Rhagada setzeri setzeri Maassen, 2009

Köhler, Frank, 2014, On the land snail Rhagada Albers, 1860 (Gastropoda: Camaenidae) from across the Lesser Sunda Islands, Raffles Bulletin of Zoology 62, pp. 115-123 : 119

Rhagada setzeri setzeri Maassen, 2009

Rhagada setzeri Maassen, 2009: 66–69 , figs 13–16 (from ‘ Alor Island , near the airport’, 8° 15' 0" S, 124° 45' 0" E; holotype RMNH 113381 ).

Material examined. AM C.471156 ( Indonesia, Alor Island, 3 dry).

Diagnosis. Shell comparatively large (D = 16–19 mm), with usually well-developed axial ribs, umbilicus closed. Penis with long epiphallus, short flagellum, inner penial anatomy unknown (based on Maassen, 2009).

Comparative remarks. Comparatively large shell with well-developed axial sculpture and closed umbilicus is most characteristic. Well-differentiated from R. solorensis and R. abbasi n. sp. in a molecular phylogeny based on mitochondrial COI and 16S sequences ( Fig. 3 ). Specimens from Atauro Island differ by consistently smaller shell, weaker sculpture, narrowly open umbilicus but are differentiated by comparatively low genetic distances ( Table 2 ); therefore being recognised as distinct subspecies.

Figures

Fig. 3. Bayesian consensus phylogram based on analyses of concatenated 16S and COI sequences, adapted from Köhler & Criscione (2013). Lesser Sunda species Rhagada solorensis, R. setzeri setzeri, R. setzeri atauroensis n. subsp., and R. abbasi n. sp. form the sister clade of Rhagada from the Australian Pilbara. Clades from the Australian Kimberley and Pilbara regions collapsed; outgroup pruned from tree; see Köhler & Criscione (2013) for further details.