Cheilosia ( Cheilosia ) inarmata, Bot & Mengual & Meutter & Skevington, 2025

Bot, Sander, Mengual, Ximo, Meutter, Frank Van de & Skevington, Jeffrey H., 2025, Review of the genus Cheilosia Meigen, 1822 (Diptera: Syrphidae) from the Caucasus, with the description of 14 new species, European Journal of Taxonomy 1023, pp. 1-181 : 71-77

Cheilosia ( Cheilosia) inarmata sp. nov.

urn:lsid:zoobank.org:act:

Figs 36–37

Cheilosia impressa Loew, 1840 – Stackelberg & Richter 1968: 246 (Caucasian records). — Stackelberg 1970: 61 (Caucasian records). — Peck 1988: 105 (Caucasian records). — Barkalov 1993: 724 (Caucasian records). — Mengual et al. 2020: 17. — Speight 2020a: 37 (Caucasian records).

Cheilosia impressa Loew, 1848 [sic] – Gujabidze 2002: 245.

Differential diagnosis

Cheilosia inarmata sp. nov. is similar to C. impressa Loew, 1840 and C. schnabli , but these two species are unique among all European Cheilosia in possessing a distinct, knob-like basolateral projection on the external surface of the procoxae ( Fig. 37C) ( Vujić et al. 1998; Speight 2020a). This diagnostic projection is absent in C. inarmata ( Fig. 37B). Given the large morphological variation within both C. inarmata and C. impressa , other differences between C. inarmata and C. impressa could not be found, except for the frontal triangle in the male usually pruinose in our new species (male frontal triangle usually shiny in C. impressa ). Besides the absence of the projection on the fore coxa, differences with C. schnabli include the rough and densely punctured scutum in C. inarmata (in C. schnabli the scutum is finely punctured), tergum III in male with a dull area in anterior half never reaching the posterior margin (in C. schnabli male tergum III with dull area extending to posterior margin), and the dorsal lobe of postgonite of the male genitalia broadened below apex ( Fig. 37F) and sclerite of the distiphallus asymmetric ( Fig. 37E, dorsal lobe of postgonite narrow and elongated in apical part and sclerite of the distiphallus symmetric in C. schnabli ). Females of C. inarmata have a short and adpressed pile on scutum, while females of C. schnabli have a semi-adpressed short pile mixed with some longer pile on scutum.

Etymology

The species name derives from the Latin prefix ‘ in -’ meaning ‘without’ ( Brown 1956: 436) and the Latin word ‘ armatus ’ meaning ‘furnished with weapons’ ( Brown 1956: 806), and it refers to the lack of a knob-like projection on the fore coxae. Species epithet is to be treated as an adjective.

Material examined

Holotype

GEORGIA • ♂; Samtskhe-Javakheti, 41.8234° N, 42.8400° E; 2025 m a.s.l.; 11 Jun. 2019; X. Mengual leg.; ZFMK, ZFMK-DIP-00066425 = ZFMK-TIS-8008794 .

