Lista haraldusalis ( Walker, 1859 )

Rong, Hua, Wang, Yiping, Qi, Mujie & Li, Houhun, 2021, Taxonomic review of the genus Lista Walker, 1859 from China (Lepidoptera, Pyralidae, Epipaschiinae), with descriptions of five new species, Zootaxa 5081 (2), pp. 237-262 : 248-249

Lista haraldusalis ( Walker, 1859)

( Figs 4, 5 , 11 )

Locastra haraldusalis Walker, 1859: 160 . TL: Malaysia ( Sarawak). TD: BMNH.

Lista genisusalis Walker, 1859: 877 . TL: Sarawak.

Stericta haraldusalis Hampson, 1896: 121 .

Craneophora haraldusalis (Walker) : Janse, 1931: 474.

Lista haraldusalis (Walker) : Solis, 1992: 283.

Material examined. CHINA, Beijing: 1 ♀, Mutianyu, Huairou , 23.VII.2010, coll. AH Zhang and HM Chen ; Tianjin: 2 ♀, Mt. Baxian , Ji County, 550 m, 24. VI.2001, coll. HH Li et al.; 1 ♀, Mt. Jiulong , Ji County, 200 m, 26. VI.2004, coll. HH Li et al.; 1 ♀, Heishuihe, Mt. Baxian , Ji County, 550 m, 15. VI.2009, coll. ZP Liang et al., slide No. RH15148; 1 ♂, Limutai , Ji County, 25. VI.2009, coll. ZP Liang et al.; 1 ♂, 7 ♀, Heishuihe, Mt. Baxian , Ji County, 600 m, 7–16.VII.2010, coll. YH Mou and SR Liu ; 1 ♂, Mt. Baxian , Ji County, 23.VII.2015, coll. HH Li and PX Cong , slide No. RH16131 ; 5 ♀, Mt. Jiulong , Ji County, 10–12.VII.2009, coll. WC Li ; Hebei Province: 1 ♀, Suanzaoping, Neiqiu , 670 m, 28.VII.2000, coll. HL Yu ; 1 ♀, Suobao , She County, 700 m, 1.VIII.2000, coll. HL Yu ; 1 ♀, Qingta , She County, 700 m, 3.VIII.2000, coll. HL Yu , slide No. ZDD01101 ; 1 ♀, Bulaoqingshan, Xingtai County, Xingtai , 952 m, 11.VII.2016, coll. SH Lu and H Rong ; 2 ♀, Liuzhaizi Village, Wulingshan National Nature Reserves , 539 m, 16.VII.2016, coll. SN Zhao and SR Li , slide No. RH16170 ; Shanxi Province: 1 ♀, Zhongtiaoshan Nature Reserves , 10.VII.2012, coll. Q Gao and N Chen ; 2 ♀, Zhongtiaoshan Nature Reserves , 11.VII.2012, coll. Q Gao and N Chen , slide No. RH15160 ; 1 ♀, Dahe Forest Farm, 1340 m, 14.VII.2012, coll. Q Gao and N Chen ; 2 ♀, Taikuanhe Nature Reserves, Yuncheng , 893 m, 6.VIII.2014, coll. SL Hao and YF Peng , slide No. RH16187 ; Liaoning Province: 9 ♂, 2 ♀, Laotuding , Huanren County, 28–29.VII.2012, coll. DD Zhang and LJ Yang , slide Nos. RH16341 ♂, RH16342 ♀, RH16343 ♂, RH16344 ♂; 1 ♂, Laotudingzi, Huanren , 8–9.VIII.2009, coll. WC Li and JY Liu , slide No. RH15159 ; 1 ♂, 1 ♀, Qianchang, Benxi , VI.1994, coll. JQ Ma , slide No. RH16381 ♂; 1 ♂, 1 ♀, Liaoning Forest College , VI.1994 , slide No. RH16382 ♀; 3 ♀, Mt. Tiecha , Benxi County, 28–29. VI.2010, coll. JY Liu and YP Cai, slide No. RH15154 ; 3 ♀, Wenquansi , Benxi County, 2.VII.2010, coll. JY Liu and YP Cai ; 1 ♀, Mt. Qian, Anshan , 5.VII.2010, coll. JY Liu and YP Cai ; 1 ♂, Bingyugou, Zhuanghe , 30.VII.2012, coll. DD Zhang and LJ Yang ; 1 ♀, Lianshanguan Forest Farm, 26. VI.2015; 2 ♀, Miaogou, Moshiyu Village , Xiaoshi, 29. VI.2015, slide No. RH16384 ; 2 ♀, Nongqiaoshanzhuang, Mt. Qian, Anshan , 225 m, 4.VIII.2016, coll. MJ Qi et al., slide No. RH16164; 1 ♀, Mt. Qian, Anshan , 245 m, 