Stalisyne lutetiorum, Szwedo, Jacek, Bourgoin, Thierry & Lefebvre, Fabrice, 2006

Szwedo, Jacek, Bourgoin, Thierry & Lefebvre, Fabrice, 2006, New Mnemosynini taxa (Hemiptera, Fulgoromorpha: Cixiidae) from the Palaeogene of France with notes on their early association with host plants, Zootaxa 1122, pp. 25-45 : 27-31

Stalisyne lutetiorum sp. nov.

( Figs. 1 –12)

MATERIAL: Holotype, female. Specimen MNHN PA 2427, 1/2; collection “Langlois­ Meurinne / De Ploëg”, mounted in Canada balsam, deposited in Muséum national d’Histoire naturelle, Paris [ MNHN]. State of preservation: anterior part of body partly hardly visible, covered with milky veil, left tegmen incomplete, only basal part and claval portion preserved. Facial portion partly covered with milky veil. Fore and mid femora, tibiae and tarsi missing.

TYPE LOCALITY: Farm Le Quesnoy, Chevrière, region of Creil, Oise department (Northern France).

STRATIGRAPHIC POSITION: Lowermost Eocene, circa 53 Ma, Sparnacian, level MP7, of the mammal fauna of Dormaal ( Feugueur 1963; De Ploëg et al. 1998; Nel et al. 1999, Nel et al. 2004; Cavagnetto 2000).

ETYMOLOGY: Named after Gallo­Celtic tribe inhabiting environs of Paris—the Lutetes.

DIAGNOSIS: Vertex narrow at base, angulately incised, its anterior margin slightly before the level of compound eyes (in M. cubana vertex wide at base, with straight angle incision, almost straight in middle portion, anterior margin of vertex not exceeding level of anterior angles of compound eyes). Postocular carinae distinct (not distinct in M. cubana ), placed more caudad to the compound eyes than in Stalisyne veromanduiorum . Mesonotum with five distinct carinae (carinae very weak in M. cubana , three carinae in Autrimpus sambiorum ). basicostal field short and wide (no distinct basicostal field in M. cubana ); first M forking apicad of ScRA and CuA forkings (M forked basad of ScRA forking in M. cubana ); RA with three terminalia (two terminalia in M. cubana and A. sambiorum ); RP with four terminalia (three terminalia in M. cubana and A. sambiorum ); CuP and Pcu not connected with transverse veinlet at tegmen base (such veinlet present in M. cubana , A. sambiorum and Mnasthaia arverniorum ).

DESCRIPTION: General coloration in amber ochraceous, head, pronotum and mesonotum lighter, tegmina with dark brown, elongate spot at the base of clavus, incomplete transverse band in the median portion, not reaching costal margin and irregular spots in apical cells. Tegmen regularly covered with short setae, including veins.

Total length (female) 12.3 mm. Head with compound eyes (2.2 mm) wider than long, Anterior margin of head in dorsal view rounded. Vertex with lateral margins distinctly elevate, posterior margin angulately incised; vertex at base (0.56 mm) slightly narrower than long in mid line (0.6 mm), narrowing apicad. Face very long (4.48 mm in mid line), with median carina and median ocellus. Two longitudinal carinae at the margin of frons and vertex, two transverse carinae, the upper delimiting vertex, and the lower uniting longitudinal carinae with lateral carinae present. Frons 1.12 mm long in mid line, 1.12 mm wide at widest point, at the level of antennae, with disc almost flat; Frontoclypeal suture distinct, arcuate; Postclypeus 1.68 long in mid line, convex, with distinct lateral carinae; Anteclypeus 1.48 mm long in mid line, with median carina; Labium long; Rostrum very long (6.28 mm), reaching far beyond end of abdomen and tip of ovipositor, with anteapical segment almost two times as long as apical one; Bases of antennae sunk in lateral concavities of the head capsule, basal segment short, the second rounded, covered with sensillae, arista relatively long; Lateral ocelli distinct, placed just below the level of lower margin of the compound eye, close to the lateral carinae of frons.

Pronotum 3.16 mm wide, wider than head with compound eyes, 0.6 mm long in mid line, about as long in mid line as in lateral portion, with median carina and postocular carinae distinct, posterior margin incised at wide angle; Tegula with median carina, about twice as wide as long.

Mesonotum slightly longer (2.72 mm) than wide (2.6 mm), with concave disc and with five longitudinal carinae, scutellum distinct, flat.

Tegmen 10 mm long, 3.04 mm wide at widest point, clavus long, reaching 2/3 of tegmen length, stigma distinct, elongately triangular; costal margin mildly curved at base, with distinct basicostal field, almost straight in the median portion, angulately rounded in apical portion of tegmen; basal cell about twice as long as wide, ScR forked near the base, ScRA1 reaching the stigma, RA with three terminalia, RP forked basad of the level of nodal line, with four terminalia; M simple at base, forked slightly apicad of ScR and Cu forkings, with five terminalia, CuA forked apicad of ScR forking, at the half distance between ScR forking and PcuA1 junction, CuA with two terminalia, CuA1 branch distinctly curved; first row of transverse veinlets uniting RP and M and M and CuA1 basad of nodal line, transverse veinlets of apical line not very regular, not forming distinct line; claval veins Pcu and A1 united apicad of half of length of clavus.

Hind wing with venation typical of Mnemosyne spp., RP with two terminalia, M with four terminalia, CuA with three terminalia, transverse veinlet r­m long curved apicad.

Hind femur 2.12 mm long, hind tibia 3.84 mm long, quardrangular in cross section, with two distinct lateral spines, the first near the base and the second, bigger just above half of tibiaís length, six apical spines organized in formula 1 ­ 2 ­ 3; hind tarsus 3.28 mm long, basitarsomere longer than combined length of mid and apical tarsomeres, 1.96 mm long with 5 apical spines, mid tarsomere shorter, 1.04 mm, with 6 apical spines, apical tarsomere, 0.72 mm long with small tarsal claws and pulvilla.

Female abdomen 4.36 mm wide, with distinct trichobothrial pits in lateral portion of sternites, pygofer capsule oblique in lateral view, concave, with wax pores; ovipositor elongate, not exceeding length of anal tube, gonapophysis VIII shortened, 1 mm long, with tip covered with a few setae, gonapophysis IX 2.04 mm long, with short, stout setae on internal margin, gonoplac foliaceous, 2.32 mm long, 0.48 mm wide, with margins covered with long setae.

Female anal tube 1.16 mm long, slightly flattened, anal style 1 mm long.

Figures

FIGURES 1 – 4. Stalisyne lutetiorum gen. et sp. nov. Fig. 1. Anterior part of body in dorsal view; Fig. 2. Face in ventrolateral view; Fig. 3. Face and rostrum in ventrolateral view; Fig. 4. Anterior part of body in right lateral view; Scale bar: 1 mm.