Lasioglossum (Dialictus) deludens, Gardner & Gibbs, 2023

Gardner, Joel & Gibbs, Jason, 2023, Revision of the Nearctic species of the Lasioglossum (Dialictus) gemmatum species complex (Hymenoptera: Halictidae), European Journal of Taxonomy 858 (1), pp. 1-222 : 32-52

Lasioglossum (Dialictus) deludens sp. nov.

urn:lsid:zoobank.org:act:657A63A6-BF79-4EF1-A2B3-F2BE12AC0E6D

Figs 12–14 , 76C , 78D , 102C

Dialictus deludens (nomen nudum) – De Luca et al. 2019: 4879.

Diagnosis

Females of L. deludens sp. nov. have the tegula relatively small (barely reaching posterior margin of mesoscutum in dorsal view if at all), translucent yellow-orange, impunctate, and with inner posterior margin straight; mesoscutum punctures smaller and finer in comparison to mesepisternum punctures, dense (IS <1 PD) and becoming moderately sparse anteromedially and usually submedially (IS = 1–3 PD); mesoscutum usually becoming shiny posteriorly; metapostnotum shiny with strong anastomosing rugae; mesepisternum densely punctate (IS <1 PD) and usually shiny; paraocular area with dense tomentum; and metasoma black to brown (rarely somewhat reddish, but never bright red-orange).

Females of L. deludens sp. nov. are most similar to those of L. profundum sp. nov., L. rufodeludens sp. nov., and some L. diabolicum sp. nov. Females of L. profundum have the tegula dark brown, mesoscutum punctures about as large and coarse as mesepisternum punctures, and mesepisternum imbricate. Females of L. rufodeludens have the mesoscutum and mesepisternum completely dull and metasoma red-orange. Females of L. diabolicum have the tegula with at least some punctures medially (may be difficult to see if the tegula is very pale yellow) and the mesoscutum punctures about as large and coarse as mesepisternum punctures.

Males of L. deludens sp. nov. have the same characters as for females, in addition to the face completely covered with dense appressed tomentum extending above eye emargination but not reaching median ocellus. They are most similar to those of L. profundum sp. nov. and L. rufodeludens sp. nov. Males of L. profundum have the tegula dark brown and, in direct comparison, the mesosoma and metasoma punctures are larger and coarser. Males of L. rufodeludens sp. nov. have the mesoscutum mostly or entirely tessellate and slightly more densely punctate (IS = 1–2 PD) and face with dense tomentum not extending above eye emargination.

An undescribed species unrelated to the L. gemmatum complex and occurring sympatrically with L. deludens sp. nov. is superficially similar in both sexes, but the tegula is small and ovoid and the propodeum lacks an oblique carina.

Etymology

The specific epithet ʻ deludens ʼ is the Latin present participle of ʻ deludo ʼ (deceive, dupe). It refers to this species’ frequent misidentification in the past. An appropriate translation would be the deceiving sweat bee.

Material examined

Holotype MEXICO – Sonora • ♀; 30 km E of Agua Prieta ; 31.30972° N, 109.25417° W; 20Apr. 2005; R.L. Minckley leg.; PCYU SBV047813 .

[Verbatim label: MEXICO: Sonora / 30 km east Agua Prieta / 31°18′35″N 109°15′15″W / 20 Apr 2005, RL Minckley / ex. blue bowl / Bw7. SBV 047813 // 1849C12 // HOLOTYPE / Lasioglossum (Dialictus) deludens Gardner and Gibbs ]

