Agyrtodini, Jeannel, 1936

Seago, Ainsley E., 2005, Male description and generic review of Agyrtolasia Szymczakowski, with key to genera of Agyrtodini (Coleoptera: Leiodidae: Camiarinae: Agyrtodini), Zootaxa 1103 (1103), pp. 1-15 : 8-12

Key to world genera of Agyrtodini

1. Lacinia expanded distally, (more so than galea) bearing a broad, articulated “sporebrush” or extremely dense patch of short spines ( Fig. 12a) ......................................... 2

– Lacinial apex with loose clump of spines or setae, but neither articulated nor substantially wider than galea ( Fig. 12b) ................................................................................. 7

2. Male with protarsus apparently triungulate with bifid claw ( Fig.13a); epistomal suture absent ( Fig. 13b); elytra with vivid “cloverleaf” pattern of pale spots (SE Australia) .. ................................................................................... Cholevomorpha Blackburn, 1891

– Male protarsus biungulate, without bifid claw; epistomal suture usually present; elytra solidly brown or without “cloverleaf” pattern .............................................................. 3

3. Labrum as long as wide, antennae slender, habitus slender­legged ( Fig.14b), pronotum constricted anteriorly; elytra tapering posteriorly. Mesosternum always distinctly carinate; male protarsus not broadly expanded ( New Zealand) (“ Zeagyrtes group”) ...... 4

– Labrum short and transverse; body compact and convex with relatively stout, robust legs and distinct antennal club ( Fig. 14a). Mesosternum carinate or not; male protarsus always expanded ................................................................................................... 6

4. Distinctive elytral color pattern of 3+ transverse chevron­shaped stripes. Epistomal suture present. Parameres bent medially or hooked at apex ...................................... 5

– Elytra medium­to­dark brown, marked evenly with white spots (on which pubescence is silver) or with indistinct posterior “wash” of pale tan. Epistomal suture absent. Parameres distally curved, but not hooked ........................ Zearagytodes Jeannel, 1936

5. Elytra with faint “chevron” stripes; pronotum with dark medial stripe and often with paired dark spots at the lateral margins. Median lobe short and truncate, not tapering along length. Aedeagal parameres distinctly hooked, bearing two small apical setae ( Fig. 15). Labrum elongate ...................................................... Zeagyrtes Broun, 1917

– Elytra with three dark fasciae. Median lobe short and triangular in shape, tapering evenly from base to apex. Parameres slender, bent medially but not hooked, without short apical setae. Labrum transverse .................... Zeagyrtoma Szymczakowsi, 1966

6. Elytra with “S”­shaped, tricolor pattern ( Fig. 16) and conspicious, long, shaggy, dorsal vestiture; elytral lateral striation indistinct. Body length <1.7 mm (E Australia) ......... ................................................................................. Agyrtolasia Szymczakowski, 1973

– Elytra solid brown, without tricolor pattern; dorsal vestiture short, even, and dense; elytra with conspicuous lateral striation. Body length> 2 mm ( Australia, New Zealand, Chile) ..................................................................... Agyrtodes Portevin, 1907

7. Mesosternal carina acute, conspicuously pointed. Hind tibia curved along length. Large for agyrtodines, body length 3–3.5 mm ( Chile) .......... New Genus “F” (in prep)

– Mesosternal carina absent, or low and rounded (if present, then body length less than 2 mm). Hind tibiae not curved; body length typically <3 mm ...................................... 8

8. Hind wings absent; body length 2 mm or less ............................................................ 9

– Hind wings present; body length almost always> 2 mm .......................................... 11

9. Protibia bent medially and expanded apically ( Fig.17); smoothly rounded mesosternal carina present ( New Zealand) ............................. Chelagyrtodes Szymczakowsi, 1973

– Protibia straight, neither bent nor apically expanded; mesosternal carina absent or extremely low ............................................................................................................ 10

10. Maxilla with apical palpomere expanded ( Fig. 18a); epistomal suture with stem. Eyes reduced. Elytra with faint transverse striae. Tarsal formula 4­4­5 (Tasmania) ............ .................................................................................................. New Genus 1 (in prep.)

– Maxilla with penultimate palpomere expanded, substantially more so than apical palpomere ( Fig. 18b). Epistomal suture without stem. Elongate, non­striate elytra. Tarsal formula 5­5­5 ( New Zealand) ............................................ New Genus 2. (in prep)

11. Male sternite VII with paired hook­like processes ( Fig. 19a); large, curved spine on ventral face of mesofemur ( Fig. 19b) (S Chile, Argentina) ........................................... ............................................................................................ Dasypelates Portevin, 1907

– Male sternite VII and mesofemur without hooks or spurs ......................................... 12

12. Aedeagal parameres expanded and flattened apically, with paddle­like or spoon­like shape (e.g., Fig. 21) ................................................................................................... 14

– Aedeagal parameres slender, without notable apical expansion ( Figs. 20a, b) (S Africa)......................................................................................................................... 13

13. Parameres with strong median bend; median lobe short, broad, and apically truncate ( Fig. 20a). Elytra with transverse striae ............................... Dictydiella Jeannel, 1936

– Parameres straight or slightly curved; median lobe with “nipple” or gentle hook at apex ( Fig. 20b). Elytra without transverse striae ................. Afropelates Jeannel, 1964

14. Aedeagal parameres broadly expanded, with spoon­like flattened apex bearing at least one row of long setae on interior face ( Fig. 21); male mesotarsus with segments 1–2 dilated. Male sternite VI cleft; body length> 3.5 mm ( Australia) ................................. .............................................................................. Paragyrtodes Szymczakowski, 1966

– Parameres expanded at apex but typically blade­like in shape, often sharply truncate (never rounded); male mesotarsus with first segment only dilated (except Eupelates , in which segments 1–2 are expanded). Male sternite VI whole; body length typically <3 mm ( Chile and Argentina) ......................................................................................... 15

15. Penultimate maxillary palpomere expanded, longer than terminal palpomere ( Fig. 22); pronotum bearing six–ten extremely deep punctures along posterior margin. Epistomal suture without stem ( Fig. 24a). Femora without distinct pale stripe. Pronotum and elytra frequently tuberculate and without transverse striae ..................................... ............................................................................................... Ragytodina Jeannel, 1957

– Terminal maxillary palpomere longer than other segments; elytra and pronotum without deep punctures, never tuberculate. Epistomal suture with stem ( Fig. 24b). All femora with pale median stripes ( Fig. 23). Elytra smooth, with transverse striolations .................................................................................................................................... 16

16. No conspicuous color pattern on dorsum. Aedeagal parameres broad and flattened, extending well past apex of median lobe ( Fig. 25a) ............. Eupelates Portevin, 1907

– Elytra with or without vivid dorsal color pattern. Aedeagal parameres slender, only slightly longer than median lobe ( Fig. 25b) ...................... Chiliopelates Jeannel, 1957