6.VIII.2016, coll. MJ Qi et al.; Jilin Province: 1 ♀, Huinan , 8.VII.1991 , slide No. RH16383 ; 1 ♀, Dongliao , 16.VII.1991 ; 1 ♂, Wanbao, Antu County, Yanbian , 24.VII.2012, coll. DD Zhang and LJ Yang , slide No. RH16350 ; 1 ♂, Duanmusonggou, Huashan, Linjiang , 26.VII.2012, coll. DD Zhang and LJ Yang , slide No. RH16345 ; 1 ♂, 1 ♀, Duanmusonggou, Huashanzhen, Linjiang , 26.VII.2012, coll. LJ Yang, XH Chen , slide Nos. RH16346 ♀, RH16349 ♂; Heilongjiang Province : 1 ♀, Laoshan, Mao’ershan, Shangzhi , 15.VII.2009, coll. WC Li and JY Liu ; Zhejiang Province: 1 ♂, Chanyuansi, Mt. Tianmu , 350 m, 15.VIII.1999, coll. HH Li et al., slide No. WYP05014 ; 1 ♀, Mt. Tianmu , 1140 m, 17.VIII.1999, coll. HH Li et al., slide No. WYP05248 ; 1 ♂, Sanmuping, Mt. Tianmu , 800 m, 19.VIII.1999, coll. HH Li et al., slide No. DYL00118 ; 1 ♀, Mt. Longtang , Qingliangfeng, 500 m, 21. V.2012, coll. LL Yang and ZG Zhang; 2 ♀, Mt. Jiulong , 400 m, 5.VIII.2014, coll. LL Yang and N Chen , slide No. RH16182 ; 2 ♀, Renchuan, Jinhua , 909 m, 30.VII.2015, coll. AH Yin et al.; 1 ♀, Simingshan National Nature Reserves, Ningbo , 822 m, 2.VIII.2016, coll. QY Wang et al., slide No. RH16191; 1 ♀, Laofoyan Village, Shuangxikou, Jiangkou , 424 m, 8.VIII.2016, coll. QY Wang et al.; 2 ♀, Linkeng Village, Yongjia , 387 m, 26.VIII.2016, coll. QY Wang et al.; Anhui Province: 1 ♂, Wenquan, Yuexi , 25.VIII.1995, coll. FX Hu , slide No. YP01326 ; 1 ♀, Jiuhuajie, Mt. Jiuhua , 9.VIII.2004, coll. JS Xu and JL Zhang ; Henan Province: 2 ♀, Shibanyan, Linzhou , 550 m, 21.VII.2006, coll. H Zhen and DH Kuang ; 1 ♀, Guanshan , Hui County, 550 m, 26.VII.2006, coll. DH Kuang and H Zhen ; Hubei Province: 1 ♂, Houhe , Wufeng County, 1000 m, 12.VII.1999, coll. HH Li et al., slide No. WYP05258 ; 1 ♂, Shayuan, Hefeng , 1260 m, 17.VII.1999, coll. H Li et al.; 1 ♂, 1 ♀, Mahe, Xianfeng , 1280 m, 25.VIII.1999, coll. HH Li et al., slide No. WYP05259 ♀; 3 ♀, Luotian , 24.VII.2012, coll. YP Wang , slide No. RH15192 ; 1 ♀, Maoba, Mt. Xingdou, Enshi , 780 m, 31.VII.2012, coll. J Zhang and XB Fu ; 2 ♀, Baimiaohe , 18.VIII.2012, coll. YL Xiao ; 1 ♂, 1 ♀, Taohuachong, Mt. Dabie , 661 m, 24. VI.2014, coll. ZH Liu and C Pan; 1 ♀, Taohuachong, Mt. Dabie , 590 m, 25. VI.2014, coll. LJ Xu; 5 ♂, 6 ♀, Qingtaiguan, Mt. Dabie , 590 m, 1–2.VII.2014, coll. LJ Xu , slide Nos. RH16423 ♂, RH16424 ♀; 3 ♂, 3 ♀, Qingtaiguan, Luotian , 524 m, 2.VII.2014, coll. ZH Liu and C Pan , slide Nos. RH16331 ♂, RH16332 ♀; 3 ♂, Qingtaiguan , Luotian County, 570 m, 1–4.VII.2014, coll. W Guan and MQ Yang , slide No. RH16121 ; Guangxi Zhuangzu Autonomous Region: 1 ♂, Jiman, Anchui, Rongshui , 650 m, 18.VII.2004, coll. JS Xu , slide No. WYP05015 ; 1 ♂, Peixiu, Rongshui , 30.VIII.2011, coll. DD Zhang , slide No. RH16335 ; 1 ♀, Langping, Cenwanglaoshan , 1450 m, 7.VIII.2014, coll. XL Wei and C Ran ; 1 ♀, Jiuniutang, Mt. Mao’er, Guilin , 1016 m, 24.VII.2015, coll. MJ Qi and SN Zhao , slide No. RH16193 ; Hainan Province: 1 ♂, Jianfengling Nature Reserves, Ledong , 