HYDROPHILIDAE, Latreille, 1802

Hubbard, Todd, 2017, Aquatic Coleoptera (Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae) of Iowa: Distributional Records and Notes, The Coleopterists Bulletin 71 (3), pp. 471-505 : 497-503

HYDROPHILIDAE

Anacaena lutescens (Stephens) . Allamakee: Hecker Cr., Postville, TMM, 7/24/07; Buchanan: Pine Cr., Quasqueton, CTH, 8/9/11; Clayton: Unnamed trib. Silver Cr., Luana, CTH, 8/7/06; Delaware: Bear Cr., Dyersville, CTH, 7/27/ 11; Fayette: Unnamed trib. to Little Turkey R., St. Lucas, CTH, 8/12/10; Floyd: Cedar R., Charles City, Mark Johnston, 9/9/08; Franklin: Bailey Cr., Sheffield, CTH, 7/20/11; Kossuth: Buffalo Cr., Titonka, KLS, 8/23/00; Mahaska: Buck Cr., Barnes City, CTH, 7/26/12; Mitchell: Cedar R., St. Ansgar, CTH, 7/5/06; Winneshiek: South Pine Cr., Sattre, CTH, 8/4/10. Distribution: Western USA, extending east to the upper Midwest.

Berosus aculeatus LeConte. Clayton : Peck Cr., Osterdock, CTH, 7/25/11; Mitchell: Cedar R., St. Ansgar, CTH, 7/5/2006. Distribution: Eastern USA, extending west to Minnesota in the north, Iowa, and Texas in the south .

Berosus fraternus LeConte. Black Hawk : Cedar Hills Sand Prairie ( UV light), ELF, 6/19/12; Butler: Palmer Cr., Greene , CTH, 5/29/2007 ; Dubuque: North Fk. Maquoketa R ., Dyersville , CTH , 7/25/05; Hancock: East Branch Iowa R ., Goodell, JML, 10/17/00; Henry: Skunk R ., Oakland Mills , CTH , 9/24/08; Jackson: North Fk. Maquoketa R ., Maquoketa , CTH , 9/13/12; Linn: Cedar R ., Cedar Rapids , CTH , 8/24/15; Muscatine: Mud Cr., Wilton , CTH , 9/15/03. Distribution: Transcontinental in the USA.

Berosus infuscatus LeConte. Clinton : Brophy Cr., McCausland, CTH, 8/14/12; Henry: Big Cedar Cr., Oakland Mills, CTH, 8/29/12; Jackson: Tetes Des Morts Cr., St. Donatus, CTH, 8/20/14; Johnson: Iowa R., Iowa City, CTH, 9/18/13; Linn: Cedar R., Cedar Rapids, CTH, 9/19/13; Louisa: Honey Cr., Conesville, CTH, 8/24/04; Mahaska: Muchakinock Cr., Eddyville, CTH, 9/ 2/11; Muscatine: Pike Run, West Liberty, CTH, 7/21/05; Polk: Des Moines R., Des Moines, KLS, 9/24/13; Washington: English R., Riverside, CTH, 9/24/02. Distribution: Transcontinental in the USA.

Berosus pantherinus LeConte. Allamakee : Little Paint Cr., Harpers Ferry, CTH, 8/27/14; Cedar: Cedar R., West Branch, CTH, 9/12/06; Clinton: Brophy Cr., McCausland, CTH, 8/14/12; Johnson: Old Mans Cr., Hills, CTH, 9/18/13; Linn: Cedar R., Cedar Rapids, CTH, 9/19/13; Lucas: Wolf Cr., Chariton, TMM, 9/12/12; Muscatine: Cedar R., Conesville, CTH, 8/29/ 12; Polk: Des Moines R., Des Moines, CTH, 8/ 27/12; Wayne: Jordan Cr., Millerton, TMM, 8/ 24/12. Distribution: Eastern USA, extending west to Minnesota in the north, Nebraska, Missouri, and Texas in the south.

Berosus peregrinus (Herbst) . Appanoose: Chariton R., Rathbun, CTH, 8/17/04; Buchanan: Pine Cr., Quasqueton, CTH, 8/9/11; Butler: Shell Rock R., Shell Rock, CTH, 9/8/14; Cedar: Sugar Cr., Wilton, CTH, 8/23/01; Clayton: Silver Cr., Monona, CTH, 8/7/06; Clinton: Brophy Cr., McCausland, CTH, 10/2/14; Dallas: South Raccoon R., Redfield, TMM, 8/8/12; Davis: Soap Cr., Drakesville, CTH, 8/16/04; Decatur: Weldon R., Woodland, TMM, 8/8/ 13; Delaware: Bear Cr., Dyersville, CTH, 7/ 27/11; Des Moines: Flint Cr., Danville, CTH, 7/15/03; Dubuque: Little Maquoketa R., Dubuque, CTH, 8/19/08; Floyd: Cedar R., Charles City, CTH, 9/9/08; Franklin: Beaver Cr., Ackley, CTH, 8/22/13; Guthrie: Middle Raccoon R., Coon Rapids, MWB, 6/16/07; Hamilton: Boone R., Stratford, TMM, 8/28/ 14; Henry: Big Cr., New London, CTH, 8/30/ 05; Jackson: Farmers Cr., Fulton, CTH, 8/8/07; Johnson: ( Wickham 1911); Kossuth: Lotts Cr., West Bend, MWB, 9/25/02; Linn: Indian Cr., Cedar Rapids, CTH, 8/27/13; Louisa: Honey Cr., Conesville, CTH, 8/21/12; Lucas: Wolf Cr., Chariton, TMM, 9/12/12; Madison: Middle R., Wintereset, TMM, 7/16/12; Marion: English Cr., Knoxville, CTH, 7/18/05; Mitchell: Deer Cr., Carpenter, CTH, 8/21/13; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; Polk: Des Moines R., Des Moines, TMM, 8/27/12; Sioux: Sixmile Cr., Hawarden, KBK, 8/3/00; Story: Ballard Cr., Cambridge, TMM, 8/2/13; Taylor: Honey Cr., Bedford, TMM, 8/18/04; Van Buren: Fox R., Mt. Sterling, CTH, 7/16/13; Wayne: Jordan Cr., Millerton, TMM, 9/12/12; Webster: Des Moines R., Ft. Dodge, TMM, 9/29/14; Winnebago: Buffalo Cr., Forest City, MWB, 9/16/06; Woodbury: West Fk. Little Sioux R.,

Bronson, TMM, 8/9/05. Distribution: Transcontinental in the USA.