Paratypes

GEORGIA – Adjara Region • 1 ♀; Kintrishi Nature Reserve ; 41.7553° N, 42.1129° E; 2280 m a.s.l.; 14–28 Jul. 2018; GGBC-members leg.; ZFMK, ZFMK-TIS-8010475 • 1 ♂; Kintrishi Nature Reserve ; 41.7344° N, 41.9888° E; 458 m a.s.l.; 20 Jul. 2018; X. Mengual leg.; ZFMK, ZFMK-DIP-00053910 = ZFMK-TIS-8005513 • 1 ♀; same data as for preceding; ZFMK, ZFMK-DIP-00053911 = ZFMKTIS-8005520 • 1 ♀; Mtirala National Park, Mount Mtirala ; 41.7990° N, 41.6576° E; 1318 m a.s.l.; 25 Jul. 2018; A. Reimann leg.; MTD, Dip-AR-4545 = ZFMK-TIS-8002677 . – Imereti • 1 ♀; road from Abastumani to Sairme ; 41.86433° N, 42,77842° E; 1830 m a.s.l.; 10 Jun. 2019; X. Mengual leg.; ZFMK, ZFMK-DIP-00066415 . – Kakheti • 2 ♂♂; Lagodekhi N.P.; 41.8635° N, 46.3412° E; 1851 m a.s.l.; 2 Jul. 2021; S. Bot leg.; SBA, SB.003170 , SB.003171 • 2 ♀♀; same data as for preceding; SBA, SB.003172 , SB.003173 • 1 ♀; same data as for preceding; ZFMK, ZFMK-TIS-8011515 • 1 ♀; Lagodekhi N.P.; 41.877° N, 46.384° E; 2652 m a.s.l.; 3 Jul. 2021; S. Bot leg.; SBA, SB.003169 • 2 ♀♀; Lagodekhi N.P.; 41.9032° N, 46.3876° E; 2807 m a.s.l.; 4 Jul. 2021; S. Bot leg.; SBA, SB.003167 , SB.003168 . – Mtskheta-Mtianeti • 2 ♀♀; Stepantsminda ; 42.6591° N, 44.6366° E; 1740–2500 m a.s.l.; 3 Aug. 2001; J.-H. Stuke leg.; ZFMK, ZFMK-DIP-00058257 , ZFMK-DIP-00058258 • 1 ♂, 1 ♀; Kazbeg, Dzhuta ; 42.564° N, 44.735° E; 2183 m a.s.l.; 11 Jul. 2012; Th. Zeegers leg.; GPA • 2 ♀♀; Tbilisi National Park ; 41.879° N, 45.016° E; 22 Jun. 2019; 1251 m a.s.l.; S. Bot leg.; SBA, SB.003175 , SB.003176 • 1 ♀; Tbilisi N.P. ; 41.8808° N, 45.0203° E; 1270 m a.s.l.; 22 Jun. 2019; X. Mengual leg.; ZFMK, ZFMKDIP-00066405 = ZFMK-TIS-8008785 • 1 ♂; Stepantsminda ; 42.6722° N, 44.6113° E; 2070 m a.s.l.; 4 Jul. 2019; A. Reimann leg.; MTD, ZFMK-TIS-8002827 • 1 ♀; same data as for preceding; MTD, Dip-AR-5259 = ZFMK-TIS-8002826 • 1 ♀; Stepantsminda ; 44,6492° N, 42.6546° E; 1825 m a.s.l.; 6–12 Jul. 2019; A. Ssymank leg.; Malaise trap; ASW, ZFMK-DIP-00069439 = ZFMK-TIS-8009526 • 1 ♀; Gveleti , road to waterfall; 44.6200° N, 42.7053° E; 1560 m a.s.l.; 6 Jul. 2019; A. Reimann leg.; MTD, Dip-AR-4749 = ZFMK-TIS-8002801 • 1 ♂; Juta , 42.5794° N, 44.7433° E; 2150 m a.s.l.; 10 Jul. 2019; A. Ssymank