770 m, 15.VII.2014, coll. PX Cong et al.; 1 ♀, Tianchi, Jianfengling , 787 m, 11.VII.2015, coll. QY Wang et al.; 2 ♀, Limushan Forest Park, 607 m, 24–25.VII.2014, coll. PX Cong et al.; 3 ♂, 3 ♀, Limushan Forest Park, 607 m, 13–17. V.2015, coll. PX Cong et al.; 1 ♀, Limushan Nature Reserves , 632 m, 1.VII.2015, coll. QY Wang et al.; 1 ♀, Limushan Forest Park, 607 m, 15.VII.2016, coll. X Bai et al.; 1 ♀, Diaoluoshan Nature Reserves , 922 m, 25. V.2015, coll. PX Cong et al.; 2 ♀, Wuzhishan Nature Reserves , 742 m, 18– 22. V.2015, coll. PX Cong et al.; 8 ♂, 5 ♀, Wuzhishan Nature Reserves , 738 m, 23–30.VII.2016, coll. X Bai et al.; 1 ♂, 3 ♀, Shuiman, Wuzhishan , 766 m, 31.VII–4.VIII.2016, coll. X Bai et al.; Sichuan Province: 1 ♂, Diaojiaolou, Mt. E’mei, 863 m, 23.VII.2011, coll. JB Cao , slide No. RH16349 ; 1 ♂, Nuoshuihe, Tongjiang , 700 m, 5.VII.2013, coll. GQ He and LJ Xu ; 1 ♀, Huagaoxi, Xuyong , 417 m, 25.VIII.2013, coll. LJ Xu ; 1 ♂, 1 ♀, Huagaoxi , 621 m, 5.IX.2014, coll. DD Xu and XL Wei ; Guizhou Province: 1 ♀, Lanjiang, Xishui , 500 m, 3. VI.2000, coll. YL Du, slide No. WYP05257 ; 1 ♂, Chishuisuoluo , 390 m, 22.IX.2000, coll. HL Yu , slide No. WYP05256 ; 1 ♂, 1 ♀, Lanjiang, Xishui , 500 m, 27–28.IX.2000, coll. HL Yu , slide Nos. WYP05254 ♂, WYP05262 ♀; 1 ♂, Dashahe, Daozhen , 1350 m, 24.VIII.2004 , YL Xiao, slide No. WYP05008 ; 2 ♂, 1 ♀, Taojiang , Leishan County, 27.VIII.2012, coll. JW Li and XH Chen , slide Nos. RH16337 ♂, RH16338 ♀; Yunnan Province : 4 ♀, Laiyanghe Forest Park, Pu’er , 26.VIII.2009, coll. XC Du , slide No. RH16436 ; 13 ♂, 3 ♀, Baihualing, Mt. Gaoligong, Baoshan , 1473 m, 29– 31.VII.2013, coll. SR Liu et al., slide Nos. RH15147 ♂, RH16190 ♂; 4 ♂, 4 ♀, Taiyanghe Nature Reserves , 1450 m, 30.VII–23.VIII.2014, coll. ZG Zhang , slide No RH15153 ♂; 3 ♂, Baihualing, Baoshan , 1473m, 5–6.VIII.2014, coll. KJ Teng et al.; 1 ♀, Xishuangbannan Botanical Garden , 565 m, 2. VI.2015, coll. MF Tao; 1 ♀, Hanlongzhai, Mangkuan, Baoshan , 1537 m, 20.VIII.2015, coll. XL Wei ; 1 ♂, 1 ♀, Taiyanghe Nature Reserves, Pu’er , 1450 m, 14.VIII.2016, coll. KJ Teng et al., slide Nos. RH16166 ♀, RH16169 ♂; Tibet Autonomous Region : 1 ♀, Motuo County, 1103 m, 8.VII.2013, coll. JW Li , slide No. RH16368 ; 1 ♀, Dexing , Motuo County, 835 m, 9.VII.2013, coll. JW Li ( SYSBM) ; Shaanxi Province: 1 ♂, Shangdong, Ziyang , 350 m, 21. V.1994, coll. J Zhou, slide No. WYP05249 ; 3 ♀, Xunyangba, Ningshan , 1360 m, 26. VI –1.VII.2007, coll. HL Yu, slide No. RH16184 ; 1 ♂, 1 ♀, Haopingsi, Yingtou , 1251 m, 17.VII.2012, coll. JW Li ( SYSBM) ; 1 ♀, Houzhenzi , Zhouzhi County, 1310 m, 19.VII.2012, coll. JW Li ( SYSBM) ; 1 ♂, 5 ♀, Huangbaiyuan, Taibai , 1295 m, 30. VI.2016, coll. JX Zhao and Y Fei; 1 ♂, Hengxi, Langao , 877 m, 13.VIII.2016, coll. WX Feng and WT Shi ; 4 ♀, Tiancheng Village, Langao , 577 m, 16–17.VIII.2016, coll. WX Feng and WT Shi , slide No. RH16426 ; 2 ♀, Tiancheng Village, Langao , 577 m, 16–17.VIII.2016, coll. JX Zhao and P Liu , slide No. RH16425 .

