Spogostylum ocyale ( Wiedemann, 1828 )

Greathead, D. J., 2003, A sympatric species pair: Spogostylum ocyale (Wiedemann, 1828) and S. griseipenne Macquart, 1850 (Diptera: Bombyliidae), Zootaxa 274, pp. 1-6 : 2-4

Spogostylum ocyale ( Wiedemann, 1828)

Anthrax ocyale Wiedemann, 1828: 285 ( Sudan)

Anthrax umbra Walker, 1849: 245 (patria ignota)

Spogostylum arugotense Zaitzev, 1997: 909 ( Israel)

Spogostylum sordidum Sack, Greathead, 1980: 320 ( Saudi Arabia, females)

The syntype specimen of S. ocyale Wiedemann labelled ‘Nubien Ruppel’ in ZMHB was examined by N. L. Evenhuis and designated lectotype ( Evenhuis & Greathead, 1999). This specimen has no spot on the interradial vein. The holotype of S. umbra Walker in BMNH was also examined and is a very dark specimen without the spot on the interradial vein. Similar dark specimens in the author’s collection have a pointed aedeagus, thus S. umbra was designated a synonym of S. ocyale . Theodor (1983: Figs 594­597) correctly identified S. ocyale . However, Zaitzev (1997) reversed Theodor’s designation of the two forms and described S. ocyale as S. arugotense . Greathead (1980b) identified very dark specimens resembling S. ocyale as S. sordidum but re­examination shows that the females are S. ocyale .

Diagnosis. Interradial vein not infuscated ( Fig. 1 ). Male with aedeagus shorter and pointed ( Fig. 3 : Theodor, 1983: Fig. 596: Greathead, 1967: Fig. 50). Female with spermathecal bulb broad at base and tapering toward apex ( Fig. 5 : Theodor, 1983: Fig. 597). Specimens show the full range of coloration from very dark to very pale.

Distribution. Afrotropical: Somalia, Sudan, Yemen. Palaearctic: Egypt, Libya, Oman, ‘Palestine’, Saudi Arabia, United Arab Emirates.

Material Examined. Afrotropical. SOMALIA: miles E Berbera, coastal plain, vi.1949, K.M. Guichard 1 male in BMNH. SUDAN: Nubien, Ruppel, lectotype male of Anthrax ocyale Wiedemann in ZMHB; Khartoum, 2 males in UMO. SOMALIA: 4 miles S. of Silil, 10.vi.1957, J. Roffey, 1 male in DJG. YEMEN: Jebel Harir, 5200 feet, 26.x­ 6.xi.1937, H. Scott & E.B. Britton, 1 male, 2 females in BMNH; Lahej, 27.xi.1899, W.R.O. Grant, 1 male in BMNH; Lahej­Aden, dates between vi­xi.1895, C.G. Nurse, 6 males, 1 female in BMNH; Reidat Al Ma’ara, 30.viii.1956, D.J. Greathead 1 male in DJG; near Sana’a, 20.viii.1962, G.B. Popov, 1 female in DJG; Seiyun, 25.viii.1956, D.J. Greathead, 1 female in DJG.

Palaearctic. EGYPT: Baharein, 20.vi.1935, J. Omer­Cooper, 2 females in BMNH; Gara, 4­5.viii.1935, J. Omer­Cooper, 1 male in BMNH; Maragi, 26.vi.1935, J. Omer­Cooper, 1 male in BMNH; Sitra, 15.vi.1935, J. Omer­Cooper, 1 male in BMNH; Siwa, 2.vi.1935, J. Omer­Cooper, 2 males in BMNH; same, 30.vi.1935, 1 female; same, 12.vii.1935 1 female; same, 22.vii.1935, 2 females; same, 24.vii.1935, 1 female; no data, Magdi, 1 female in DJG. JORDAN: Al Balqa Gov., Jordan University Farm, Dayr Alla, 20.viii.1994, light trap, 1 male. LIBYA: ix­x.1957, British Ennedi Expedition, 1 female in BMNH. OMAN: Muscat, Lt. Col. Jayakar, 1 male, 1 female in BMNH; Qara Mountains, Ain Al Rizat, 250 feet, xi.1930, B.S. Thomas, 1 male in BMNH; Wadi Tizi, 19.xi.1982, I.L. Hamer, 1 male in BMNH; Jiddat Al Harasis, Yalooni, 10­18.iv.1982, C.F. Dewhurst, 1 male, 2 females in BMNH and DJG; Khabura, alt. 10 m, 28.vi.1980, R. Whitcombe, 1 female in DJG; Salalah, 5.xi.1976, D.J. Greathead, 1 female in DJG; Salalah, Rabat, 6.xi.1976, D.J. Greathead, 4 males in DJG; Dhofar, Qara Mountains, 21.ix.1977, K.M. Guichard, 2 males in DJG. ‘ PALESTINE ’: Jericho, 14.vii.1918, E.A. Austen, 1 female in BMNH; same, 8.vii.1922, P.A. Buxton, 1 female; same, ix.1918, 1 female in BMNH. SAUDI ARABIA: near Abha, 11.vii.1962, G.B. Popov, 1 female in DJG; Mahail, 18.30N 41.49E, 23.xii.1971, 1 female in DJG; 90 km E of Jedda, 14.iv.1980, K.M. Guichard, 1 female in DJG. UNITED ARAB EMIRATES: Salalah, 24.vii.1981, A.R. Pittaway, 1 female in BMNH; Ras Al Kheima, 2.vi.1957, G.B. Popov, 1 female in DJG; same, 20.vi.1957, I female in DJG. PATRIA IGNOTA: Holotype male of Anthrax umbra Walker , ‘presented by Captain Lord Byron’ in BMNH.

Figures

FIGURES 1 ­ 2. Wings of Spogostylum species. 1. S. ocyale (Wiedemann). 2. S. griseipenne (Macquart).

FIGURES 3 ­ 6. Genitalia of Spogostylum species. 3. S. ocyale (Wiedemann), phallosome. 4. S. griseipenne (Macquart), phallosome. 5. S. ocyale, bulb of spermatheca. 6. S. griseipenne, bulb of spermatheca. Scale bars = 0.2 mm.