Quadrophenia Rifkind, 2017

Rifkind, Jacques, 2017, New genera and species of mimetic Cleridae from Mexico and Central America (Coleoptera: Cleroidea), Insecta Mundi 2017 (591), pp. 1-18 : 4

Quadrophenia Rifkind , new genus

( Fig. 5–13 , 17–21)

Type species. Quadrophenia townshendi Rifkind (by present designation)

Diagnosis. Members of the new genus bear similarities to some species belonging to the genus Enoclerus Gahan , and to the monotypic genus Systenoderes Spinola. Quadrophenia species can be differentiated from both genera by a unique combination of broad, subflattened pronotal dorsum, relucent pronotal and elytral integument, and the shape of antennomere 11, which is broadly emarginate internally, as well as on the distal/ external side ( Fig. 17–21). Systenoderes amoenus Spinola is patterned very similarly to some species of Quadrophenia , although the dark pronotal maculation in Systenoderes appears always to be interrupted longitudinally at the middle, whereas it is always entire (when present) in Quadrophenia .

Description. Clerinae . Form: medium sized, body subflattened; integument shining. Head: eyes mod- erate in size, finely faceted; ocular emargination triangular, moderate; frons shallowly bi-impressed, surface shining, indistinctly, shallowly, rather sparsely punctate; maxillary palpus with terminal palpomere digitiform, apex subtruncate; labial palpus with terminal palpomere securiform; antennae extending to or slightly extending past posterior margin of pronotum when laid alongside; antennomeres 3–5 slightly flattened, subcylindrical; antennomeres 6–8 subconical; antonnomeres 9–11 forming a distinct, rather abruptly expanded club, with antennomeres 9–10 subserrate and antennomere 11 enlarged, polygonal, bilaterally compressed apically, emarginate internally and on distal/ external side ( Fig. 17–21), apex subacuminate; gular process narrow. Pronotum: broader than long; subflat- tened above; transverse impression distinct, rather deeply incised, broadly U-shaped at middle; foveae shallow but rather broad; sides slightly to moderately convex; integument shining, rather sparsely, shallowly punctate; vestiture rather sparse, varied in composition, including robust, elongate, erect and suberect setae; pronotal collar rather narrow longitudinally. Scutellum: broad, rounded posteriorly. Elytra: subflattened above; approximately 1.5× as long as broad; subbasal tumescences absent; umbones prominent; sides sinuate behind rounded humeri; posterior moderately expanded laterally; apices dehiscent; surface shining; punctation moderately densely but irregularly arrayed, not arranged in striae; minute asetiferous punctures also present; vestiture inconspicuous. Legs: tibial spur formula 1-2-2; tarsal pulvilli well developed. Mesosternum: posterior process very narrow, elongate, not elevated distally. Metasternum: anterior process not elevated apically. Abdomen: ventrites shining, inconspicuously setose; pygidium without elaborate modifications; male pygidium without setal daggers; aedeagus well sclerotized, parameres rather short and subsagittate posteriorly; phallus subacuminate apically.

Distribution. The genus ranges from southern Mexico to Panama. Specimens of some undescribed South American species may eventually be assigned to this genus.

Etymology. The genus is named for The Who’s iconic 1973 concept album. Quadrophenia is treated as feminine.

Figures

Figures 5–7. Quadrophenia townshendi new species. 5) habitus of holotype; 6) habitus of paratype; 7) habitus of paratype.

Figures 8–11. Quadrophenia n.spp. 8) Quadrophenia mooni new species, habitus; 9) Quadrophenia entwistlei new species, habitus; 10) Quadrophenia entwistlei, lateral view; 11) Quadrophenia daltreyi new species, habitus.

Figures 12–13. Quadrophenia sumidero new species. 12) habitus; 13) lateral view.