Paratypes

MEXICO – Coahuila • 1 ♂; Boquillas ; 29.1883° N, 102.9374° W; 19 Jun. 1992; Robert L. Minckley leg.; ex Phacelia ; SEMC . – Durango • 1 ♀; 37 km NW of Ceballos ; [26.78° N, 104.38° W]; elev. 1230 m; 16 Mar. 1992; R. W. Brooks leg.; ex Nama hispida ; SEMC SM0342993 • 1 ♀; 69 mi. N of Durango, Highway #31; [25.03° N, 104.65° W]; 29 Jul. 1956; R.E. Beer e t al. leg.; SEMC . – Sonora • 1 ♀; 30 km E of Agua Prieta ; 31.2136° N, 109.2136° W; 11 Aug. 2004; Robert L. Minckley leg.; RLM • 8 ♀♀; 30 km E of Agua Prieta ; 31.3086° N, 109.2731° W; 2 May 2001; Robert L. Minckley leg.; RLM SBV011441 , SBV011447 , SBV011449 , SBV011453 , SBV011462 , SBV011465 , SBV011466 , SBV011471 • 1 ♀; 30 km E of Agua Prieta ; 31.31° N, 109.2778° W; 2 May 2001; Robert L. Minckley leg.; RLM SBV011349 • 3 ♀♀; 30 km E of Agua Prieta ; 31.315° N, 109.2728° W; 2 May 2001; Robert L. Minckley leg.; RLM SBV011426 , SBV011429 , SBV011436 • 5 ♀♀; 30 km E of Agua Prieta ; 31.3167° N, 109.2783° W; 2 May 2001; Robert L. Minckley leg.; RLM SBV011357 , SBV011372 , SBV011381 , SBV011400 , SBV011405 • 1 ♀; 30 km E of Agua Prieta ; 31.3183° N, 109.2514° W; 18 May 2004; Robert L. Minckley leg.; RLM SBV024134 • 2 ♀♀; 30 km E of Agua Prieta ; 31.3183° N, 109.2689° W; 30 Mar. 2002; A. Romero leg.; RLM SBV013275 , SBV013301 • 1 ♀; 30 km E of Agua Prieta ; 31.3189° N, 109.2511° W; 30 Mar. 2002; A. Romero leg.; RLM SBV013237 • 11 ♀♀; 30 km E of Agua Prieta ; 31.3222° N, 109.2769° W; 2 May 2001; Robert L. Minckley leg.; RLM SBV011480 , SBV011481 , SBV011488 , SBV011490 , SBV011495 , SBV011498 , SBV011500 , SBV011507 , SBV011509 , SBV011510 , SBV011511 • 5 ♀♀; 30 km E of Agua Prieta ; 31.3331° N, 109.2272° W; 3 May 2001; Robert L. Minckley leg.; RLM SBV011556 , SBV011570 , SBV011575 , SBV011576 , SBV011583 • 1 ♀; 30 km E of Agua Prieta ; 31.3331° N, 109.265° W; 21 May 2001; Robert L. Minckley leg.; RLM SBV012912 • 1 ♂; ibid.; 21 May 2001; Robert L. Minckley leg.; RLM SBV012924 • 1 ♀; 30 km E of Agua Prieta ; 31.3333° N, 109.2653° W; 20 Apr. 2005; Robert L. Minckley leg.; RLM SBV047719 • 1 ♂; ibid.; 20 Apr. 2005; Robert L. Minckley leg.; RLM SBV047759 • 1 ♂; 30 km E of Agua Prieta, Rio San Bernardino and Highway 2; [31.312° N, 109.254° W]; 10 May 2002; Robert L. Minckley leg.; ex Erigeron sp. ; RLM SBV031273 • 1 ♂; 38.6 km south of Ciudad Obregon ; [27.13° N, 109.93° W]; 20 Mar. 1990; Robert L. Minckley leg.; SEMC • 2 ♀♀; 5 miles west of Santa Ana ; [30.54° N, 111.2° W]; 10 Jun. 1961; F.D. Parker leg.; UCDC • 2 ♀♀; Arrollo El Limón, Rancho Las Lajas , 26 km E of Alamos; 27.0672° N, 108.7245° W; elev. 427 m; 20–22 Jun. 2008; M.E. Irwin leg.; BBSL FDP754275 , FDP754288 • 1 ♀; Cócorit ; [27.58° N, 109.96° W]; 11 Jun. 1961; F.D. Parker leg.; UCDC • 3 ♂♂; Hermosillo ; [29.07° N, 