Berosus sayi Hansen. Allamakee : Little Paint Cr., Harpers Ferry, CTH, 8/16/07; Appanoose: Chariton R., Centerville, TMM, 7/20/12; Black Hawk: Cedar Hills Sand Prairie (UV light), ELF, 6/19/12; Clay: Big Muddy Cr., Spencer, TMM, 8/27/13; Clayton: Hewitt Cr., Volga, CTH, 8/4/ 14; Clinton: Brophy Cr., McCausland, CTH, 9/ 30/13; Davis: North Fabius R., Moulton, CTH, 7/17/06; Dickinson: Little Sioux R., Lake Park, TMM, 8/11/14; Dubuque: Middle Fk. Maquoketa R., Rickardsville, CTH, 8/10/04; Fremont: Waubonsie State Park (UV light), ELF, 8/15/12; Kossuth: Lotts Cr., West Bend, MWB, 9/25/ 02; Mitchell: Cedar R., St. Ansgar, CTH, 7/5/ 06; Story: Dye Cr., Colo, TMM, 5/27/14; Wayne: Jordan Cr., Millerton, TMM, 9/12/ 12; Winneshiek: South Pine Cr., Sattre, CTH, 8/14/07; Wright: East Br. Iowa R., Belmond, TMM, 7/23/12. Distribution: Transcontinental in the USA.

Chaetarthria atra (LeConte) . Delaware: Maquoketa R., Dundee, MWB, 6/24/01. Distribution: Eastern USA, extending west to Missouri and Iowa.

Chaetarthria pallida (LeConte) . Johnson: ( Wickham 1911). Distribution: Primarily eastern USA, with isolated records in the west.

Cymbiodyta blanchardi Horn. Allamakee : French Cr., Churchtown, CTH, 9/16/04; Clayton: North Cedar Cr., Clayton, CTH, 9/2/15; Clinton: Elk R., Teeds Grove, CTH, 8/12/13; Dubuque: White Pine Hollow Cr., Luxemburg, CTH, 9/9/ 04; Fayette: Unnamed trib. to Volga Lake outlet, Fayette, CTH, 8/4/15; Mitchell: Burr Oak Cr., Osage, CTH, 8/3/11; Winneshiek: UV light, KR, 11/3/01. Distribution: Eastern USA, extending west to Missouri and Iowa.

Cymbiodyta chamberlaini Smetana. Allamakee : Suttle Cr., Monona, CTH, 8/17/11; Clayton: Unnamed trib. to Volga R., Volga, CTH, 9/21/15; Delaware: Twin Springs Cr., Colesburg, CTH, 9/ 17/07; Dubuque: Catfish Cr., Dubuque, CTH, 7/ 28/95; Fayette: Unnamed trib. to Little Turkey R., St. Lucas, CTH, 9/28/11; Jackson: Storybook Hollow Cr., Bellevue, CTH, 8/29/07; Winneshiek: Middle Bear Cr., Highlandville, CTH, 7/17/07. Distribution: Eastern USA, extending west to Wisconsin in the north, Iowa, and Kansas in the south.

Cymbiodyta minima Notman. (Downie and Arnett 1996) . Distribution: Northeastern USA, with Iowa forming part of the western border.

Cymbiodyta semistriata (Zimmermann) . Davis: Burr Oak Cr., Troy, CTH, 6/1/15; Van Buren: Town Branch, Pittsburg, CTH, 7/29/14. Distribution: Eastern USA, extending west to Wisconsin in the north and Colorado.

Cymbiodyta toddi Spangler. Allamakee : Little Paint Cr., Harpers Ferry, CTH, 8/16/07; Black Hawk: Wolf Cr., La Porte City, CTH, 8/19/04; Buena Vista: Unnamed trib. to Maple Cr., Aurelia, TMM, 8/27/13; Butler: South Beaver Cr., Parkersburg, CTH, 7/30/96; Chickasaw: Wapsipinicon R., Twin Ponds Park, CTH, 9/29/ 14; Clay: Unnamed trib. to Little Sioux R., Peterson, TMM, 8/13/13; Clayton: Bixby Springs, Edgewood, MWB, 9/9/04; Dubuque: Catfish Cr., Dubuque, CTH, 8/1/07; Fayette: Brush Cr., Arlington, CTH, 9/10/04; Franklin: Boylan Cr., Aredale, CTH, 8/11/14; Grundy: South Beaver Cr., Parkersburg, CTH, 7/30/96, 9/ 3/08; Guthrie: Middle Raccoon R., Coon Rapids, MWB, 6/16/07; Hardin: South Fk. Iowa R., Iowa Falls, TMM, 8/2/12; Howard: North Branch Turkey R., Lime Springs, CTH, 8/5/15; Jasper: Walnut Cr., Prairie City, CTH, 8/21/95; Johnson: Muddy Cr., North Liberty, CTH, 10/2/ 06; Keokuk: North Skunk R., Sigourney, CTH, 10/7/13; Lee: Lick Cr., Farmington, KJS, 7/28/ 14; Linn: West Otter Cr., Toddville, CTH, 10/11/ 01; Madison: Howerdon Cr., Winterset, JML, 7/ 12/01; Mitchell: Rock Cr., Rock, CTH, 9/10/09; Monroe: Sage Cr., Stephens Forest, CTH, 8/8/ 14; O’ Brien: Little Floyd R., Sheldon, JML, 8/ 22/02; Page: East Nodaway R., Hawleyville, CTH, 10/20/98; Sioux: Sixmile Cr., Hawarden, JML, 8/2/00; Van Buren: Fox R., Mt. Sterling, CTH, 7/16/13; Winneshiek: Trout R., Decorah, CTH, 7/18/07, South Pine Cr., Sattre, CTH, 8/14/ 07. Distribution: Eastern USA, extending west to North Dakota in the north and Oklahoma in the south.

