Dietomorpha Reymond, 1938

Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, European Journal of Taxonomy 809, pp. 1-71 : 25-26

publication ID	https://doi.org/ 10.5852/ejt.2022.809.1719
publication LSID	lsid:zoobank.org:pub:A754493E-5466-4479-B515-AABEDDE09D93
DOI	https://doi.org/10.5281/zenodo.6449866
persistent identifier	https://treatment.plazi.org/id/03FC8666-5572-FF9F-500E-BA96FC8EF8CC
treatment provided by	Felipe
scientific name	Dietomorpha Reymond, 1938
status

Treatment

Genus Dietomorpha Reymond, 1938

Figs 3 View Fig , 17 View Fig , 23C View Fig , 26C View Fig

Dietomorpha Reymond, 1938: 143 .

Dietomorpha – Kühnelt 1957: 86, fig. 4.

Kawiria – Kaszab 1957: 295–296 (erroneous interpretation of Dietomorpha pardalis ).

Type species

Dietomorpha pardalis Reymond, 1938 , by monotypy.

Species included

Dietomorpha pardalis Reymond, 1938 , Dietomorpha gonzalesi S. Chigray & Nabozhenko sp. nov.

Diagnosis

Body robust ( Fig. 3 View Fig ), completely covered with lanceolate longitudinally striated, creamy scales (each scale with median longitudinal ridge at middle) ( Fig. 9D, F View Fig ), with spotted ( Fig. 2A View Fig ) or striped ( Fig. 3C, E–G View Fig ) pattern of dark scales on elytra and dark spots on pronotum. Body under scales black or darkbrown. Body length 9–14 mm. Epistoma and frons covered with sparse, erected yellowish setae. Eyes large, circular, convex in dorsal view; surface behind eyes concealed by anterior margin of pronotum.

Pronotum slightly transverse (1.4–1.7× as wide as long); lateral margins and partly prothoracic hypomera basally with bare and smooth black spot. Disc of pronotum (Fig. 52) convex, with two triangular deep impressions at base. Pronotum covered with sparse, erect, conical tubercles becoming larger along lateral margins, each tubercle with spine at apex. Prosternum without transverse triangular depression along anterior margin, longitudinal length 2.2× as short as longitudinal length of one procoxa. Prosternal process elongated and narrow (length 3 × width), raised or not raised between procoxae, strongly ( D. pardalis ) ( Fig. 3B View Fig ) or weakly ( D. gonzalesi S. Chigray & Nabozhenko sp. nov.) ( Fig. 3D View Fig ) protruding, not touching with mesoventrite.

Elytra oval or widely oval, with one ( D. gonzalesi S. Chigray & Nabozhenko sp. nov.) or double in middle ( D. pardalis ) humeral rib; with strong, acute spinose tubercles (with spine in base of each tubercle) ( Fig. 9E–F View Fig ) on dark spots ( D. pardalis ) or with five longitudinal rows of spinose tubercles ( D. gonzalesi S. Chigray & Nabozhenko sp. nov.). Scutellar shield completely or partly concealed by base of pronotum; surface around scutellar shield with triangular depression. Transverse length of metacoxae subequal to intercoxal process of abdominal ventrite 1.

Trochanters with elongate, sparse setae. Meso- and metafemora curved outward. Protibiae with projecting process at apex of outer margin, armed with tuft of spines ( Fig. 26B View Fig ). Lateral margins of protibiae with long, fine setae and short spines. Protibial terminal spurs elongated, extending to base of protarsomere 4; mesotibial terminal spurs extending to apex of mesotarsomere 2; metatibial terminal spurs extending to midlength of metatarsomere 1. Outer and inner protibial terminal spurs subequal in length, inner meso- and metatibial terminal spurs longer than outer ones, spurs flattened from sides. All tarsi flattened from sides, covered with long setae dorsally and short setae ventrally. Tarsal claws elongated, thin, weakly curved outward.

Male genitalia ( Fig. 17A–H View Fig )

Inner sternite VIII ( Fig. 17H View Fig ) weakly sclerotized on margins, densely covered with long, dense setae; anterior margin deeply emarginated in middle; gland is absent (or at least, was not observed).

Rods of spiculum gastrale ( Fig. 17G View Fig ) widely spaced, thickened, arcuately connected at apex, with wide membrane along this connection and near derivatives of inner sternite IX; these derivatives elongated, ladle-shaped, evenly sclerotized in middle and slightly more sclerotized terminally; apical margin of derivatives covered with short dense setae.

Tegmen of aedeagus thickened, obtuse ( Fig. 17A–C View Fig ). Basal piece of tegmen much shorter and wider than apical piece, with tubercle-shaped apex ( Fig. 17A–B View Fig ). Apical piece strongly curved, narrowly rounded at apex, covered with short setae in apical half; ventral apophyses not expressed, dorsal apophyses moderately long, triangular. Median lobe ( Fig. 17D, F View Fig ) widened in basal ⅔ and narrowed in apical third, with weakly separated apex, strongly curved; basal part with additional four sclerotized armatures; baculi not merged at apex.

Female genitalia

Spiculum ventrale ( Fig. 17I View Fig ) very short and strongly widened, sternite VIII with long acutely angulate curved apophyses.

Ovipositor ( Fig. 17G–L View Fig ) short, weakly sclerotized. Coxite with four distinct not merged lobes. Baculi of coxite lobe I short, shorter than paraproct baculi; lobe II small, sclerotized; lobe III larger, conical, sclerotized; lobe IV membranous (only at apex slightly sclerotized), conical, with dense pubescence of long setae. Paraproct V-shaped on each side ventrally, baculi sharply widened in base and narrow in apical third. Vulva with narrow sclerotization in the middle of ventral side. Apical margin of proctiger with deep V-shape emargination, baculi of proctiger strongly widened.

Female genital ducts ( Fig. 23C View Fig ) Vagina moderately widened, sacciform, narrower before oviduct and after spermatheca, apical part of vagina not elastically curved. Spermatheca very short, single-tube. Accessory gland long, with two tubelike constrictions, short in base and long in middle. Basal part between two constrictions not sacciform.

Distribution

Iran and Pakistan (Balochistan region), Afghanistan (Registan Desert).

Notes

Koch (1964) synonymized Dietomorpha pardalis and Kawiria szekessyi Kaszab, 1957 (junior synonym). Koch also noted that Reymond (1938) should be considered as the author of the genus Dietomorpha and the species Dietomorpha pardalis , because he presented figures for this species. Kaszab (1957) noted that Reymond (1938) did not give a description or a differential diagnosis of the genus, and the author of Dietomorpha pardalis should therefore be Kühnelt (1957), who distinctly described, compared and figured this taxon. Later Kwieton (1982) cited Koch’s work but incorrectly established the following synonymy: Kawiria pardalis Reymond, 1938 , comb. n. = Kawiria szekessyi and Dietomorpha Reymond, 1938 = Kawiria Schuster, 1957 . Medvedev (2005) was not aware of these works and re-established the synonymy previously proposed by Koch, and he also recognised the author of the binomen Dietomorpha pardalis Kühnelt, 1957 .