110.96° W]; 18 Jun. 1961; F.D. Parker leg.; UCDC • 1 ♂; Municipio de Agua Prieta ; 31.2908° N, 109.2636° W; 16 Apr. 2005; A. Romero leg.; RLM SBV142487 • 1 ♂; Municipio de Agua Prieta ; 31.3164° N, 109.2881° W; 3 May 2007; Robert L. Minckley leg.; ex Prosopis velutina ; RLM SBV073513 • 1 ♂; ibid.; 10 May 2007; Robert L. Minckley leg.; ex Lycium andersonii wrightii ; RLM SBV074220 • 1 ♂; ibid.; 14 May 2007; Robert L. Minckley leg.; ex Lycium berlandieri ; RLM SBV074294 • 3 ♂♂; Municipio de Agua Prieta, Rancho Las Anitas ; 31.3139° N, 109.3142° W; 11 Aug. 2007; Robert L. Minckley leg.; RLM SBV076940 , SBV076982 , SBV077166 • 1 ♂; Municipio de Agua Prieta, Rancho Nogalitos ; 31.3118° N, 108.9862° W; 25 May 2008; Robert L. Minckley leg.; RLM SBV083792 • 1 ♀; Nogales ; [31.3° N, 110.94° W]; 30 Jun. 1941; USNM • 1 ♂; Rancho Puerta Blanca ; 31.3139° N, 109.2503° W; 29 Apr. 2007; Robert L. Minckley leg.; ex Physaria fendleri ; RLM SBV073048 • 1 ♂; Rancho Puerta Blanca ; 31.3222° N, 109.0797° W; 8 May 2007; Robert L. Minckley leg.; ex Opuntia engelmannii × phaeacantha; RLM SBV074046 • 1 ♂; ibid.; 18 May 2007; Robert L. Minckley leg.; ex Porophyllum gracile ; RLM SBV074434 • 1 ♂; ibid.; 25 May 2007; Robert L. Minckley leg.; ex Dasyliron wheeleri ; RLM SBV074699 • 1 ♀; Rancho Puerta Blanca ; 31.3251° N, 109.0916° W; 8 Apr. 2005; Robert L. Minckley e t al. leg.; ex Malacothrix fendleri ; RLM SBV143077 • 1 ♀; Rancho San Bernardino ; [31.31° N, 109.25° W]; 22 Jun. 2000; K. Toal e t al. leg.; RLM SBV001862 • 4 ♀♀; ibid.; 7 May 2001; Robert L. Minckley leg.; RLM SBV008452 , SBV008472 , SBV019814 , SBV019842 • 1 ♀; Rancho San Bernardino ; [31.32° N, 109.27° W]; 25 Aug. 2001; Robert L. Minckley leg.; RLM SBV137594 • 1 ♂; Rancho San Bernardino ; 31.3002° N, 109.2625° W; 3 May 2002; A. Romero leg.; RLM SBV066335 • 1 ♂; ibid.; 4 Jul. 2002; Robert L. Minckley leg.; RLM SBV064051 • 1 ♂; Rancho San Bernardino ; 31.3002° N, 109.2626° W; 19 Apr. 2002; Robert L. Minckley leg.; RLM SBV064280 • 1 ♂; Rancho San Bernardino ; 31.3071° N, 109.2568° W; 12 Jun. 2004; N. de la Torre leg.; ex Tamarix ramosissima ; RLM SBV054512 • 1 ♂; Rancho San Bernardino ; 31.3072° N, 109.2568° W; 3 Jun. 2004; A. Romero leg.; ex Tamarix ramosissima ; RLM SBV025649 • 1 ♂; ibid.; 12 Jun. 2004; N. de la Torre leg.; ex Tamarix ramosissima ; RLM SBV054540 • 2 ♂♂; ibid.; 12 Jun. 2004; N. de la Torre leg.; RLM SBV054335 , SBV054378 • 6 ♂♂; Rancho San Bernardino ; 31.3074° N, 109.2569° W; 12 Jun. 2004; N. de la Torre leg.; RLM SBV054389 , SBV054397 , SBV054399 , SBV054405 , SBV054431 , SBV054432 • 1 ♂; Rancho San Bernardino ; 31.3075° N, 109.2572° W; 16 Jun. 2004; A. Romero leg.; RLM SBV079866 • 5 ♀♀; Rancho San Bernardino ; 31.3083° N, 109.2736° W; 21 Apr. 2005; A. Romero leg.; PCYU