Cymbiodyta vindicata Fall. Benton : Mud Cr., Van Horne, CTH, 9/30/15; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Clayton: Roberts Cr., Gunder, CTH, 8/11/08; Clinton: Silver Cr., DeWitt, CTH, 7/17/12; Dubuque: Hickory Cr., New Vienna, CTH, 9/18/08; Fayette: Deep Cr., Wadena, CTH, 9/28/05; Ida: Odebolt Cr., Ida Grove, CTH, 8/5/97; Jackson: South Big Mill Cr., Bellevue, CTH, 9/17/14; Monroe: Sage Cr., Stephens Forest, CTH, 8/8/14; Ringgold: Lotts Cr., Beaconsfield, JML, 6/21/06; Sioux: Deep Cr., Granville, KLS, 8/2/06; Woodbury: Dutch Cr., Danbury, TMM, 6/20/16. Distribution: transcontinental in the USA, exclusive of the Southwest.

Enochrus cinctus (Say) . Davis: Unnamed trib. to North Fabius R., Bloomfield, CTH, 7/15/13; Jackson: Farmers Cr., LaMotte, CTH, 8/8/07; Johnson: ( Wickham 1911); Van Buren: Fox R., Mt. Sterling, CTH, 7/16/13. Distribution: Eastern USA, extending west to South Dakota in the north, Kansas, and Texas in the south.

Enochrus collinus Brown. Black Hawk : Cedar Hills Sand Prairie (UV light), ELF, 6/19/12; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Henry: Little Potomac Cr., Mt. Pleasant, CTH, 8/7/15; Mahaska: Buck Cr., Barnes City, CTH, 7/26/12; Ringgold: Lotts Cr., Beaconsfield, JML, 6/21/06. Distribution: Northeastern USA, extending west to Minnesota in the north. Iowa forms the southwestern border of its range. New state record.

Enochrus consortus Green. Davis : Soap Cr., Peach Ave., CTH, 7/17/13; Linn: Wapsipinicon R., Viola, DPH, 6/9/01; Louisa: Iowa R., Wapello, MWB, 10/8/13; Muscatine: Pike Run, Nichols, CTH, 8/20/12; Van Buren: Shimek State Forest ( UV light), ELF, 5/18/12. Distribution: eastern USA, extending west to Minnesota in the north, Iowa, Missouri, and Louisiana in the south .

Enochrus diffusus (LeConte) . Howard: ( Wickham 1911). Distribution: Northwestern USA, extending east to Michigan.

Enochrus fimbriatus (Melsheimer) . Johnson: ( Wickham 1911); Scott: ( Wickham 1911); Story: ( Wickham 1911; Gunderson 1967). Distribution: Transcontinental in the USA.

Enochrus hamiltoni (Horn) . Allamakee: French Cr., Churchtown, CTH, 7/1/02; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Butler: Dry Cr., Clarksville, CTH, 5/29/07; Clay: Elk Cr., Gillett Grove, TMM, 9/24/13; Dickinson: ( Wickham 1911); Dubuque: Middle Fk. Little Maquoketa R., Rickardsville, CTH, 9/4/12; Fayette: Volga R., Maynard, CTH, 8/30/10; Hancock: Unnamed trib. to East Branch Iowa R., Goodell, TMM, 6/26/13; Jackson: South Big Mill R., Bellevue, CTH, 9/3/10; Jasper: Walnut Cr., Prairie City, CTH, 8/21/95; Linn: East Otter Cr., Alburnett, CTH, 7/10/02; Poweshiek: Walnut Cr., Malcom, CTH, 7/28/09; Ringgold: Lotts Cr., Beaconsfield, JML, 6/21/06; Story: Dye Cr., Colo, TMM, 6/19/13; Winneshiek: North Bear Cr., Highlandville, CTH, 7/17/07; Woodbury: Dutch Cr., Danbury, TMM, 6/20/16. Distribution: Transcontinental in the USA.

Enochrus ochraceus (Melsheimer) . Allamakee: Yellow R., Postville, CTH, 8/14/00; Black

Hawk: Cedar R., Waterloo, CTH, 8/6/12; Buchanan: Lime Cr., Brandon, CTH, 8/21/08; Clayton: Roberts Cr., Postville, CTH, 8/14/08; Clinton: Sugar Cr., Preston, CTH, 8/2/05; Crawford: Otter Cr., Deloit, TMM, 9/30/13; Decatur: Thompson Fk. Grand R., Davis City, TMM, 9/3/13; Dickinson: Little Sioux R., Lake Park, TMM, 7/23/03; Dubuque: Lytle Cr., Zwingle, CTH, 7/17/12; Fayette: Bell Cr., Clermont, CTH, 9/2/04; Floyd: Cedar R., Charles City, MWB, 9/14/01; Franklin: Boylan Cr., Aredale, TMM, 8/11/14; Fremont: Nishnabotna R., Hamburg, TMM, 10/1/13; Guthrie: Middle Raccoon R., Coon Rapids, MWB, 6/16/ 07; Hardin: South Fk. Iowa R., Iowa Falls, TMM, 8/2/12; Henry: Big Cedar Cr., Oakland Mills, CTH, 8/29/12; Jackson: Unnamed trib. to Big Mill Cr., La Motte, CTH, 9/13/11; Jasper: Walnut Cr., Prairie City, CTH, 5/1/96; Johnson: Muddy Cr., North Liberty, CTH, 10/2/06; Jones: Helmer Cr., Prairieburg, CTH, 8/31/05; Kossuth: East Fk. Des Moines R., Irvington, MWB, 8/25/ 03; Mahaska: Buck Cr., Barnes City, CTH, 7/26/ 12; Mitchell: Wapsipinicon R., McIntyre, CTH, 8/27/08; Monroe: Sage Cr., Stephens Forest, CTH, 8/8/14; Montgomery: Pilot Branch, Stennett, TMM, 9/16/13; Muscatine: Honey Cr., Conesville, CTH, 7/18/06; O’ Brien: Unnamed trib. to Waterman Cr., Sutherland, TMM, 5/20/ 14; Polk: Camp Cr., Mitchellville, KBK, 9/26/ 01; Pottawattamie: Jordan Cr., Macedonia, TMM, 9/17/13; Poweshiek: Walnut Cr., Hartwick, CTH, 7/27/09; Sac: North Raccoon R., Auburn, TMM, 10/1/13; Scott: Lost Cr., Princeton, CTH, 8/ 30/04; Van Buren: Town Branch, Pittsburg, CTH, 9/14/15; Wapello: Des Moines R., Ottumwa, CTH, 10/8/12; Wayne: South Fk. Chariton R., Corydon, TMM, 8/24/12; Winnebago: Winnebago R., Lake Mills, CTH, 8/16/95; Winneshiek: Yellow R., Postville, CTH, 7/25/07; Woodbury: Little Sioux R., Smithland, TMM, 10/3/12. Distribution: Eastern USA, extending west to the Midwest, although West Coast collections exist.