leg.; ASW • 1 ♂; Stepantsminda ; 42.7047° N, 44.6210° E; 1540 m a.s.l.; 12 Jul. 2019; A. Ssymank leg.; ASW, Ssy9506-03 = ZFMK-TIS-8009417 • 3 ♂♂; Gergeti ; 42.6669° N, 44.6127° E; 2175 m a.s.l.; 13 Jul. 2019; A. Ssymank leg.; ASW • 1 ♀; Gergeti ; 42.6616° N, 44.6155° E; 2080 m a.s.l.; 13 Jul. 2019; A. Ssymank leg.; ASW • 1 ♀; Stepantsminda ; 42.6577° N, 44.6568° E; 1950 m a.s.l.; 17 Jul. 2019; A. Ssymank leg.; ASW, Ssy9523-02 = ZFMK-TIS-8009418 ; ASW • 1 ♀; Tbilisi N.P. ; 41.880° N, 45.023° E; 1289 m a.s.l.; 4 May 2023; S. Bot leg.; SBA, SB.003157 • 2 ♂♂, 1♀; Tbilisi N.P. ; 41.877° N, 45.0138° E; 1250 m a.s.l.; 4 May 2023; F. Van de Meutter leg.; FMT • 1 ♂; Kithoki ; 42.4147° N, 44.7549° E; 1240 m a.s.l.; 8 May 2023; F. Van de Meutter leg.; FMT • 2 ♀♀; Tbilisi N.P. ; 41.8770° N, 45.0137° E; 1248 m a.s.l.; 9 May 2023; S. Bot leg.; SBA, SB.003158 , SB.003159 . – Racha-Lechkhumi and Kvemo Svaneti • 4 ♂♂; Tsana ; 42.909° N, 43.142° E; 1900 m a.s.l.; 29 Jun. 2018; S. Bot leg.; SBA, SB.003199 to SB.003202 • 1 ♂; Zeskho ; 42.888° N, 43.233° E; 1900 m a.s.l.; 30 Jun. 2018, S. Bot leg.; SBA, SB.003195 • 2 ♀♀; same data as for preceding; SBA, SB.003196 , SB.003197 • 1 ♂; Zeskho ; 42.888° N, 43.233° E; 1900 m a.s.l.; 1 Jul. 2018; S. Bot leg.; SBA, SB.003193 • 1 ♀; same data as for preceding; SBA, SB.003194 • 1 ♂; Tsana ; 42.90° N, 43.14° E; 1830 m a.s.l.; 18 Jun. 2019; J. van Steenis leg.; JSB, 2019-01.061 • 2 ♂♂; Tsana ; 42.8887° N, 43.1429° E; 1757 m a.s.l.; 18 Jun. 2019; X. Mengual leg.; ZFMK, ZFMK-DIP-00066432 , ZFMK-DIP-00066440 • 1 ♂; Tsana ; 42.915° N, 43.143° E; 1969 m a.s.l.; 19 Jun. 2019; S. Bot leg.; SBA, SB.003178 • 1 ♀; same data as for preceding; SBA, SB.003177 • 1 ♂; same data as for preceding; L. Hofstee leg.; LHH • 6 ♂♂; Tsana ; 42.916° N, 43.142° E; 1975 m a.s.l.; 19 Jun. 2019; X. Mengual leg.; ZFMK, ZFMK-DIP-00066428 , ZFMKDIP-00066429 = ZFMK-TIS-8008795 , ZFMK-DIP-00066430 , ZFMK-DIP-00066431 , ZFMKDIP-00069433 = ZFMK-TIS-8009513 , ZFMK-DIP-00066436 • 3 ♀♀; same data as for preceding; ZFMK, ZFMK-DIP-00066416 , ZFMK-DIP-00066438 , ZFMK-DIP-00069441 = ZFMK-TIS-8009511 • 2 ♂♂; Tsana ; 42.8862° N, 43.1563° E; 2252 m a.s.l.; 19 Jun. 2019; J.H. Skevington leg.; CNC, CNC1386850 , CNC1386878 • 1 ♀; same data as for preceding; CNC, CNC1386883 • 1 ♀; 42.82111° N, 43.16069° E; 1450 m a.s.l.; 20 Jun. 2019; X. Mengual leg.; ZFMK, ZFMK-DIP-00066417 = ZFMKTIS-8008789 • 1 ♀; near Adishi ; 42.9986° N, 42.9088° E; 2140 m a.s.l.; 11 Sep. 2019; J. Mortelmans leg.; FMT • 1 ♂; above Glola ; 42.668° N, 43.630° E; 2598 m a.s.l.; 9 Jul. 2021; S. Bot leg.; SBA, SB.003160 • 1 ♂; same data as for preceding; XLS, SB.003161 • 1 ♀; same data as for preceding; ZFMK, ZFMK-TIS-8011487 . – Samegrelo-Zemo Svaneti • 1 ♂; Mestia ; 43.080° N, 42.762° E; 1800 m a.s.l.; 22 Jun. 2018; S. Bot leg.; SBA, SB.003214 • 2 ♀♀; Ushguli ; 42.914° N, 43.010° E; 2100 m a.s.l.; 23 Jun. 2018; S. Bot leg.; SBA, SB.003212 , SB.003213 • 1 ♂; Ushguli ; 42.950° N, 43.075° E; 2275 m a.s.l.; 24 Jun. 2018; S. Bot leg.; SBA, SB.003209 • 2 ♀♀; same data as for preceding; SBA, SB.003210 , SB.003211 • 1 ♂; 42.948° N, 43.021° E; 2800 m a.s.l.; 25 Jun. 2018; S. Bot leg.; SBA, SB.003207 • 1 ♀; same data as for preceding; SBA, SB.003208 • 2 ♂♂; 42.963° N, 42.990° E; 2300 m a.s.l.; 26 Jun. 2018; S. Bot leg.; SBA, SB.003203 , SB.003204 • 2 ♀♀; same data as for preceding; SBA, SB.003205 , SB.003206 • 1 ♀; Ushguli ; 42.916° N, 43.018° E; 2140 m a.s.l.; 29 Jun. 2018; S. Bot leg.; SBA, SB.003198 • 1 ♂; Mestia ; 43.042° N, 42.768° E; 1473 m a.s.l.; 13 Jun. 2019; S. Bot leg.; SBA, SB.003182 • 1 ♀; Mestia ; 43.0424° N, 42.7681° E; 1473 m a.s.l.; 13 Jun. 2019; J.H. Skevington leg.; CNC, CNC1386425 • 2 ♂♂; 42.948° N, 43.070° E; 2258 m a.s.l.; 15 Jun. 2019; S. Bot leg.; SBA, SB.003180 , SB.003181 • 3 ♂♂; 42.9437° N, 43.0539° E; 2220 m a.s.l.; 15 Jun. 2019; X. Mengual leg.; ZFMK, ZFMK-DIP-00066433 , ZFMK-DIP-00066434 = ZFMK-TIS-8008796 , ZFMK-DIP-00066441 • 1 ♀; same data as for preceding; ZFMK, ZFMK-DIP-00066437 • 3 ♂♂; 42.9142° N, 43.0076° E; 2082m a.s.l.; 15 Jun. 2019; X. Mengual leg.; ZFMK, ZFMK-DIP-00066435 , ZFMK-DIP-00066439 = ZFMK-TIS-8008797 , ZFMK-DIP-00066442 • 4 ♂♂; Ushguli ; 42.56° N, 43.04° E; 2260 m a.s.l.; 15 Jun. 2019; J. van Steenis leg.; JSB, 2019-01.043 to 2019-01.046 • 1 ♀; same data as for preceding; JSB, 2019-01.047 • 3 ♂♂; Ushguli ; 42.91° N, 43.01° E; 2100 m a.s.l.; 15 Jun. 2019; J. van Steenis leg.; JSB, 2019-01.048 to 