Atheta (Dimetrota auct.) fenyesi Bernhauer, 1907

Gusarov, Vladimir I., 2003, Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes, Zootaxa 353 (1), pp. 1-134 : 46-49

Atheta (Dimetrota auct.) fenyesi Bernhauer, 1907

( Figs. 47–55 )

Atheta (Dimetrota) fenyesi Bernhauer, 1907: 395 .

Atheta (s. str.) querula Casey, 1910a: 36 , syn. nov.

Dimetrota (s. str.) resima Casey, 1910a: 100 , syn. nov.

Dimetrota (s. str.) vigilans Casey, 1910a: 103 , syn. nov.

Atheta (s. str.) socors Casey, 1911a: 108 , syn. nov.

Dimetrota (s. str.) immerita Casey, 1911a: 141 , syn. nov.

Dimetrota (s. str.) incredula Casey, 1911a: 141 , syn. nov.

Dimetrota (s. str.) opinata Casey, 1911a: 142 , syn. nov.

Dimetrota (s. str.) cerebrosa Casey, 1911a: 142 , syn. nov.

Atheta (Dimetrota) fenyesi: Fenyes, 1920: 205 (as valid species).

Atheta (s. str.) querula: Fenyes, 1920: 214 (as valid species).

Atheta (Dimetrota) resima: Fenyes, 1920: 205 (as valid species).

Atheta (Dimetrota) vigilans: Fenyes, 1920: 205 (as valid species).

Atheta (s. str.) socors: Fenyes, 1920: 215 (as valid species).

Atheta (Dimetrota) immerita: Fenyes, 1920: 205 (as valid species).

Atheta (Dimetrota) incredula: Fenyes, 1920: 205 (as valid species).

Atheta (Dimetrota) opinata: Fenyes, 1920: 205 (as valid species).

Atheta (Dimetrota) cerebrosa: Fenyes, 1920: 205 (as valid species).

Atheta (Dimetrota) fenyesi: Bernhauer & Scheerpeltz, 1926: 663 (as valid species).

Atheta (s. str.) querula: Bernhauer & Scheerpeltz, 1926: 649 (as valid species).

Atheta (Dimetrota) resima: Bernhauer & Scheerpeltz, 1926: 665 (as valid species).

Atheta (Dimetrota) vigilans: Bernhauer & Scheerpeltz, 1926: 665 (as valid species).

Atheta (s. str.) socors: Bernhauer & Scheerpeltz, 1926: 649 (as valid species).

Atheta (Dimetrota) immerita: Bernhauer & Scheerpeltz, 1926: 663 (as valid species).

Atheta (Dimetrota) incredula: Bernhauer & Scheerpeltz, 1926: 663 (as valid species).

Atheta (Dimetrota) opinata: Bernhauer & Scheerpeltz, 1926: 664 (as valid species).

Atheta (Dimetrota) cerebrosa: Bernhauer & Scheerpeltz, 1926: 662 (as valid species).

Atheta (Dimetrota) fenyesi: Moore & Legner, 1975: 360 (as valid species).

Atheta (s. str.) querula: Moore & Legner, 1975: 371 (as valid species).

Atheta (Dimetrota) resima: Moore & Legner, 1975: 372 (as valid species; misspelled as remisa).

Atheta (Dimetrota) vigilans: Moore & Legner, 1975: 377 (as valid species).

Atheta (s. str.) socors: Moore & Legner, 1975: 374 (as valid species).

Atheta (Dimetrota) immerita: Moore & Legner, 1975: 363 (as valid species).

Atheta (Dimetrota) incredula: Moore & Legner, 1975: 363 (as valid species).

Atheta (Dimetrota) opinata: Moore & Legner, 1975: 368 (as valid species).

Atheta (Dimetrota) cerebrosa: Moore & Legner, 1975: 356 (as valid species).

Dimetrota fenyesi: Seevers, 1978: 259 (as valid species).

Xenota querula: Seevers, 1978: 271 (as valid species).

Dimetrota resima: Seevers, 1978: 260 (as valid species).

Dimetrota vigilans: Seevers, 1978: 260 (as valid species).

Xenota socors: Seevers, 1978: 271 (as valid species).

Dimetrota immerita: Seevers, 1978: 259 (as valid species).

Dimetrota incredula: Seevers, 1978: 259 (as valid species).

Dimetrota opinata: Seevers, 1978: 260 (as valid species).

Dimetrota cerebrosa: Seevers, 1978: 259 (as valid species).