Enochrus pygmaeus (Fabricius) . Allamakee: Waterloo Cr., Dorchester, CTH, 8/1/12; Appanoose: Unnamed trib. to North Cr., Cincinnati, CTH, 9/15/15; Benton: Bear Cr., Shellsburg, CTH, 10/2/13; Boone: Beaver Cr., Beaver, KBK, 7/25/01; Butler: West Fk. Cedar R., Lake Considine Park, CTH, 10/2/12; Clayton: Peck Cr., Osterdock, CTH, 7/25/11; Dallas: Beaver Cr., Perry, JML, 7/25/01; Davis: North Fabius R., Moulton, CTH, 7/17/06; Decatur: Long Cr., Van Wert, CTH, 9/14/95; Dubuque: White Pine Hollow Cr., Luxemburg, CTH, 10/2/03; Fayette: Brush Cr., Wadena, CTH, 9/27/05; Floyd: Cedar R., Charles City, MWB, 9/14/01; Franklin: Beaver Cr., Ackley, MWB, 8/15/05; Guthrie: Middle Raccoon R., Coon Rapids, MWB, 6/16/07; Harrison: Soldier R., Pisgah, JML, 10/25/00; Jackson: Farmers Cr., LaMotte, CTH, 8/8/07; Johnson: Old Mans Cr., Hill, CTH, 9/18/13; Keokuk: North Skunk R., Sigourney, CTH, 10/7/ 13; Linn: Cedar R., Bertram, CTH, 10/10/05; Louisa: Long Cr., Columbus Junction, CTH, 9/ 14/95; Mahaska: North Skunk R., Rose Hill, CTH, 8/22/11; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; Page: West Tarkio Cr., Coin, TMM, 9/7/04; Pottawattamie: Mosquito Cr., Manawa, TMM, 9/ 24/12; Shelby: East Branch West Nishnabotna R., Harlan, KLS, 8/4/05; Van Buren: Town Branch, Pittsburg, CTH, 9/14/15; Wapello: Des Moines R., Ottumwa, CTH, 10/7/13; Webster: Allen Cr., Stratford, JML, 8/15/07; Woodbury: West Fk.Little Sioux R., Hornick, TMM, 8/18/14. Distribution: Transcontinental in the USA, except the West Coast.

Enochrus sayi Gundersen. Des Moines : Flint Cr., Danville, CTH, 7/15/03; Floyd: Cedar R., Carrville, CTH, 8/7/12; Jackson: Maquoketa R., Maquoketa, CTH, 9/13/12. Distribution: Eastern USA, extending west to Iowa in the north, Missouri, and Texas in the south. Iowa forms the northwestern edge of the range.

Helochares maculicollis Mulsant. Buchanan : Wapsipinicon R., Independence, CTH, 8/6/12; Linn: Wapsipinicon R., Central City, CTH, 9/27/ 06; Louisa: Iowa R., Wapello, CTH, 10/15/03; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; Scott: Wapsipinicon R., DeWitt, CTH, 8/30/ 12. Distribution: Northeastern USA, extending west to Missouri and Iowa.

Helocombus bifidus (LeConte) . Black Hawk: Cedar Hills Sand Prairie ( UV light), ELF, 6/19/ 12; Howard: ( Wickham 1911). Distribution: Eastern USA, extending west to Minnesota in the north, Iowa, and Missouri in the south .

Hydrobius fuscipes (Linnaeus) . Black Hawk: Cedar Hills Sand Prairie (UV light), ELF, 6/19/ 12; Jackson: South Big Mill Cr., Bellevue, CTH, 9/17/14. Distribution: Northern transcontinental in the USA, Iowa forms part of the southern border in the Midwest.

Hydrobius melaenus (Germar) . Allamakee: Clear Cr., Lansing, MWB, 8/16/07; Clayton: North Cedar Cr., Clayton, CTH, 9/24/07; Dubuque: Catfish Cr., Dubuque, CTH, 8/1/07; Fayette: Unnamed trib. to Volga Lake outlet, Fayette, CTH, 8/4/15; Winneshiek: North Bear Cr., Highlandville, CTH, 7/17/07. Distribution: Eastern USA, extending west to Wisconsin and Iowa.

Hydrochara leechi Smetana. Winneshiek : Decorah, HFW, 7/20/33; (Downie and Arnett 1996). Distribution: Central USA, from North Dakota to New Mexico in the west, and Pennsylvania, Indiana, and Texas in the east .

Hydrochara obtusata (Say) . Hancock: Pilot Knob Wetland, Forest City, MTF, 7/18/07; Linn: Cedar R., Cedar Rapids, CTH, 8/24/15. Distribution: Northern transcontinental in the USA, Iowa forms part of the southern border in the Midwest.