2019-01.050 • 2 ♀♀; same data as for preceding; JSB, 2019-01.051 , 2019-01.052 • 1 ♂; Ushguli ; 42.94° N, 43.05° E; 2220 m a.s.l.; 15–17 Jun. 2019; J. van Steenis leg; Malaise trap; JSB, 2019- 01.058 • 1 ♀; same data as for preceding; JSB, 2019-01.059 • 1 ♂; 42.9471° N, 43.0671° E; 2257 m a.s.l.; 15 Jun. 2019; J.H. Skevington leg.; CNC, CNC1386559 • 1 ♀; Ushguli ; 42.909° N, 43.007° E; 2294 m a.s.l.; 16 Jun. 2019; S. Bot leg.; SBA, SB.003179 • 1 ♂; Ushguli ; 42.56° N, 43.04° E; 2260 m a.s.l.; 16 Jun. 2019; J. van Steenis leg.; JSB, 2019-00.908 = ZFMK-TIS-8009608 • 1 ♀; same data as for preceding; JSB, 2019-01.055 • 1 ♀; Ushguli ; 42.94° N, 43.07° E; 2300 m a.s.l.; 16 Jun. 2019; J. van Steenis leg.; JSB, 2019-01.056 • 2 ♀♀; Ushguli ; 42.91° N, 43.01° E; 2135 m a.s.l.; 16 Jun. 2019; J. van Steenis leg.; JSB, 2019-01.053 , 2019-01.054 • 1 ♂; Ushguli ; 42.910° N, 43.000° E; 2120 m a.s.l.; 17 Jun. 2019; L. Hofstee leg.; LHH • 1 ♀; near Ushguli , up to the ruins; 42.910° N, 43.007° E; 2295 m a.s.l.; 17 Jun. 2019; X. Mengual leg.; ZFMK, ZFMK-DIP-00066418 = ZFMK-TIS-8008790 • 1 ♂; Ushguli ; 42.91° N, 42.93° E; 2290 m a.s.l.; 17 Jun. 2019; J. van Steenis leg.; JSB, 2019-01.057 • 1 ♂; near Shovi ; 42.713° N, 43.701° E; 2022 m a.s.l.; 7 Jul. 2021; S. Bot leg.; SBA, SB.003166 • 2 ♀♀; same data as for preceding; SBA, SB.003164 , SB.003165 • 1 ♂; near Chiora ; 42.7625° N, 43.5540° E; 2027 m a.s.l.; 8 Jul. 2021; S. Bot leg.; SBA, SB.003162 • 1 ♀; same data as for preceding; SBA, SB.003163 ; SBA • 2 ♂♂; 42.9125° N, 43.0549° E; 2299 m a.s.l.; 11 Jul. 2021; S. Bot leg.; SBA, SB.003152 , SB.003153 • 1 ♂; same data as for preceding; ZFMK, ZFMK-TIS-8011554 • 1 ♀; same data as for preceding; ZFMK, ZFMK-TIS-8011555 • 1 ♀; 42.9159° N, 43.0191° E; 2151 m a.s.l.; 12 Jul. 2021; S. Bot leg.; SBA, SB.003154 • 1 ♂; 42.9616° N, 43.0914° E; 2447 m a.s.l.; 12 Jul. 2021; S. Bot leg.; SBA, SB.003155 • 2 ♂♂; 42.9091° N, 43.0031° E; 2166 m a.s.l.; 13 Jul. 2021; S. Bot leg.; SBA, SB.003149 , SB.003150 • 1 ♂; same data as for preceding; ZFMK, ZFMK-TIS-8011494 • 1 ♀; same data as for preceding; SBA, SB.003151 • 1 ♀; 43.0275° N, 42.9115° E; 2887 m a.s.l.; 14 Jul. 2021; S. Bot leg.; SBA, SB.003156 . – Samtskhe-Javakheti • 1 ♀; Borjomi