Adult ( Fig. 11 ) wingspan 21.0–26.0 mm.

Diagnosis. This species is characterized by the uncus with two curved and thorn-shaped lateral processes, distal process of the gnathos short column-shaped and serrated at apex, and sclerotized sub-rectangular median plate obtusely rounded on the outer margin in the male genitalia ( Fig. 4 ). It is similar to L. furciillatusa Wang, Chen & Wu, 2019 in wing pattern and the male genitalia, and the differences between the two species are stated in the diagnosis of L. furciillatusa .

Distribution. China (Anhui, Beijing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hebei, Heilongjiang, Henan, Hubei, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shanxi, Sichuan, Tianjin, Tibet, Yunnan, Zhejiang), India, Indonesia, Japan, Malaysia, Nepal ( Wang et al. 2017).

Figures

FIGURE 3–5. Morphology of Lista spp. 3. Adult of L. strumiformis sp. nov.; 4. Male genitalia of L. haraldusalis; 5. Female genitalia of L. haraldusalis. Scale bar: Fig. 3=2.0 mm; Figs 4, 5=0.5 mm.

FIGURES 6–13. Adults of Lista spp. 6. L. angustusa, male; 7. L. ficki, male; 8. L. furcellata sp. nov., holotype, male; 9. L. furciillatusa, male; 10. L. gilvasa, male; 11. L. haraldusalis, male; 12. L. insulsalis, female; 13. L. longifundamena, female. Scale bars: 2.0 mm.