SBV048073 , SBV048079 , SBV048082 , SBV048087 , SBV048088 • 9 ♀♀; ibid.; 21 Apr. 2005; A. Romero leg.; RLM SBV048075 , SBV048076 , SBV048078 , SBV048080 , SBV048081 , SBV048086 , SBV048091 , SBV048092 , SBV048093 • 1 ♂; ibid.; 12 Aug. 2006; Robert L. Minckley leg.; RLM SBV060698 • 1 ♂; Rancho San Bernardino ; 31.3086° N, 109.2731° W; 25 Apr. 2001; Robert L. Minckley leg.; RLM SBV209834 • 1 ♂; ibid.; 2 May 2001; Robert L. Minckley leg.; PCYU SBV011472 • 1 ♂; ibid.; 22 May 2001; Robert L. Minckley leg.; RLM SBV030151 • 1 ♀; ibid.; 19 Jul. 2001; Robert L. Minckley leg.; RLM SBV134259 • 1 ♂; ibid.; 6 May 2002; Robert L. Minckley leg.; RLM SBV068366 • 1 ♂; Rancho San Bernardino ; 31.3097° N, 109.2542° W; 19 Aug. 2003; A. Nelson leg.; RLM SBV034617 • 3 ♂♂; Rancho San Bernardino ; 31.3097° N, 109.2778° W; 19 Aug. 2003; A. Nelson leg.; RLM SBV034551 , SBV034572 , SBV034573 • 1 ♀; ibid.; 8 Aug. 2004; Robert L. Minckley leg.; ex Lepidium thurberi ; RLM SBV026385 • 1 ♀; ibid.; 10 Aug. 2004; Robert L. Minckley leg.; ex Chenopodium neomexicanum ; RLM SBV026469 • 1 ♀; Rancho San Bernardino ; 31.3099° N, 109.2778° W; 2 May 2001; Robert L. Minckley leg.; RLM SBV011344 • 2 ♂♂; ibid.; 11 May 2001; A. Romero leg.; RLM SBV029242 , SBV029256 • 1 ♂; ibid.; 11 Sep. 2001; Robert L. Minckley leg.; RLM SBV018150 • 1 ♂; ibid.; 19 May 2004; Robert L. Minckley leg.; RLM SBV024523 • 1 ♂; ibid.; 25 Jul. 2007; Robert L. Minckley leg.; RLM SBV075348 • 1 ♀; Rancho San Bernardino ; 31.3101° N, 109.2535° W; 11 May 2001; A. Romero leg.; RLM SBV029274 • 1 ♂; ibid.; 11 May 2001; A. Romero leg.; RLM SBV029268 • 1 ♀; ibid.; 20 Jun. 2001; Robert L. Minckley leg.; RLM SBV016859 • 1 ♂; ibid.; 19 Jul. 2001; Robert L. Minckley leg.; RLM SBV134195 • 2 ♀♀; ibid.; 1 Aug. 2004; Robert L. Minckley leg.; RLM SBV026094 , SBV026097 • 2 ♀♀; Rancho San Bernardino ; 31.3112° N, 109.2662° W; 2 May 2001; Robert L. Minckley leg.; RLM SBV011309 , SBV011316 • 1 ♂; ibid.; 8 Jul. 2001; Robert L. Minckley leg.; RLM SBV133170 • 1 ♂; ibid.; 6 May 2002; Robert L. Minckley leg.; RLM SBV068158 • 1 ♀; Rancho San Bernardino ; 31.3136° N, 109.2547° W; 18 Apr. 2001; Robert L. Minckley leg.; RLM SBV208414 • 1 ♀; ibid.; 6 May 2001; Robert L. Minckley leg.; RLM SBV012240 • 1 ♂; ibid.; 12 May 2001; Robert L. Minckley leg.; RLM SBV029363 • 4 ♂♂; ibid.; 7Aug. 2001; Robert L. Minckley leg.; RLM SBV135710 , SBV135711 , SBV135712 , SBV135744 • 1 ♀; ibid.; 31 Jul. 2004; Robert L. Minckley leg.; RLM SBV026070 • 1 ♂; Rancho San Bernardino ; 31.3139° N, 109.2503° W; 3 Jun. 2002; Robert L. Minckley leg.; RLM SBV016829 • 2 ♂♂; ibid.; 25 Aug. 2003; N. de la Torre leg.; RLM SBV035522 , SBV035523 • 1 ♂; Rancho San Bernardino ; 31.314° N, 109.2553° W; 6 Jun. 2004; A. Romero leg.; ex Tamarix ramosissima ; RLM SBV025428 • 