro, 2005: 264.
Brock & Büscher, 2022: 512.
= Bostra intermedia Carl, 1913: 40 View in CoL . syn. n.
Phanocles intermedius, Zompro & Brock, 2003: 15 View in CoL . [Type data] Otte & Bock, 2005: 262.
Brock & Büscher, 2022: 513.
Further material examined [1 ♂]:
♂: Moca, Guatalon, 1.000 m, Mar. Apr. 31; Guatemala; J. Bequaert [ ANSP] .
Diagnosis. The previously unknown ♂♂ of this species are very similar to those of L. incolum is (Brunner v. Wattenwyl, 1907 ) and share with this species the strongly deflexed and almost semi-circular lateral margins of abdominal tergum IX ( Fig. 40Q View FIGURE 40 ). They can however be separated by the slightly less elongate head, less narrowed posterior portion of the anal segment ( Fig. 40R View FIGURE 40 ), which has the lateral margins almost entirely parallel-sided, more obtose central protuberance of the poculum ( Fig. 40Q View FIGURE 40 ) and buff, differently shaped vomer, which is much narrower and elongate and has a deeply impressed medio-longitudinal furrow along its ventral surface ( Fig. 95F View FIGURE 95 ; grey, broadly oval, flattened and with several fine, transverse ridges in incolumis ). Also ♀♀ strongly resemble those of L. incolumis but differ by the shape of the head, which is gradually narrowed towards the posterior (rather parallel-sided in incolumis ), much less pronounced spines of the meso- and metapleuare and notably more distinct and longer spines of the praeopercular organ ( Fig. 39K View FIGURE 39 ). From the similar L. freygessneri ( Redtenbacher, 1908) comb. n. ♀♀ can be distinguished by the considerably stockier shape (mesonotum 7.6x vs. 8.8x longer than pronotum in freygessneri ), much more prominent spines of the praeopercular organ and slender basitarsi (with a rounded medio-dorsal lobe in freygessneri ).
Description. ♂♂ ( Fig. 36G View FIGURE 36 ): Medium sized (body length 94.5 mm), very slender and typical for the genus with a large and obtuse central protuberance of the poculum and slender club-like cerci. Body surface smooth and slightly glossy. General colour pale buff, the abdomen somewhat darker (most certainly due to preservation. Head chestnut brown between the eyes, genae with a faint and washed brown postocular streak, the lower portion dark yellow. Lateral margins of pronotum broadly black, the posterior transverse membrane dark brown. The lateral surfaces of abdominal tergum IX with an indistinct washed whitish longitudinal streak. All tibiae with three faint pale cream annulae. Antennae dark reddish brown at the base and gradually becoming lighter with the red tone fading towards the apex.
Head: Moderately elongate, flattened, 1.5x longer than wide, broadest at the eyes and gradually narrowing towards the posterior. Frons with a distinct C-shaped impression and area between the eyes with two very shallow swellings. Eyes circular in outline, projecting hemispherical and their diameter contained about 2.1x in length of genae. Antennae reaching one-third the way along abdominal segment V. Scapus somewhat compressed dorsoventrally and oval in cross-section, roundly rectangular in dorsal aspect and about 1.3x longer than wide. Pedicellus round in cross-section and about three-fifth the length of scapus. III strongly constricted towards the apex and notably longer than pedicellus.
Thorax: Pronotum about as long but slightly narrower than head, basically rectangular in dorsal aspect with a slight narrowing pre-medially; about 1.6x longer than wide. Transverse median sulcus moderately distinct, straight and expanding only over about half of width of segment, the medio-longitudinal line distinctly impressed in anterior half of segment. Mesothorax 9.4x longer than pronotum, uniform in diameter and just slightly weakly at posterior margin. Mesonotum with a fine, punctured furrow parallel to lateral margins. Metanotum like mesonotum, about 0.4x of whose length. Mesosternum with a very weak medio-longitudinal carina, metasternum smooth and simple.
