Schoutedeniastes Burgeon, 1941

Volkovitsh, Mark G., 2016, Svatactesis gen. nov., a new genus for Polyctesis johanidesi with notes on the tribe Polyctesini (Coleoptera: Buprestidae: Polycestinae), Acta Entomologica Musei Nationalis Pragae 56 (1), pp. 5-15 : 13-14

Schoutedeniastes Burgeon, 1941

S. magnifica species-group ( Figs 7 , 21 ): S. magnifica (Waterhouse, 1875) (= S. kerremansi (Obenberger, 1924)) ; Afrotropical Region.

S. amabilis species-group ( Figs 8, 12 , 22–23 ): S. amabilis (Laporte & Gory, 1835) , S. apicata (Waterhouse, 1882) , S. birmanica (Thery, 1947) (= S. rondoni (Baudon, 1962)) , S. duaulti (Baudon, 1962) , S. hieroglyphica (Théry, 1904) , S. okhurai (Akiyama & Ohmomo, 1992) ,? S. hatai (Ohmomo & Akiyama, 1994) (not studied); Oriental Region and East Palaearctic.

S. vitalisi species-group ( Figs 9 , 24–26 ): S. igorrota (Heller, 1891) (with subspecies aenea Hoscheck, 1931), S. vitalisi (Bourgoin, 1922) (= S. consobrina (Bourgoin, 1922; S. vitalisi (Obenberger, 1924)) ; Oriental Region and East Palaearctic.

The differences in some character states ( Table 1 ) between these species-groups cast doubts on the monophyly of Schoutedeniastes . The only reliable characters to distinguish this genus from Polyctesis are the colour pattern, wider subhumeral lobe of elytra, nearly covering metepisternum, and the structure of clypeus (shape of anterior emargination, degree of reduction of the lateral branches) and, correspondingly, antennal sockets (open vs. closed). The length of the 2 nd elytral stria of the species of S. magnifica and S. amabilis species-groups (2 nd stria short) are similar to Svatactesis gen. nov. while the species of S. vitalisi species-group (2 nd stria long) are similar to Polyctesis and Bellamyina . Interestingly, the penis structure of Schoutedeniastes vitalisi ( Fig. 26 ) is rather similar to those of Xenopsis violaceocyanea Volkovitsh, 2008 and X. kubani Volkovitsh, 2008 ( Polycestini : Xenopseina ) (see VOLKOVITSH 2008, Figs 78, 80). In this way, S. vitalisi species-group of Schoutedeniastes may be a link between Polyctesini and Polycestini which in turn gives rise to doubts regarding the tribal level of Polyctesini . A detailed taxonomic revision of all the taxa attributed to Polyctesini is needed to clarify its generic composition and relations to Polycestini comprising the subtribes Xenopseina Volkovitsh, 2008 and Polycestina Lacordaire, 1857 ( VOLKOVITSH 2008).

Figures

Figs 1–12. 1–9 – habitus. 1–4 – Svatactesis johanidesi (Bílý, 1997). 1 – holotype, ♁ (NMPC), 13.6 mm. 2–4 – ♀, Turkey (Aydinlar) (VKCB), 14.3 mm:2 – dorsal view, 3 – lateral view, 4 – ventral view.5 – Polyctesis rhois Marseul, 1865, ♁, Turkey (Erzinсan) (ZIN), 11.9 mm. 6 – Bellamyina hunanensis (Peng, 1992), ♀, China (Shaanxi) (ZIN), 12.0 mm. 7 – Schoutedeniastes magnifica (Waterhouse, 1875), syntype, ♀, “Limpopo” (MNHN), 15.2 mm. 8 – S. amabilis (Laporte & Gory, 1835), ♀, Sri Lanka (MNHN), 11.2 mm. 9 – S. vitalisi (Bourgoin, 1922), ♀, syntype, Laos (Muong You) (MNHN), 12.3 mm. 10–12 – head, frontal view. 10 – Svatactesis johanidesi, ♀ (VKCB). 11 – Polyctesis rhois, ♀, Turkey (ZIN). 12 – Schoutedeniastes amabilis (Laporte & Gory, 1835),♁, India (Tamil Nadu) (NMPC). Photo: 1–4, 10 –V. Kubáň; 5–9, 11, 12 – M. Volkovitsh.

Figs 13–27. 13–26 – male genitalia.13–16 – Svatactesis johanidesi (Bílý, 1997), dorsal view:13 – aedeagus,holotype; 14–16 – paratype, Turkey (Aslanli) (NMPC): 14 – tegmen; 15 – penis; 16 – apex of penis. 17–18 – Polyctesis rhois Marseul, 1865, Turkey (Erzincan), dorsal view. 17 – tegmen; 18 – penis. 19–20 – Bellamyina hunanensis (Peng, 1992), China (Shaanxi), dorsal view: 19 – tegmen; 20 – penis. 21 – Schoutedeniastes magnifica (Waterhouse, 1875), Zimbabwe (CDFA), aedeagus, ventral view.22–23 – S.okhurai (Akiyama & Ohmomo, 1992), Thailand (ZIN), dorsal view: 22 – tegmen; 23 – penis. 24–26 – S. vitalisi (Bourgoin, 1922), Thailand (NMPC): 24–25 – tegmen: 24 – dorsal view; 25 – lateral view.26 – penis, dorsal view.27 – Svatactesis johanidesi, ♀, Turkey (Aydinlar) (VKCB), ovipositor and pregenital abdominal segments, ventral view. Photo: 13, 27 –V. Kubáň; 14–26 – M. Volkovitsh.