1 ♂; ibid.; 10 Jun. 2004; A. Romero leg.; ex Tamarix ramosissima ; RLM SBV025742 • 1 ♀; ibid.; 10 Jun. 2004; N. de la Torre leg.; RLM SBV025709 • 2 ♂♂; ibid.; 10 Jun. 2004; N. de la Torre leg.; RLM SBV025696 , SBV025697 • 1 ♀; Rancho San Bernardino ; 31.315° N, 109.2729° W; 2 Jul. 2001; Robert L. Minckley leg.; RLM SBV017752 • 1 ♀; ibid.; 19 Jul. 2001; Robert L. Minckley leg.; RLM SBV134222 • 5 ♀♀; Rancho San Bernardino ; 31.3159° N, 109.2686° W; 9 May 2006; Robert L. Minckley leg.; RLM SBV059140 , SBV059142 , SBV059148 , SBV059152 , SBV059153 • 1 ♀; Rancho San Bernardino ; 31.316° N, 109.2426° W; 20 Apr. 2001; Robert L. Minckley leg.; RLM SBV208990 • 2 ♀♀; ibid.; 28 Apr. 2001; Robert L. Minckley leg.; RLM SBV210324 , SBV210325 • 1 ♀; Rancho San Bernardino ; 31.3162° N, 109.1872° W; 11 May 2005; Robert L. Minckley leg.; ex Calycoseris wrightii ; RLM SBV049278 • 3 ♀♀; ibid.; 24 May 2005; Q. McFrederick leg.; RLM SBV050681 , SBV050683 , SBV050692 • 1 ♀; Rancho San Bernardino ; 31.3166° N, 109.2784° W; 1 Apr. 2002; A. Romero leg.; RLM SBV014957 • 2 ♂♂; Rancho San Bernardino ; 31.3169° N, 109.2592° W; 6 Jun. 2004; N. de la Torre leg.; ex Tamarix ramosissima ; RLM SBV025514 , SBV025552 • 1 ♂; ibid.; 10 Jun. 2004; N. de la Torre leg.; ex Tamarix ramosissima ; RLM SBV025786 • 1 ♀; Rancho San Bernardino ; 31.3179° N, 109.2602° W; 10 Jun. 2004; Robert L. Minckley leg.; RLM SBV025669 • 1 ♂; ibid.; 10 Jun. 2004; Robert L. Minckley leg.; RLM SBV025656 • 1 ♂; Rancho San Bernardino ; 31.318° N, 109.2603° W; 29 May 2004; N. de la Torre leg.; RLM SBV025090 • 1 ♀; Rancho San Bernardino ; 31.3183° N, 109.2514° W; 31 Jul. 2004; Robert L. Minckley leg.; RLM SBV026036 • 1 ♀; Rancho San Bernardino ; 31.3189° N, 109.2511° W; 21 May 2001; A. Romero leg.; RLM SBV012662 • 1 ♂; ibid.; 10 Aug. 2001; A. Romero leg.; RLM SBV016885 • 1 ♂; ibid.; 10 Aug. 2001; Robert L. Minckley leg.; RLM SBV135906 • 1 ♀; Rancho San Bernardino ; 31.3194° N, 109.2714° W; 2 Sep. 2003; A. Romero leg.; PCYU SBV036580 • 1 ♀; Rancho San Bernardino ; 31.3194° N, 109.2797° W; 11 May 2001; Robert L. Minckley leg.; RLM SBV013083 • 1 ♂; ibid.; 20 Jun. 2001; Robert L. Minckley leg.; RLM SBV030925 • 1 ♂; ibid.; 19 Jul. 2001; Robert L. Minckley leg.; RLM SBV134250 • 1 ♀; ibid.; 1 Apr. 2002; A. Romero leg.; RLM SBV015268 • 2 ♀♀; Rancho San Bernardino ; 31.3194° N, 109.2842° W; 27 Apr. 2005; Robert L. Minckley leg.; ex Eriastrum diffusum ; RLM SBV048511 , SBV048614 • 1 ♂; ibid.; 7 May 2005; Robert L. Minckley leg.; ex Cryptantha sp. ?; RLM SBV049099 • 1 ♂; ibid.; 7 May 2005; Robert L. Minckley leg.; ex Eriogonum abertianum ; RLM SBV049201 • 1♀; ibid.; 7 May2005; Robert L. Minckley leg.; ex Prosopis velutina ; RLM SBV049142 • 2 ♀♀; ibid.; 19 May 2005; Robert L. Minckley leg.; ex Cryptantha