Type material. Lectotype of Atheta fenyesi (here designated): ♂, “ Pasadena Cal. Dr. A.Fenyes ”, “Dec.[ember]”, “781.”, “ ♂ ”, “107.”, “ fenyesi Brh. Typus” (yellow label), “Chicago NHMus M.Bernhauer Collection ” ( FMNH) . Paralectotypes: ♀, “Pasadena Cal.[ifornia] Dr. A.Fenyes ”, “1.”, “ fenyesi Brh. Cotyp. ” (yellow label), “Chicago NHMus M.Bernhauer Collection ” ; ♂, “Pasadena Cal.[ifornia] Dr. A.Fenyes ”, “1.”, “ fenyesi Brh. Cotypus. ” (yellow label), “Chicago NHMus M.Bernhauer Collection ” ; ♀, “Pasadena Cal.[ifornia] Dr. A.Fenyes ”, “Dec.[ember]”, “801.”, “ ♀ ”, “ fenyesi Brh. Cotypus. ” (yellow label), “Chicago NHMus M.Bernhauer Collection ” ( FMNH) .

Lectotype of Atheta querula (here designated): ♂, “ Calaveras Co. Cal. [ifornia]”, “ Atheta querula Csy. ”, “TYPE USNM 39263” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Dimetrota resima (here designated): ♂, “San Mateo Cal.[ifornia] Ba[ker]”, “ resima Csy. ”, “TYPE USNM 39113” (red label), “CASEY bequest 1925” ( NMNH) . Paralectotypes: ♂ (with the head and prothorax missing), “San Mateo Cal.[ifornia] Bake[r]”, “ resima ­2 PARATYPE USNM 39113” (red label), “CASEY bequest 1925” ( NMNH) .

Lectotype of Dimetrota vigilans (here designated): ♂, “Pom.[ona, Los Angeles Co.] Cal.[ifornia] 2/2/96 [2.ii.1896 ( Fall )]”, “ vigilans Csy. ”, “TYPE USNM 39121” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Atheta socors (here designated): ♀, “Cal.[ifornia, San Diego]”, “ socors Csy. ”, “TYPE USNM 39296” (red label), “CASEY bequest 1925” ( NMNH).

Holotype of Dimetrota immerita : ♂, “Riv.[erside] Cal.[ifornia] 3­20[?]­90 [20.iii.1890]”, “ immerita Csy. ”, “TYPE USNM 39126” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Dimetrota incredula (here designated): ♂, “Cal.[ifornia, Saucelito , Marin Co.]”, “ incredula Csy. ”, “TYPE USNM 39128” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Dimetrota opinata (here designated): ♂, “Cal.[ifornia, Berkeley , Alameda Co.]”, “ opinata Csy. ”, “TYPE USNM 39129” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Dimetrota cerebrosa (here designated): ♂, “Cal.[ifornia, Saucelito , Marin Co.]”, “ cerebrosa Csy. ”, “TYPE USNM 39127” (red label), “CASEY bequest 1925” ( NMNH) . Paralectotype: ♀, “Cal.[ifornia, Monterey]”, “CASEY determ.? cerebrosa ­2”, “CASEY bequest 1925” ( NMNH) .

Diagnosis. Atheta fenyesi can be recognized among the other species of the subgenus Dimetrota auct . by the presence of four projections at the posterior margin of the male tergum 8 ( Figs. 47, 49 ), by the distinct shape of the median lobe ( Figs. 52–54 ) (particularly the fused and expanded ostial lamellae forming a wide rhomboid sclerite on the abparameral side of the median lobe ( Fig. 54 )) and the distinct shape of the spermatheca ( Fig. 55 ).

Discussion. The types of At. fenyesi , At. querula , Dim. resima , Dim. vigilans , At. socors , Dim. immerita , Dim. incredula , Dim. opinata and Dim. cerebrosa are similar in external characters and in the shape of the aedeagus and spermatheca. All these names are considered to be synonyms.

Distribution. Atheta fenyesi is known from California.

Figures

FIGURES 47–51. Abdominal segment 8 of Atheta fenyesi Bernhauer (male, lectotype of Dimetrota vigilans Casey (47, 49); male, lectotype of At. querula Casey (48); and female, paralectotype of At. fenyesi (50–51)). 47 — male tergum 8; 48 — male sternum 8; 49 — apex of male tergum 8; 50 — female tergum 8; 51 — female sternum 8. Scale bar 0.2 mm (49), 0.4 mm (47–48, 50–51).

FIGURES 52–55. Genitalia of Atheta fenyesi Bernhauer (male, lectotype of At. querula Casey (52–54); and female, paralectotype of At. fenyesi (55)). 52 — median lobe, parameral view; 53 — median lobe, lateral view; 54 — median lobe, abparameral view; 55 — spermatheca. Scale bar 0.2 mm.