Abdomen: Median segment about two-thirds the length of metanotum and constricted medially. Segment II somewhat longer than median segment, II–IV uniform in length, V–VI slightly gradually decreasing in length; all rather uniform in width but constricted medially; IV 6x and VI 5x longer than wide. VII shorter than all preceding and somewhat widening in the posterior one third; about four-fifth the length of VI. Tergum VIII trapezoidal, gradually widening towards the posterior, a little less than two-thirds the length of VII and with a fairly obtuse longitudinal ridge laterally. IX widest segment and almost 2x wider than II–VI, the posterior portion narrowed and the lateral margins strongly deflexed and almost semi-circular in outline; lateral surface with a prominent and obtuse, gently curved longitudinal bulge. Anal segment only about two-thirds the length of IX, roundly rectangular in dorsal aspect with the posterior margin somewhat labiate, swollen and weakly indented medially and the outer angles obtusely rounded; 1.5x longer than wide ( Fig. 40R View FIGURE 40 ). Ventral surfaces of the outer lateral angles set with minute denticles. Epiproct very small, triangular and fully hidden under anal segment. Vomer elongate-triangular in general shape and gradually narrowing towards a single, arcuate terminal hook; ventral surface with a distinctly impressed medio-longitudinal furrow and the lateral portions notably inflated in median portion ( Fig. 95F View FIGURE 95 ). Cerci slender, gently curved towards the posterior and club-shaped with the swollen apical portion somewhat indented interiorly. Poculum moderately bulgy, cup shaped and angular in lateral aspect with a prominent and obtusely conical central protuberance ( Fig. 40Q View FIGURE 40 ), the posterior margin rounded and not reaching to posterior of abdominal tergum IX ( Fig. 40S View FIGURE 40 ).
Legs:All long, slender and entirely unarmed, profemora almost as long as head, pro- and mesothorax combined, mesofemora slightly shorter than mesothorax and metafemora reaching to posterior margin of abdominal segment V with metatibiae projecting considerably beyond apex of abdomen. Basitarsi elongate, slender and about as long as remaining tarsomeres combined.
Comments. Although meant to support future studies on the group and nomenclatural stability, the lectotype designation for Clonistria guatemalensis by Zompro & Brock (2003: 13) was very unlucky and is questionable in several aspects. The type series consists of three specimens all from Guatemala but which are two distinct species. One species is represented by the adult ♂ in the collection of MHNG, which perfectly matches with the original description and measurements provided by Redtenbacher (1908: 405). The other two specimens, a ♀ in MNHU and a ♂ in MHNG are both immature but congeneric, the latter of which was not definitely cited by Redtenbacher. Having based his description of Clonistria guatemalensis on the adult ♂ in MHNG (No. 3/28), Redtenbacher added the immature ♀ in MNHU as the opposite sex with doubt. Unfortunately, Zompro & Brock (2003: 13) selected the only specimen that had not been definitely cited by Redtenbacher as being part of the type-series, the immature ♂ in MHNG (No. 2/14), as the lectotype of C. guatemalensis . This specimen does not at all match with the original description, measurements or key features used by Redtenbacher and sadly results in a synonymy of C. intermedia Carl, 1913 . The lectotype bears exactly the same data and labels as the holotype of B. intermedia and agrees in most important features, hence is the opposite sex and here synonymised (syn. n.). The MNHU specimen is obviously an immature ♀ of intermedia . As a further effect of Zompro’s unlucky lectotype designation, now a new name becomes necessary for the adult ♂ paralectotype in MHNG, the specimen on which C. guatemalensis had in fact been based on by Redtenbacher. It is therefore here named Phanocles mexicanus sp. n. The previously unknown ♂ is here described based on a specimen in the collection of ANSP. Eggs unknown.
Table 17: Measurements of Lanceobostra guatemalensis ( Redtenbacher, 1908) comb. n.
* According to Carl (1913: 41), tip of subgenital broken off
| ANSP |
Academy of Natural Sciences of Philadelphia |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Lanceobostra guatemalensis ( Redtenbacher, 1908 )
| Hennemann, Frank H. & Conle, Oskar V. 2024 |
Phanocles intermedius
| Zompro, O. & Brock, P. D. 2003: 15 |
Clonistria guatemalensis
| Redtenbacher, J. 1908: 405 |