Syzeuctus tenuifasciatus SCHMIEDEKNECHT , 1900

Riedel, Matthias, 2021, The genus Syzeuctus FÖRSTER in the Western Palaearctic region and Central Asia (Hymenoptera, Ichneumonidae, Banchinae), with descriptions of 17 new species, Linzer biologische Beiträge 52 (2), pp. 1013-1098 : 1070-1071

Syzeuctus tenuifasciatus SCHMIEDEKNECHT, 1900 (Figs 90, 136)

O r i g i n a l: Syzeuctus tenuifasciatus SCHMIEDEKNECHT, 1900: 345 , new name for S. punctiventris THOMSON.

syn. Lissonota punctiventris THOMSON, 1894: 2128 , preoccupied by THOMSON, 1877.

H o l o t y p e: (♀) "29/VI Triest" (Lund), type not studied.

syn. nov. Syzeuctus quadripunctorius KISS, 1929

H o l o t y p e: (♀) "Száskézd Silbernagel; Typus; Syzeuctus maculatorius F. var. 4-punctorius KISS " (Budapest), type studied .

syn. nov. Syzeuctus rufipes KISS, 1933

H o l o t y p e: (♂) " Budapest Kamaraedö 7.IX.927 A. Müller; Typus; Syzeuctus maculatorius F. v. rufipes KISS " (Budapest), type studied .

S t u d i e d m a t e r i a l: 56♀♀, 58♂♂; Austria, Bulgaria, Croatia, Czech Republic, Germany, Greece *, Hungary, Italy, Kazakhstan *, Kyrgyzstan *, Serbia, Slovakia, Switzerland *, Turkey,

Uzbekistan *. D e s c r i p t i o n: ♀: Body length 8-12.5 mm. Antenna with 33-42 flagellomeres ; 1st

flagellomere 4.5-5.0× longer than wide and 0.6× as long as eye; preapical flagellomere 1.0-1.1× longer than wide. Temple short, strongly and roundly narrowed behind eye, dorsally 0.25-0.35× as long as eye. OED 1.6-2.2× and OOD 2.8× ocellar diameter. Frons strongly impressed medially, sides with distinct swellings and sometimes ± distinct transverse ridge or lamella, with dense punctures. Face densely punctate, inner eye margins slightly convergent ventrally. Clypeus 1.5-1.7× wider than long, with very few setiferous punctures, apical margin rounded. MI 0.6-0.75. Genal carina reaching mandibular base.

Mesosoma 1.35-1.4× longer than wide. Mesoscutum and mesopleuron with rather dense punctures. Frontoventral edge of pronotum strongly curved, lateral pronotal groove finely rugose or ± punctate. Speculum punctate. Epicnemial carina low, with dentiform lamella on transversal part. Propodeum coarsely and densely punctate, with strong and complete apical transversal carina. Hind femur 5.0-5.7× longer than wide. Hind metatarsus 0.6-0.75× as long as hind femur. 2 nd hind tarsomere 1.4-1.45× longer than 3 rd one. Hind claw with short teeth, moderately bent apically. Areolet stalked; 2 nd recurrent vein in or slightly distal to its middle. Nervulus postfurcal by 1× its width. Hairs on basal cell about as long as their intervals. Hind wing with 10 distal hamuli.

Metasoma slender. 1 st tergite 1.55-1.9× longer than wide, with coarse but scattered lateral punctures. 2 nd tergite 1.0-1.3× as long as wide, densely punctate. 3 rd tergite 1.0- 1.4× as long as wide, densely punctate. Ovipositor 1.1-1.5× longer than fore wing, 1.1- 1.3× longer than metasoma and 2.4-2.9× longer than hind tibia.

Color: Black. Flagellum black, sometimes reddish-brown distally. Palps, clypeus in apical 2/3 (sometimes reddish), wider inner and narrower outer orbits, sometimes frontal spot on upper margin of pronotum, anterolateral and sometimes central spots on mesoscutum, sometimes spot on mesopleuron, tegula, sometimes spot on subtegular ridge, scutellum apically or mainly except anteromedian black spot, often postscutellum, and sometimes spot on propodeum yellow. Metasoma black, sometimes 1 st tergite with subbasal yellow spot and 2 nd and 3 rd tergites with narrow apical yellow bands. Coxae and trochanters black; sometimes fore coxa and trochanter with yellow spot; fore trochantellus reddish. Legs otherwise reddish; hind tibia often yellow basally and sometimes darkened in apical 0.3; hind tarsus ± brown or black, hind metatarsus often ± yellow basally. Wings with apical brown stripes (sometimes weak); pterostigma reddish-brown.

♂: Body length 7-11 mm. Antenna with 35-40 flagellomeres; 1 st flagellomere 4.5× longer than wide, preapical flagellomere 1.1-1.2× longer than wide. Temple shortly parallel behind eye, moderately narrowed apically, dorsally 0.4× as long as eye. OED 1.4-1.8× ocellar diameter. Clypeus 1.7× wider than long. MI 0.75. Lateral pronotal groove smooth ventrally. Speculum partly with scattered punctures. Epicnemial carina low, with strong denticular lamella on transversal part. Apical transversal carina of propodeum complete. Hind femur 3.9-5.5× longer than wide. Hind metatarsus 0.7× as long as hind femur. 2 nd hind tarsomere 1.45× longer than 3 rd one. Areolet stalked; 2 nd recurrent vein in the middle. Nervulus postfurcal by 1× its width.

1 st tergite 1.9-2.1× longer than wide, with scattered punctures. 2 nd tergite 1.05-1.1× longer than wide, with fine scattered or rather dense punctures. 3 rd tergite 1.0-1.3× longer than wide, with rather dense punctures.

Color: Black. Flagellum reddish, darker basally. Palps, mandible, clypeus, face (often except black central or paramedian stripes), malar space, wide orbits, and scape ventrally yellow. Mesosoma with extended yellow coloration: at least spot on collare, anterolateral spot on mesoscutum, tegula, subtegular ridge, spot on mesopleuron, apical scutellum and postscutellum and median spot on propodeum yellow. 1 st to 4 th tergites with apical yellow bands; 1 st tergite often with subbasal yellow spot. Fore and mid coxae and trochanters brown, with yellow spots; hind coxa and trochanters black. Fore and mid legs largely yellow, ± reddish-yellow caudally; hind femur red; hind tibia reddish-yellow, dark in apical 0.2; hind tarsus brown in distal 2/3. Fore wing with brown apical spot, pterostigma yellowish-red.

Hosts unknown.

R e m a r k: The separation of this species from the closely related Syzeuctus fuscator (PANZER) which is used here is in accordance with SCHMIEDEKNECHT (1907: 1262), but differs from the suggestions made by HORSTMANN 1999: 53. In HORSTMANN´ s view, the taxon S. fuscator (PANZER) contains specimens with very variable coloration, especially of hind legs, and therefore he included many specimens into his concept of S. fuscator that are determined as S. tenuifasciatus here.

In my opinion, all specimens with yellow-red or reddish hind femora (which have the same coloration as the mid femora) belong to S. tenuifasciatus whereas specimens with reddish-brown or blackish hind femora (with or without apical yellow margins and always darker than the mid femora) belong to S. fuscator . If one accepts this separation, the types of S. quadripunctorius and S. rufipes should be transferred to S. tenuifasciatus (syn. nov.)