Tapinopterus (Molopsis) wiedemanni ( Chaudoir, 1850 )

Lohaj, Roman, Guéorguiev, Borislav, Dubault, Gérard & Lassalle, Bernard, 2012, Revision of Molopsis Schatzmayr (Coleoptera: Carabidae: Pterostichini), with descriptions of four new species, Zootaxa 3185, pp. 1-35 : 26-27

Tapinopterus (Molopsis) wiedemanni ( Chaudoir, 1850)

(Figs. 21–23, 40–44, 75–79, 91–94, 106–109, 121–123, 135–137, 143, 150, 153, 161–162, 166)

Type locality. Nord de l´Anatolie.

Taxonomic decision. Tapinopterus (Molopsis) wiedemanni ( Chaudoir, 1850) , combinatio novum of Feronia (Molops) wiedemanni Chaudoir, 1850: 145 (nec Fairmaire, 1866: 252).

References

Feronia (Molops) wiedemanni Chaudoir: Chaudoir, 1850 : 145; Marseul, 1882: 51.

Feronia wiedemanni Chaudoir : Kraatz, 1875: 419.

Feronia (Steropus) wiedemanni Chaudoir : Marseul, 1880: 313.

Molops wiedemanni Chaudoir : Ganglbauer, 1889: 52; Bousquet, 2003: 475.

Platysma (Molops) wiedemanni Chaudoir : Jakobson, 1907: 355.

Material examined. 103 specimens.

Type material. Two syntypes (3 Ƥ) of Feronia wiedemanni (lectotype male and paralectotype female by present designation), well preserved, “ Wiedemanni Chaud. Anatolie C. Falderm.” (h label pinned on the bottom of box)/ LECTOTYPE and PARALECTOTYPE Feronia (Molops) wiedemanni Chaudoir D.Dubault désign. 2003 (red p)/ Tapinopterus (Molopsis) wiedemanni G.Dubault détermin. (p) (Fig. 21) ( MNHN, “Collection Chaudoir”, box no 216 “ Argutor ”).

Other material examined. Abant: 29 33 ƤƤ, “Aband 1.300 m. Turkey J. y S. Klapperich.” ( MNHN); 1 3, TR.bor.occ. ABANT lake, 5.6.1992, V.Skoupý lgt. (cVS); 1 3 1 Ƥ, TURKEY occ. Abant Gölü env. (35 km SW of BOLU), 1993-06-21, Kuboň lgt. (cGD); 2 33, Türkie bor., Abant, 9.– 10.5.1994, A.Trmal lgt. (cRL); 1 3 3 ƤƤ, TURKEY (Bolu), Elmacik dagi, Golu lake, 10.V.1994, Major (cDW); 3 33 1 Ƥ, N TURKEY, Abant env., 5.1995, F.Moravec lgt. (cRL) (Fig. 22); 1 3, TR-prov. Bolu, Abant gölü, 1200 m, 4.6.1996, Z.Malinka lgt. (cVS); 1 3 1 Ƥ, TURKEY, Abant Golu, 30 km W Bolu, 4.5.1997, leg J.Bašta (cDW); 1 3, TURKEY, Abant Golu, W Bolu, 8.6.1998, lgt. E.& P. Hajdaj (cDW); 1 Ƥ, TR vill. Bolu, Abant Gölü, 23.– 24.5.2000, Josef Mertlik lgt. (cVS).

Aladaglari: 15 33 8 ƤƤ, Anatolia bor., Aladaġ ca 20 km südl. Bolu, 1400–1600 m, 27.IV.1981, Heinz leg. ( MCSNM, cWH, cBL, cRL).

Köroġlu daġlari: 1 Ƥ, Anatolia bor, Tasliyayla, ca 21 km ne Seben (Bolu), 1500–1600 m, 27.IV.1984, W. Heinz lgt. (cWH); 13 33 9 ƤƤ, Kartalkaya, 1800 Bolu Turquie, 19.05.94, G. Dubault (cGD, cRL, cBL).

Sündiren daġlari: 7 33 1 Ƥ, Anatolia centr., Wald n. Mihaliççik, 1500–1600m, Heinz leg. (cBL, cGD) (Fig. 23).

Diagnosis. BL 11.3–15.8 mm.

Male genitalia. Median lobe of aedeagus in dorsal view long, with long, thin, and slightly curved to the left apical lamella, which formed more or less distinct transverse disc in lateral position (Figs. 40–44, 75–79); apical orifice deflected to left lateral position; basal bulb shorter than apical, with basal orifice concave, apical part long, copulatory sclerite curved, with widened medial part, situated in paramedial position, apex of lobe bent downwards, but reflexed before end. Right paramere relatively short, strongly curved and slightly constricted medially, with thinner apical and thicker basal part (Figs. 91–94, 106–109). Left paramere with long transverse apophysis and long, oblique paramedial apophysis (Figs. 121–123, 135–137).

Female genitalia. Tergum VIII relatively long, with convex, subpyramidal distal margin, straight margin and short “legs” in proximal position (Fig. 143). Sternum VIII unified, in proximal position with two closed “holes” and without “legs” (Fig. 150). Syntergum IX+X long, narrow, bearing pair of small, articulated, distant each other ovipositors (Fig. 153). Ovipositor with valvifer and stylus (Figs. 161–162); valvifer with twin-peaked internal margin, convex external margin, and process across at distal part; stylus consist of one, basal, subelongate stylomere (apical stylomere disappeared), having strongly incised internal margin. Spermatheca with seminal canal and receptaculum discrete; receptaculum shorter than seminal canal, slightly curved apically (Fig. 166); spermathecal gland elongate, spermathecal canal short, inserted at junction of receptaculum and seminal canal.

Taxonomic notes. This species was described from three males and one female as Feronia (Molops) wiedemanni from the north of Anatolia ( Chaudoir, 1850: 145–146). Later, Gemminger & Harold (1868: 333) placed it under Molops , but Chaudoir (1876: 348) specified that it did not belongs to that genus. Kraatz (1875: 419) is first to indicate that F. molopina and F. wiedemanni were close relatives. Marseul (1882) put F. wiedemanni Chaudoir among the Molops species. For the lack of basolateral setigerous punctures and presence of single basal impression of pronotum, Ganglbauer (1889: 52) supposed that this taxon belonged to Tapinopterus . It seems that since the work of Jakobson (1907) this taxon disappeared from the literature. It appears again only recently, when F. wiedemanni Chaudoir, 1850 was considered either as a representative of Molops ( Mlynář, 1977) or was placed among the species “incertae sedis” of the same genus ( Bousquet, 2003: 475).

In fact, no real explanation about the systematic position of this species has been taken after the above note of Chaudoir (1876). After prolonged search for the Chaudoirś types in MNHN by one of us (GD), two syntypes of Feronia wiedemanni were eventually found. Their study revealed that both belong to a valid species of Tapinopterus , and not to Molops .

Distribution. Mountain ranges around Bolu: Abant, surrounding of the Abant Lake, Aladaġlari, Köroġlu daġlari, Sündiren daġlari, NW Turkey.