Tingiometra secunda, Golub & Heiss, 2020

Golub, Viktor B. & Heiss, Ernst, 2020, Three new species of the lacebug genus Tingiometra from Cretaceous Burmese amber (Hemiptera: Heteroptera: Tingidae: Tingiometrinae), Zootaxa 4767 (4), pp. 531-542 : 536-538

Tingiometra secunda sp. nov.

Fig. 11

Type material. Holotype female, Fig. 11 , BUB-TING-09 (16/19-2), CEHI.

Diagnosis. Tingiometra secunda sp. nov. differs from Tingiometra burmanica by shorter and wider body (1.79 times as long as wide; vs 2.5–3.0 times as long as wide), strongly rounded lateral margins of hemelytra (lateral margins of hemelytra are rather weakly rounded), much wider discoidal area including five large cells with 17 rows of small areolae at the widest part (vs discoidal area with three large cells with 7–8 rows of areolae at widest part), shorter antennae (vs length of antennal segments I, II, III, IV is 0.13, 0.70, 0.25, 0.48), wider pronotum 1.26 times as wide as long (vs 1.33 as long) wider scutellum 1.13 as wide as long (vs 1.5 times as long as wide).

This new species is closely related to T. pankowskii sp. nov. It differs by wider body (1.79 times as long as wide; vs 2.09–2.42 times) with very rounded lateral margins of hemelytra, wider subcostal area of hemelytra having 10–11 rows of areolae at the widest part (vs 6–7 rows of areolae), narrower discoidal area with 17 rows of areolae at the widest part (cf.: 20–22 rows of areolae in holotype; 18–22 rows in paratypes). Differences to T. yuripopovi sp. nov. are mentioned in the description of the latter.

Description of holotype. Body brown, elongate oval, 1.79 times as long as width, hemelytra and paranota finely areolate. very fine punctuation on anterior part of pronotum.

Head very long, 1.78 times as long from apex of clypeus to posterior margin of eyes as width of head across eyes, and 2.55 as long, including frons, as width of head, in very small punctuation. Frons and vertex horizontal. Clypeus long, with crest on its surface. Eyes relatively small, not much protruding on the sides of the head. Vertex and frons flat, horizontal. Ocelli absent. Margins of postocular lobes of head slightly rounded. Antenniferous tubercles short, slightly directed outward. Antennae long and very thin except thickened segment I, about 4.7 times as long as width of head across of eyes; segment I shortest, II longest, III and IV shorter and thinner than II.

Pronotum narrow anteriorly and strongly widening posteriorly, 1.26 as wide as long, its anterior margin not wider than the posterior margin of head, distinctly sinuate; posterior pronotal margin rather strongly sinuate, lateral margins slightly sinuate, anterolateral angles almost sharpened, posterolateral angles narrowly rounded. Anterior part of pronotum with a insignificantly raised ring bearing a single row of small areolae. Pronotal disk almost flat, with longitudinal median low carina without distinct areolae, consisting of a longer and higher posterior part and a shorter and lower anterior one, two part of carina separated by transverse impression; posterior part of carina about 2 times as long as the anterior one. Paranota rather narrow, distinctly reflexed, in the form of very thin plates, with one row of small areolae in the posterior part and with few indistinct areolae of the second row in the anterior part. Pronotal triangular posterior process absent, therefore scutellum and clavus uncovered and plainly visible.

Scutellum large, triangular, its apex noticeably enlarged and straightly cut.

Hemelytra submacropterous, overlapping by their sutural areas only, slightly protruding posteriorly apex of abdomen. Corium divided into costal, subcostal, discoidal and sutural areas all small angular and rounded areolae. Stenocostal lamina absent. Costal area very narrow, carinate, with one row of extremely small areolae in the base only and without distinct areolae at remaining part. Subcostal, discoidal, and sutural areas distinctly separated by carinate veins. Subcostal area declivous, with 10–11 rows of areolae at widest part, divided by 3 transverse carinate veins. Discoidal area very wide, delimited by vein R exteriorly and by Cu interiorly, with 17 irregular rows of areolae at widest part, divided by one longitudinal curved carinate vein M and by four short transverse carinate veins located obliquely to longitudinal axis of the body; three short transverse veins connect M with Cu and one vein connects M with R; thus discoidal area is divided into five large cells: external basal and apical cells adjoining to subcostal area and three internal cells (basal, medial and apical) located closer to the longitudinal axis of the body and adjacent to clavus and sutural area of hemelytron; external basal cell is the largest. Every large cell of discoidal area contains many small rounded areolae. Sutural area narrow and with one row of areolae at base, with two rows at middle part of its length and with three rows of areolae at the apex of hemelytron. Clavus with 5 rows of areolae at widest part, commissure longer than scutellum. The total surface of the discoidal and sutural areas and clavus nearly flat.

Venter. Bucculae very short (see laterally), occupying only apex of head ventrally, closed anteriorly, surrounding base of labium like a cover. Labium 4-segmented, very long, its apex reaching posterior margin of hind coxae, first segment thickest. Sternal labial thoracic plates absent. Triangular median projection of the posterior margin of mesosternum reaches middle of hind coxae. External groove of scent gland apparatus not discernible. Abdominal sternites II and III completely fused, sternites IV–VIII separated by transverse furrows, sternite VII with median and 2 (1+1) lateral processes pointed at their apices. Lateral processes are curved towards the longitudinal axis of the body. Ovipositor with a half-tube shape.

Legs. Long and slender, femora and tibiae cylindrical; tarsi two-segmented, pulvilli not discernible.

Measurements of holotype (in mm). Body length from clypeus to apex of hemelytra 3.13; body width of folded wings in repose 1.75; head length from apex of clypeus to posterior margin of eyes 0.58; head length including postocular portion 0.83; length of anteocular portion 0.48, length of eye 0.10, length of postocular portion 0.25; head width across eyes 0.33; width of anteocular part of head 0.20, width of postocular part 0.30; length of antenna 1.54; length of antennal segments (I, II, III, IV) 0.15, 0.68, 0.28, 0.43; pronotum length 0.50; pronotum width 0.63; length of anterior portion of pronotal carina 0.225; length of its posterior portion 0.125; scutellum length 0.38; scutellum width 0.43; hemelytra length 1.88; length of commissure between clavus and corium 0.75; length of foreleg segments: femur 0.75, tibia 1.30; middle leg: femur 0.78, tibia 1.38, tarsus 0.15; hind leg: femur 1.20, tibia 1.75.

Etymology: Refers to the fact, that it is the second new species assigned to Tingiometra from <secundus> (Latin) second.

Distribution. Burmese amber, northern Myanmar.

Figures

FIGURES 9–12. 9,10 Tingiometra pankowskii sp. nov., 9, paratype male (?) (7303-1), dorsal view, 10, paratype male (?) (7303-1), ventral view; 11, Tingiometra secunda sp. nov., holotype female (16/19-2), dorsal view; 12, Tingiometra yuripopovi sp. nov., holotype female (18/25), dorsal view. Scales 1mm.