N.P. ; 41.831° N, 43.268° E; 1780 m a.s.l.; 17 Jun. 2018; S. Bot leg.; SBA, SB.003215 • 2 ♂♂; Borjomi N.P. ; 41.824° N, 42.848° E; 2165 m a.s.l.; 10 Jun. 2019; S. Bot leg.; SBA, SB.003187 , SB.003188 • 4 ♀♀; same data as for preceding; SBA, SB.003189 to SB.003192 • 3 ♂♂, 2 ♀♀; same data as for preceding; L. Hofstee leg.; LHH • 4 ♂♂; Borjomi N.P.; 41.86° N, 42.77° E; 1830 m a.s.l.; 10 Jun. 2019; J. van Steenis leg.; JSB, 2019-01.023 , 2019-01.027 = ZFMK-TIS-8009609 , 2019-01.028 , 2019-01.029 • 3 ♀♀; same data as for preceding; JSB, 2019-01.030 to 2019-01.032 • 1 ♀; Borjomi N.P.; 41.864° N, 42.789° E; 1798 m a.s.l.; 11 Jun. 2019; S. Bot leg.; SBA, SB.003186 • 4 ♂♂; Borjomi N.P.; 41.8234° N, 42.84° E; 2025 m a.s.l.; 11 Jun. 2019; X. Mengual leg.; ZFMK, ZFMK-DIP-00066423 = ZFMK-TIS-8008793 , ZFMK-DIP-00066424 , ZFMK-DIP-00066426 , ZFMK-DIP-00066427 • 11 ♀♀; same data as for preceding; ZFMK, ZFMKDIP-00066406 = ZFMK-TIS-8008786 , ZFMK-DIP-00066409 , ZFMK-DIP-00066410 = ZFMKTIS-8008788 , ZFMK-DIP-00066411 to ZFMK-DIP-00066414 , ZFMK-DIP-00066419 , ZFMKDIP-00066420 = ZFMK-TIS-8008792 , ZFMK-DIP-00066421 • 1 ♂; Borjomi N.P.; 41.863° N, 42.789° E; 1800 m a.s.l.; 11 Jun. 2019; X. Mengual leg.; ZFMK, ZFMK-DIP-00066443 = ZFMKTIS-8008798 • 1 ♂; Borjomi N.P.; 41.82° N, 42.84° E; 2100 m a.s.l.; 11 Jun. 2019; J. van Steenis leg.; JSB, 2019-01.033 • 6 ♀♀; same data as for preceding; JSB, 2019-01.034 to 2019-01.039 • 1 ♀; Abastumani ; 41.79° N, 42.82° E; 1515 m a.s.l.; 11 Jun. 2019; J. van Steenis leg.; JSB, 2019-01.041 • 1 ♂; Borjomi N.P. ; 41.8181° N, 42.8316° E; 2004 m a.s.l.; 11 Jun. 2019; J.H. Skevington leg.; CNC, CNC1386312 • 4 ♀♀; same data as for preceding; CNC, CNC1386258 , CNC1386259 , CNC1386261 , CNC1386275 • 1 ♂; Borjomi N.P. ; 41.8224° N, 42.8444° E, 2098 m a.s.l.; 11 Jun. 2019; J.H. Skevington leg.; CNC, CNC1386170 • 1 ♀; Borjomi N.P. ; 41.8697° N, 42.7923° E; 1683 m a.s.l., 11 Jun. 2019; J.H. Skevington leg.; CNC, CNC1386364 • 2 ♂♂; Borjomi N.P. ; 41.823° N, 42.841° E; 2049 m a.s.l.; 11 Jun. 2019; S. Bot leg.; SBA, SB.003183 , SB.003184 • 1 ♀; same data as for preceding; SBA, SB.003185 • 1 ♂; Borjomi N.P. ; 41.8312° N, 43.2686° E; 1773 m a.s.l.; 28 Jun. 2021; S. Bot leg.; SBA, SB.003174 .