Hydrochara simula Hilsenhoff and Tracy. Jackson : South Big Mill Cr., Bellevue, CTH, 8/ 28/07. Distribution: Great Lakes region, with Iowa forming the southwestern border.

H. soror Smetana. Allamakee : Little Paint Cr., Harpers Ferry, CTH, 8/27/14; Appanoose: Sharon Bluffs State Park (UV light), ELF, 7/26/ 11; Black Hawk: Beaver Cr., Cedar Falls, MWB, 8/6/12; Butler: West Fk. Cedar R., Dumont, CTH, 10/11/99; Davis: Fox R., Moulton, CTH, 6/29/06; Decatur: UV lighting, ELF, 5/19/12; Jackson: South Big Mill Cr., Bellevue, CTH, 8/6/14. Distribution: Eastern USA, extending west to Texas in the south, Oklahoma, Iowa, and Wisconsin in the north.

H. spangleri Smetana. Allamakee : Dousman Cr., Ion, CTH, 8/12/15; Clinton: Brophy Cr., McCausland, CTH, 8/19/14; Fremont: Waubonsie State Park (UV light), ELF, 6/22/12; Guthrie: Springbrook Cr., Guthrie Center, TMM, 7/16/13; Johnson: Pechman Cr., Hills, CTH, 4/17/17; Kossuth: Private Wetland, Algona, MTF, 7/14/08; Story: Bear Cr., Ames, TMM, 7/7/13. Distribution: Eastern, exclusive of extreme northeastern USA, extending west to Nebraska in the north and Texas in the south. Iowa forms part of the northern border in the Midwest.

Hydrophilus ovatus Gemminger and Harold. Johnson : ( Wickham 1911). Distribution: Eastern USA, extending west to Iowa in the north, Kansas, and Texas in the south. Iowa forms part of the northwestern border .

Hydrophilus triangularis Say. Cerro Gordo : Private Wetland, Ventura, MTF, 7/20/07; Dickinson: Freda Haffner Kettlehole, Milford, MTF, 7/24/07; Franklin: Hampton, FHS, 9/5/ 1896; Fremont: Sapp Bros. Truck stop ( UV light), ELF, 4/17/04; Greene: Snake Cr., Rippey, MTF, 8/14/07; Johnson: State Hygienic Laboratory, Coralville, KJS, 5/20/13; Winnebago: Private Wetland, Thompson, MTF, 7/18/07; Worth: Shell Rock R., Kensett, CTH, 10/15/02. Distribution : Transcontinental in the USA .

Laccobius agilis (Randall) . Madison: North Branch North R., Winterset, CTH, 6/14/96. Distribution: Northern transcontinental in the USA, with Iowa forming the southern border in the Midwest.

Laccobius minutoides (Orchymont) . Allamakee: Upper Iowa R., Dorchester, MWB, 9/11/13; Wayne: South Fk. Chariton R., Corydon, TMM, 8/24/12. Distribution: Northeastern USA, extending west to Minnesota in the north and Kansas in the south. Iowa forms part of the western border .

Laccobius reflexipenis Cheary. Allamakee : Hecker Cr., Postville, CTH, 7/24/07; Appanoose: Packard Cr., Exline, CTH, 9/15/15; Clayton: Maquoketa R., Strawberry Point, CTH, 9/12/07; Dallas: Puckerbush Access (UV light), ELF, 11/ 1/12; Davis: North Fabius R., Moulton, CTH, 7/ 17/06; Delaware: Maquoketa R., Dundee, MWB, 6/24/01; Des Moines: Flint Cr., Danville, CTH, 7/15/03; Dubuque: Hickory Cr., New Vienna, 9/18/08; Fayette: Brush Cr., Wadena, CTH, 9/27/05; Guthrie: Middle Raccoon R., Coon Rapids, MWB, 6/16/07; Hancock: Unnamed trib. to East Branch Iowa R., Goodell, TMM, 6/26/13; Jackson: Farmers Cr., LaMotte, CTH, 8/8/07; Jones: Maquoketa R., Monticello, CTH, 9/6/05; Keokuk: North Skunk R., Sigourney, CTH, 10/7/13; Lee: Shimek State Forest (UV light), ELF, 4/3/05; Linn: Cedar R., Bertram, MWB, 10/10/05; Mitchell: Wapsipinicon R., McIntyre, MWB, 8/27/08; Story: Ballard Cr., Cambridge, TMM, 8/2/13; Wapello: Middle Avery Cr., Chillicothe, MWB, 8/27/02; Winnebago: Buffalo Cr., Forest City, MWB, 9/ 16/06; Winneshiek: North Bear Cr., Highlandville, CTH, 7/17/07. Distribution: Eastern USA, extending west to Minnesota in the north and Kansas in the south. Iowa forms part of the western border.

Laccobius spangleri Cheary. Cherokee : Ashton Cr., Washta, TMM, 7/6/06; Chickasaw: East Fk. Wapsipinicon R., New Hampton, CTH, 8/26/14; Clay: Unnamed trib. to Little Sioux R., Peterson, TMM, 6/29/15; Dubuque: Cloie Branch, Durango, CTH, 8/5/14; Fayette: North Branch Volga R., Randalia, CTH, 9/18/01; Ida: Ashton Cr., Washta, JML, 7/6/06; Jackson: Prairie Cr., Maquoketa, CTH, 8/30/06; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; Sioux: Floyd R., Alton, KLS, 8/6/03; Van Buren: Town Branch, Pittsburg, CTH, 9/14/16; Wayne: Jordan Cr., Millerton, TMM, 9/12/12. Distribution: Northeastern USA, extending west to Minnesota in the north and Kansas in the south. Iowa forms part of the western border.

