Enema endymion Chevrolat 1843, 1985

Alvarez, Héctor Jaime Gasca, da, Claudio Ruy Vasconcelos, Fonseca & Ratcliffe, Brett C., 2008, Synopsis of the Oryctini (Coleoptera: Scarabaeidae: Dynastinae) from the Brazilian Amazon, Insecta Mundi 2008 (61), pp. 1-62 : 11-13

Enema endymion Chevrolat 1843

( Fig. 14 )

Enema endymion Chevrolat 1843: 29

Enema lupercus Burmeister 1847: 236 (synonym)

Enema paniscus Burmeister 1847: 236 (synonym)

Enema gibbicollis Sternberg 1908: 24 (synonym)

DESCRIPTION. Length: 32.4. Width: 18.5 mm (Range species. Length: 30.0- 36.3 mm; width: 17.0-20.0 mm ( Ratcliffe 2003)). Color: Light to dark reddish brown.

Males. Head: Frons with short to moderately long horn; horn narrowly triangular, acuminate, recurved. Margin of eye canthus weakly crenulate, surface punctate, apex rounded. Clypeus with surface rugopunctate, apex broad, emarginate. Antenna with 10 segments, club subequal in length to segments 2-7. Mandibles broad, apex strongly bidentate. Pronotum: Surface rugose along lateral margins and surrounding discal depression, with small punctures. Disc in anterior half with strong, longitudinal depression. Anterior margin with strong emargination at center. Elytra: Surface with strongly impressed sutural stria, with small dense punctures and larger sparse punctures, sides strongly rugose and explanate. Pygidium : Base with transverse row of large, setigerous punctures. Apical margins rugose. In lateral view, surface convex. Legs: Protibia quadridentate, occasionally with all teeth equidistant. Apex of posterior tibia weakly crenulate and with small tooth. Basal segment of posterior tarsus triangularly elongated at apex. Venter: Prosternal process short, laminar, triangular. Parameres: Similar to those of E. pan .

Females. As males except in the following respects: Head: Frons with horn usually shorter. Pygidium: Surface nearly flat, weakly convex ( Ratcliffe 2003; Ratcliffe and Cave 2006).

DIAGNOSIS. The reddish brown color and the short acuminate head horn in both sexes will distinguish this species from E. pan .

DISTRIBUTION. Enema endymion is found from southern Mexico to Brazil and Bolivia, including the Amazon region. It is a species with few records in Brazil, especially in the Amazon region, and is very rare in research collections.

LOCALITY RECORDS. ( Fig. 14 ) 1 specimen examined (1 male). A specimen was seen from the following collection : MPEG.

PARÁ (1). Benevides.

TEMPORAL DISTRIBUTION. December (1).

BIOLOGY. The larval stages were described by Ratcliffe (2003). Larvae probably feed on humus, roots and organic debris in the soil ( Ratcliffe 2003; Morón et al. 1997). Adults are nocturnal and are attracted to lights. It has been reported that the larvae are preyed upon by birds and mammals ( Ratcliffe 2003).

Figures

Figure 14. Distribution of Enema endymion in Brazilian Amazonia.