Description

Male

LENGTH. Body 6–9 mm, wing 6–8 mm.

HEAD. Face bare, black, with facial tubercle, pruinose except facial tubercle, area under antenna more densely pruinose, below lunule narrower than an eye. Mala black, pruinose except ventrally and anteriorly where bordering clypeus. Parafacia black, pruinose, white or black and white pilose, about half as wide as postpedicel. Clypeus pruinose, ranging from as wide as long to 1.5 times as long as wide measured from the midline. Frontal triangle black, pruinose, most densely along eye margins, with long black pile. Length of eye contiguity about 1.2 to 1.5 times as long as frons. Angle of approximation of eyes 90–110°. Vertical triangle black, slightly pruinose, ocellar triangle long black pilose. Occiput pruinose, dorsally long black pilose. Lunule dark orange to blackish, with distinct medial arm, separating acetabula. Scape black, anteriorly with black setae; pedicel black, anteriorly with black setae; postpedicel black, sometimes predominantly orange or basoventral corner orange, squarish, about as high as wide or slightly wider than high, pruinose; arista black, with pile, pile shorter than diameter of arista at base. Eye with pile, colour of pile variable, yellow, brown or black.

THORAX. Thorax black. Scutum shiny, coarsely punctured, with short dense black erect to semi-adpressed pile and long black almost erect pile, sometimes anteriorly with portion of pile yellow instead of black, laterally with some long black setae. Anterior part of notopleuron pruinose. Scutellum shiny, with long black erect and denser short black erect pile, with black setae along posterior margin. Pleura slightly pruinose, with black pile except ventral part of katepisternum with long white pile and metasternum usually bare, but sometimes with few white pile; dorsal and ventral pile patches on katepisternum connected. Haltere pedicellum and capitulum yellow.

WING. Wing including alula entirely microtrichose. Pterostigma yellow-brown. Veins brown to black, except at wing base, where veins are yellow and contrasting with remainder dark veins.

LEGS. With black and yellow pile. Coxae and trochanters black, procoxa without basolateral knob-like projection on the external surface. Femora black, sometimes the apex dark orange. Tibiae black, sometimes the base dark orange. Tarsi black.

ABDOMEN. Abdomen black. Tergum I with long white pile, longest along lateral margins, terga II–IV with long white pile, medially shorter and more adpressed, posterior margin with black pile, most abundant in posterolateral corners, in rare cases pile on II–IV black; tergum I pruinose, tergum II pruinose except lateral margins shiny, tergum III pruinose anteromedially, pruinosity decreasing posteriorly, lateral margins and posterior margin shiny, tergum IV pruinose anteromedially, extending medially in decreasing width, V-shaped, not reaching posterior margin. Sterna II–IV pruinose; sternum I white pilose, length of pilosity increasing from anterolaterally short to long medially, medially pile bent posteriorly in apical half, sternum II with erect long white pile, sterna III–IV laterally with long erect white pile, medially in anterior part with short adpressed white pile, in posterior part with short adpressed black pile. Genitalia with surstylus 1.5 to 1.7 times as long as wide, with keel, area above keel with field of microtrichia ( Fig. 37G); dorsal lobe of postgonite broadened below apex; sclerite of the distiphallus asymmetric with right apical extension longer than left apical extension ( Fig. 37E).

Female

LENGTH. Body 6–9 mm, wing 6–8.5 mm.

Similar to the male, except for normal sexual dimorphism and the following characters. Frons shiny except anterolateral corners with small triangular area of grey pruinosity, anteriorly with erect yellow, black or mixed yellow and black pile, posteriorly with longer black pile, with paravertica. Occiput dorsally wide and shiny. Pile on eye yellow. Scutum and scutellum with short adpressed pile, colour variable, black, golden-yellow or mixed golden-yellow and black. Colour pile on pleura variable, black, yellow or mixed yellow and black. Terga II–V shiny except tergum II anteromedially pruinose. Sterna yellow pilose.

Genetics

We recover a cluster in our NJ tree with the DNA barcodes of C. impressa and C. inarmata sp. nov., together with three barcodes of C. transcaucasica (BS = 99.7%). Although all included sequences of C. impressa cluster together (BS <90%), DNA barcodes of C. inarmata are placed in a group with the three sequences of C. transcaucasica and another barcode of C. inarmata (ZFMK-TIS-8009609) is resolved sister to all other barcodes in this large cluster.

Remarks

We consider all published records of C. impressa from the Caucasus as belonging to C. inarmata sp. nov. and therefore C. impressa itself is removed from the Caucasus checklist.

Biology

Collected in mountains between 980–2887 m a.s.l. Behaviour similar to that of C. impressa .

Distribution

Common in the Caucasus, known from Armenia, Georgia and Russia.