Laccobius truncatipenis Miller. Clayton : Bear Cr., Edgewood, MWB, 9/9/04; Fayette: Bell Cr., Clermont, CTH, 7/20/04; Harrison: Soldier R., Pisgah, JML, 7/11/00; Jackson: North Fk. Maquoketa R., Maquoketa, CTH, 9/16/13; Shelby: Unnamed trib. to West Nishnabotna R., Irwin, JML, 7/6/05; Winneshiek: South Pine Cr., Sattre, CTH, 8/27/14. Distribution: Northwestern USA, extending east to Michigan. Iowa forms part of the southern border in the Midwest.

Paracymus despectus (LeConte) . Davis: Fox R., Moulton, CTH, 6/29/06. Distribution: Northeastern USA, extending west to Minnesota in the north and Kansas in the south. Iowa forms part of the western border .

Paracymus subcupreus (Say) . Allamakee: Yellow R., Postville, CTH, 8/14/00; Black Hawk: Beaver Cr., Cedar Falls, CTH, 10/9/14; Buchanan: Bear Cr., Brandon, CTH, 8/3/10; Buena Vista: Unnamed trib. to Maple Cr., Aurelia, TMM, 8/27/13; Butler: Palmer Cr., Greene, CTH, 5/29/07; Cedar: Unnamed trib. to Bennett Cr., Tipton, CTH, 7/16/12; Chickasaw: East Fk. Wapsipinicon R., New Hampton, CTH, 9/20/10; Clay: Little Sioux R., Spencer, TMM, 9/6/05; Clayton: Roberts Cr., Postville, CTH, 8/14/08; Davis: Fox R., Moulton, CTH, 6/29/06; Delaware: Twin Springs Cr., Colesburg, CTH, 9/17/07; Dubuque: Tetes Des Morts Cr., St. Donatus, CTH, 8/1/05; Fayette: Little Volga R., Maynard, CTH, 7/23/14; Floyd: Cedar R., Charles City, CTH, 9/14/01; Franklin: Bailey Cr., Sheffield, CTH, 7/20/11; Guthrie: Middle Raccoon R., Bagley, KLS, 9/20/05; Henry: Big Cedar Cr., Oakland Mills, 9/2/11; Howard: North Branch Turkey R., Lime Springs, CTH, 8/ 5/15; Iowa: Deep Cr., Guernsey, CTH, 8/25/11; Jackson: Farmers Cr., LaMotte, CTH, 8/8/07; Johnson: Muddy Cr., North Liberty, CTH, 10/2/ 06; Jones: Helmer Cr., Prairieburg, CTH, 8/31/ 05; Lee: Lick Cr., Farmington, CTH, 7/28/14; Mills: Mud Cr., Hastings, JML, 8/2/05; Mitchell: Wapsipinicon R., McIntyre, MWB, 8/27/08; Montgomery: Pilot Branch, Stennett, CTH, 9/ 9/96; Muscatine: Mud Cr., Wilton, CTH, 9/15/ 03; Pocahontas: Big Cedar Cr., Fonda, JML,9/ 27/06; Poweshiek: Walnut Cr., Brooklyn, CTH, 7/28/09; Story: Ballard Cr., Cambridge, KJS, 7/ 18/07; Van Buren: Town Branch, Pittsburg, CTH, 8/3/15; Wayne: East Fk. Medicine Cr., Allerton, CTH, 8/11/05; Winneshiek: East Pine Cr., Burr Oak, MWB, 9/29/08. Distribution: Transcontinental in the USA, except extreme western states.

Sperchopsis tessellata (Ziegler) . Allamakee: Yellow R., Ion, CTH, 8/29/02; Buchanan: Pine

Cr., Quasqeton, CTH, 8/28/97; Butler: Coldwater Cr., Greene, CTH, 7/22/96; Cedar: Rock Cr., Tipton, CTH, 8/9/95; Chickasaw: Wapsipinicon R., New Hampton, CTH, 10/6/04; Clayton: Roberts Cr., St. Olaf, CTH, 7/31/00; Clinton: Elk R., Teeds Grove, CTH, 8/6/07; Delaware: Plum Cr., Hopkinton, CTH, 8/13/ 15; Dubuque: Little Maquoketa R., Dubuque, CTH, 9/21/95; Fayette: Volga R., Maynard, CTH, 9/20/95; Floyd: Little Cedar R., Colwell, CTH, 10/3/95; Franklin: Bailey Cr., Sheffield, CTH, 7/22/96; Howard: Crane Cr., Lourdes, CTH, 7/19/94; Jackson: Prairie Cr., Maquoketa, CTH, 8/14/01; Jones: Silver Cr., Monticello, MWB, 7/31/01; Linn: East Big Cr., Springville, CTH, 7/18/01; Mitchell: Burr Oak Cr., Osage, CTH, 8/14/95; Muscatine: Pike Run, Nichols, CTH, 8/20/12; Poweshiek: North Skunk R., Millgrove County access, CTH, 7/26/12; Scott: Lost Cr., Princeton, CTH, 8/9/12; Taylor: West Branch 102 R., New Market, CTH, 10/10/95; Winneshiek: North Bear Cr., Highlandville, CTH, 7/17/07; Wright: Otter Cr., Goldfield, KLS, 8/11/00. Distribution: Eastern USA, extending west to Iowa and Arkansas.

Tropisternus affinis Motschulsky. Allamakee : Upper Iowa R., Dorchester, MWB, 9/27/02; Butler: West Fk. Cedar R., Lake Considine County park, CTH, 10/2/12; Clayton: Silver Cr., Monona, CTH, 8/7/06; Delaware: Twin Springs Cr., Colesburg, CTH, 9/17/07; Dubuque: Middle Fk. Little Maquoketa R., Rickardsville, CTH, 9/28/04; Fayette: Deep Cr., Wadena, CTH, 9/28/ 05; Guthrie: Unnamed trib. Middle Raccoon R., Panora, TMM, 9/30/14; Jackson: Farmers Cr., Fulton, CTH, 8/8/07; Kossuth: Plum Cr., Algona, MWB, 8/24/05; Linn: Cedar R., Cedar Rapids, CTH, 9/28/12; Mitchell: Deer Cr., Carpenter, CTH, 8/28/08; Muscatine: Pike Run, West Liberty, CTH, 7/19/06; Winnebago: Buffalo Cr., Forest City, JML, 9/16/06; Winneshiek: Canoe Cr., Decorah, CTH, 9/14/11; Worth: Beaver Cr., Fertile, KLS, 9/19/06. Distribution: Western USA, extending east to Wisconsin in the north and Texas in the south.

Tropisternus blatchleyi Orchymont. Appanoose : Little Shoal Cr., Cincinnati, CTH, 10/4/11; Clinton: Elk R., Teeds Grove, CTH, 8/6/07; Muscatine: Pike Run, Nichols, CTH, 8/20/12; Wapello: Des Moines R., Ottumwa, CTH, 10/7/ 13. Distribution: Eastern USA, with Iowa forming the northwestern border.

Tropisternus collaris (Fabricius) . Appanoose: Little Shoal Cr., Cincinnati, CTH, 10/4/11; Henry: Big Cr., New London, CTH, 8/30/05; Sac: North Raccoon R., Auburn, TMM, 10/1/13;

Van Buren: Des Moines R., Farmington, CTH, 9/6/06; Wayne: East Fk. Medicine Cr., Allerton, CTH, 8/11/05. Distribution: Southeastern USA, with Iowa forming the northwestern border.

Tropisternus columbianus Brown. Allamakee : Unnamed trib. Yellow R., Postvile, CTH, 7/24/ 07; Clayton: Brownfield Cr., Colesburg, CTH, 9/18/07; Muscatine: Pike Run, West Liberty, CTH, 7/19/06; Winneshiek: South Pine Cr., Sattre, CTH, 10/27/10. Distribution: Western USA, extending east to Ohio in the north and Texas in the south.

T. glaber (Herbst) . Allamakee: Irish Hollow Cr., New Albin, DPH, 8/22/05; Appanoose: Unnamed trib. to North Cr., Cincinnati, CTH, 9/15/ 15; Buchanan: Lime Cr., Brandon, CTH, 9/4/07; Clayton: Peck Cr., Osterdock, CTH, 7/25/11; Clinton: Elk R., Teeds Grove, CTH, 8/13/12; Delaware: Little Turkey R., Colesburg, MWB, 9/8/06; Fayette: Brush Cr., Wadena, CTH, 9/27/ 05; Floyd: Little Cedar R., Colwell County Park, CTH, 9/9/09; Jackson: Farmers Cr., Fulton, CTH, 8/8/07; Muscatine: Cedar R., Conesville, CTH, 8/29/12; Winneshiek: Coldwater Cr., Bluffton, CTH, 9/22/04. Distribution: Northeastern USA, extending west to South Dakota and Iowa.

Tropisternus lateralis (Fabricius) . Allamakee: Little Paint Cr., Harpers Ferry, CTH, 8/16/07; Appanoose: Cooper Cr., Centerville, CTH, 8/11/ 05; Benton: Bear Cr., Shellsburg, CTH, 10/2/13; Black Hawk: Black Hawk Cr., Hudson, CTH, 8/ 8/02; Bremer: Cedar R., Janesville, CTH, 9/9/ 14; Buchanan: Bear Cr., Brandon, CTH, 8/26/ 13; Butler: Dry Cr., Clarksville, CTH, 5/29/07; Cherokee: Unnamed trib. to Little Sioux R., Larrabee, TMM, 5/15/12; Chickasaw: Wapsipinicon R., New Hampton, CTH, 9/6/94; Clay: Little Sioux R., Spencer, TMM, 9/9/14; Clayton: Maquoketa R., Strawberry Point, CTH, 9/12/07; Clinton: Elk R., Andover, CTH, 8/7/07; Dallas: North Raccoon R., Adel, TMM, 9/29/05; Davis: Fox R., Bloomfield, CTH, 7/15/13; Decatur: Little R., Leon, CTH, 9/23/02; Delaware: Maquoketa R., Manchester, CTH, 9/25/13; Des Moines: Cottonwood Drain, Burlington, CTH, 8/5/03; Dubuque: North Fk. Maquoketa R., Dyersville, CTH, 7/25/05; Emmet: Drainage Ditch 40, Dolliver, TMM, 7/30/15; Fayette: Volga R., Randalia, CTH, 8/31/10; Floyd: Cedar R., Charles City, CTH, 9/26/02; Franklin: Beaver Cr., Ackley, CTH, 9/10/14; Greene: West Buttrick Cr., Spring Lake Park, TMM, 7/31/14; Hancock: Pilot Knob Wetland, Forest City, MTF, 7/5/07; Hardin: Private Wetland, Buckeye, MTF, 8/14/07; Harrison: Soldier R., Pisgah, TMM, 9/5/12; Henry: Big Cedar Cr., Oakland Mills, CTH, 9/2/11; Jackson: Tetes Des Morts Cr., St. Donatus, CTH, 8/27/07; Jasper: Walnut Cr., Prairie City, CTH, 8/15/01; Johnson: Muddy Cr., North Liberty, CTH, 10/2/06; Jones: Bear Cr., Wyoming, CTH, 7/13/04; Kossuth: Lotts Cr., West Bend, MWB, 9/25/02; Lee: Skunk R., Augusta, CTH, 10/8/13; Linn: Wapsipinicon R., Troy Mills, CTH, 9/24/03; Louisa: Long Cr., Columbus Junction, CTH, 9/14/95; Lucas: Wolf Cr., Chariton, TMM, 9/12/12; Lyon: Little Rock R., George, TMM, 9/28/11; Mahaska: Buck Cr., Barnes City, CTH, 7/26/12; Mitchell: Cedar R., Osage, CTH, 7/6/06; Montgomery: Pilot Branch, Stennett, TMM, 9/16/13; Muscatine: Mud Cr., Wilton, CTH, 9/15/03; O’ Brien: Unnamed trib. to Waterman Cr., Sutherland, TMM, 5/20/14; Plymouth: Big Sioux R., Akron, TMM, 10/10/ 12; Polk: Des Moines R., Des Moines, TMM, 9/ 24/13; Pottawattamie: Mosquito Cr., Manawa, TMM, 9/24/12; Sac: North Raccoon R., Sac City, TMM, 7/31/12; Scott: Lost Cr., Princeton, CTH, 8/9/12; Sioux: Sixmile Cr., Hawarden, KBK, 8/3/ 00; Story: Ballard Cr., Cambridge, TMM, 8/2/13; Union: Unnamed trib. to Thompson R., Lorimor, TMM, 9/17/15; Van Buren: Town Branch, Pittsburg, CTH, 10/1/14; Wayne: South Fk. Chariton R., Corydon, TMM, 9/12/12; Winnebago: Buffalo Cr., UWW10, JML, 9/16/06; Winneshiek: North Bear Cr., Highlandville, CTH, 7/17/07; Woodbury: Wolf Cr., Correctionville, TMM, 9/22/14; Worth: Elk Cr., Kensett, CTH, 8/26/ 08; Wright: Lower Morse Lake Wetland Complex, Belmond, MTF, 7/15/09. Distribution: Eastern USA, extending west to the Rocky Mountains.

Tropisternus mixtus (LeConte) . Allamakee: Dousman Cr., Ion, CTH, 8/12/15; Clayton: Maquoketa R., Strawberry Point, CTH, 9/12/07; Clinton: Sugar Cr., Preston, CTH, 8/2/05; Fayette: Dibble Cr., Clermont, CTH, 9/24/14; Floyd: Cedar R., Floyd, CTH, 9/23/08; Howard: Crane Cr., Lourdes, CTH, 9/28/09; Johnson: Pechman Cr., Hills, CTH, 4/17/17; Linn: Cedar R., Cedar Rapids, CTH, 9/28/12; Muscatine: Muscatine Slough, Muscatine, CTH, 9/17/02; Winnebago: Private Wetland, Thompson, MTF, 7/18/07; Winneshiek: East Pine Cr., Burr Oak, MWB, 9/29/08; Worth: Beaver Cr., Fertile, KLS, 9/19/06. Distribution: Transcontinental in the USA.

Tropisternus natator Orchymont. Allamakee : Clear Cr., Lansing, MWB, 8/16/07; Appanoose: Chariton R., Rathbun, CTH, 8/17/04; Black Hawk: West Fk. Cedar R., Finchford, MWB, 9/ 15/08; Bremer: Cedar R., Janesville, CTH, 8/6/ 12; Buchanan: Lime Cr., Brandon, CTH, 8/21/ 08; Butler: West Fk. Cedar R., Dumont, CTH,

10/11/06; Clarke: West Long Cr., Grand River, TMM, 8/25/15; Clay: Little Sioux R., Spencer, TMM, 9/8/14; Clayton: Turkey R., Garber, MWB, 10/3/03; Clinton: Sugar Cr., Preston, CTH, 8/2/05; Dallas: South Raccoon R., Redfield, TMM, 8/8/12; Davis: Fox R., Moulton, CTH, 6/29/06; Decatur: Little R., Leon, CTH, 9/ 23/02; Delaware: Little Turkey R., Colesburg, MWB, 9/8/06; Des Moines: Cottonwood Drain, Burlington, CTH, 8/5/03; Dubuque: Little Maquoketa R., Dubuque, CTH, 8/19/08; Fayette: Little Turkey R., West Union, CTH, 9/10/03; Floyd: Cedar R., Charles City, CTH, 9/14/01; Greene: West Buttrick Cr., Spring Lake Park; Guthrie: South Raccoon R., Stuart: Hamilton: Boone R., Stratford, TMM, 8/16/12; Hancock: Winnebago R., Forest City, KLS, 9/18/06; Henry: Skunk R., Oakland Mills, CTH, 9/24/ 08; Howard: Crane Cr., Lourdes, CTH, 9/24/ 13; Jackson: Maquoketa R., Canton, CTH, 9/ 7/05; Jasper: Walnut Cr., Prairie City, CTH, 8/15/01; Jefferson: Cedar Cr., Stockport, CTH, 7/20/05; Johnson: Iowa R., Iowa City, CTH, 9/ 22/05; Jones: Helmer Cr., Prairieburg, CTH, 8/31/05; Keokuk: Twin Grove Cr., Ollie, CTH, 5/22/14; Kossuth: East Fk. Des Moines R., Irvington, 8/25/03; Linn: Wapsipinicon R., Troy Mills, CTH, 9/24/03; Louisa: Long Cr., Columbus Junction, CTH, 10/1/13; Lucas: Whitebreast Cr., Woodburn, JML, 9/24/02; Madison: North Branch North R., Winterset, CTH, 8/25/95; Mahaska: Muchakinock Cr., Eddyville, CTH, 9/2/11; Marion: English Cr., Knoxville, CTH, 8/1/13; Mills: West Nishnabotna R., Malvern, TMM, 9/4/12; Mitchell: Wapsipinicon R., McIntyre, CTH, 9/10/07; Muscatine: Pike Run, Nichols, CTH, 8/23/04; Polk: Four Mile Cr., Ankeny, TMM, 8/9/06; Poweshiek: Walnut Cr., Holiday Lake, CTH, 8/25/10; Scott: Lost Cr., Princeton, CTH, 8/9/12; Tama: Richland Cr., Haven, CTH, 10/6/10; Van Buren: Fox R., Mt. Sterling, CTH, 7/19/05; Wapello: Des Moines R., Ottumwa, CTH, 10/7/13; Warren: South R., Ackworth, TMM, 7/23/12; Washington: English R., Riverside, CTH, 10/9/12; Wayne: East Fk. Medicine Cr., Allerton, CTH, 8/11/05; Winneshiek: Upper Iowa R., Decorah, CTH, 7/25/06; Worth: Elk Cr., Kensett, CTH, 8/26/08. Distribution: Eastern USA, extending west to Texas in the south, Colorado, and Minnesota in the north.