sp. ?; RLM SBV049789 , SBV049794 • 12 ♂♂; ibid.; 19 May 2005; Robert L. Minckley leg.; ex Cryptantha sp. ?; RLM SBV049786 , SBV049791 , SBV049792 , SBV049795 , SBV049796 , SBV049798 , SBV049799 , SBV049801 , SBV050054 , SBV050057 , SBV050059 , SBV050061 • 3 ♂♂; ibid.; 19 May 2005; Robert L. Minckley leg.; ex Eriogonum abertianum ; RLM SBV049893 , SBV049944 , SBV049949 • 1 ♂; ibid.; 19 May 2005; Robert L. Minckley leg.; ex Lepidium thurberi ; RLM SBV049925 • 2 ♂♂; ibid.; 19 May 2005; Robert L. Minckley leg.; ex Prosopis velutina ; RLM SBV050009 , SBV050020 • 1 ♂; ibid.; 19 May 2005; Robert L. Minckley leg.; RLM SBV049787 • 2 ♂♂; Rancho San Bernardino ; 31.3195° N, 109.2703° W; 8 May 2001; Robert L. Minckley leg.; ex Cryptantha sp. ?; RLM SBV019636 , SBV019637 • 1 ♂; Rancho San Bernardino ; 31.3196° N, 109.2693° W; 18 Aug. 2003; A. Nelson leg.; RLM SBV035627 • 4 ♂♂; Rancho San Bernardino ; 31.3196° N, 109.2703° W; 27 May 2004; B. O’Fallon leg.; RLM SBV025326 , SBV025361 , SBV025396 , SBV025405 • 1 ♂; Rancho San Bernardino ; 31.3197° N, 109.1933° W; 16 May 2005; Robert L. Minckley leg.; ex Malacothrix fendleri ; RLM SBV050065 • 1 ♂; ibid.; 30 May 2005; Robert L. Minckley leg.; ex Prosopis velutina ; RLM SBV052997 • 1 ♀; Rancho San Bernardino ; 31.3208° N, 109.276° W; 16 Jul. 2005; N.M. Williams leg.; ex Salsola tragus ; RLM SBV050983 • 1 ♀; Rancho San Bernardino ; 31.3211° N, 109.2114° W; 20 Apr. 2001; Robert L. Minckley leg.; RLM SBV208984 • 2 ♀♀; Rancho San Bernardino ; 31.3222° N, 109.2769° W; 25 Apr. 2001; Robert L. Minckley leg.; RLM SBV209863 , SBV209866 • 1 ♂; ibid.; 2 Sep. 2001; Robert L. Minckley leg.; RLM SBV138810 • 2 ♀♀; Rancho San Bernardino ; 31.3222° N, 109.2772° W; 21 May 2001; A. Romero leg.; RLM SBV012750 , SBV012803 • 1 ♀; ibid.; 22 Aug. 2001; Robert L. Minckley leg.; RLM SBV137203 • 1 ♂; Rancho San Bernardino ; 31.3222° N, 109.2778° W; 6 Aug. 2006; Robert L. Minckley leg.; ex Laennecia coulteri ; RLM SBV062228 • 1 ♂; ibid.; 15 Aug. 2006; Robert L. Minckley leg.; ex Laennecia coulteri ; RLM SBV062409 • 1 ♂; Rancho San Bernardino ; 31.3222° N, 109.2931° W; 2 May 2001; Robert L. Minckley leg.; RLM SBV139104 • 1 ♂; ibid.; 19 Aug. 2003; A. Romero leg.; RLM SBV034101 • 1 ♀; ibid.; 21 Apr. 2005; A. Romero leg.; RLM SBV047945 • 1 ♀; Rancho San Bernardino ; 31.3239° N, 109.2686° W; 1 Aug. 2005; Robert L. Minckley leg.; ex Salsola tragus ; RLM SBV053278 • 1 ♂; Rancho San Bernardino ; 31.3241° N, 109.2664° W; 20 Oct. 2004; Robert L. Minckley leg.; RLM SBV026763 • 1 ♂; ibid.; 28 Jul. 2006; A.A. Rahi leg.; ex Physalis acutifolia ; RLM SBV059761 • 1 ♀; Rancho San Bernardino ; 31.3242° N, 109.2666° W; 30 Apr. 2001; Robert L. Minckley leg.; RLM SBV011029 • 1 ♂; ibid.; 22 Aug. 2001; Robert L. Minckley leg.; RLM SBV137184 • 1 ♂; ibid.; 31 Aug. 2001; Robert L. Minckley leg.; RLM SBV138340 • 1 ♂; ibid.; 8 Sep. 2001; Robert L. Minckley leg.; RLM SBV139554 • 1 ♂; ibid.; 24 Jul. 2005; N.M. Williams leg.; ex Physalis acutifolia ; RLM SBV052135 • 1 ♀; Rancho San Bernardino ; 31.325° N, 109.265° W; 26 May 2005; Robert L. Minckley leg.; ex Parkinsonia aculeata ; RLM SBV050940 • 2 ♀♀; Rancho San Bernardino ; 31.3253° N, 109.223° W; 5 May 2001; Robert L. Minckley leg.; RLM SBV011997 , SBV012026 • 1 ♀; Rancho San Bernardino ; 31.3273° N, 109.2636° W; 31 Mar. 2002; A. Romero leg.; RLM SBV014747 • 2 ♀♀; Rancho San Bernardino ; 31.3286° N, 109.2684° W; 11 May 2006; Robert L. Minckley leg.; RLM SBV059414 , SBV059430 • 2♀♀; Rancho San Bernardino ; 31.3291° N, 109.2655°W; 10Jul. 2000; K.Toal e t al. leg.; RLM SBV002284 , SBV002295 • 1 ♀; Rancho San Bernardino ; 31.3295° N, 109.2636° W; 1 Sep. 2001; Robert L. Minckley leg.; RLM SBV138387 • 2 ♀♀; ibid.; 31 Mar. 2002; Robert L. Minckley leg.; RLM SBV014467 , SBV014526 • 1 ♀; Rancho San Bernardino ; 31.3296° N, 109.2546° W; 27 Apr. 2001; Robert L. Minckley leg.; RLM SBV210203 • 1 ♂; ibid.; 27 Apr. 2001; Robert L. Minckley leg.; RLM SBV210199 • 4 ♀♀; ibid.; 5 May 2001; Robert L. Minckley leg.; RLM SBV012139 , SBV012186 , SBV012191 , SBV012418 • 3 ♂♂; ibid.; 7 Aug. 2001; Robert L. Minckley leg.; RLM SBV135749 , SBV135750 , SBV135751 • 3 ♂♂; Rancho San Bernardino ; 31.3297° N, 109.2558° W; 20 Aug. 2003; N. de la Torre leg.; RLM SBV035452 , SBV035473 , SBV035474 • 2 ♀♀; Rancho San Bernardino ; 31.3297° N, 109.2559° W; 3 Apr. 2002; A. Romero leg.; RLM SBV016137 , SBV016146 • 1 ♀; Rancho San Bernardino ; 31.3301° N, 109.2666° W; 6 May 2001; Robert L. Minckley leg.; RLM SBV029324 • 1 ♀; ibid.; 14 May 2001; Robert L. Minckley leg.; RLM SBV029746 • 1 ♂; Rancho San Bernardino ; 31.3302° N, 109.2601° W; 30 Apr. 2001; Robert L. Minckley leg.; RLM SBV011008 • 3 ♂♂; ibid.; 10 Aug. 2001; Robert L. Minckley leg.; RLM SBV135928 , SBV135930 , SBV135932 • 2 ♂♂; ibid.; 22 Aug. 2001; Robert L. Minckley leg.; RLM SBV137268 , SBV137288 • 1 ♀; Rancho San Bernardino ; 31.3306° N, 109.21° W; 5 Aug. 2004; Robert L. Minckley leg.; ex Salsola tragus ; RLM SBV037203 • 1 ♀; ibid.; 11 Aug. 2004; Robert L. Minckley leg.; ex Zinnia acerosa ; RLM SBV026701 • 1 ♀; Rancho San Bernardino ; 31.331° N, 109.2676° W; 1 Apr. 2006; Robert L. Minckley e t al. leg.; RLM SBV056383 • 1 ♀; Rancho San Bernardino ; 31.3311° N, 109.2661° W; 28 Apr. 2001; Robert L. Minckley leg.; RLM SBV139036 • 1 ♂; Rancho San Bernardino ; 31.3312° N, 109.2663° W; 21 May 2001; A. Romero leg.; RLM SBV029896 • 1 ♀; Rancho San Bernardino ; 31.3317° N, 109.2582° W; 13 May 2001; A. Romero leg.; RLM SBV029663 • 1 ♂; Rancho San Bernardino ; 31.3321° N, 109.2702° W; 1 Sep. 2001; Robert L. Minckley leg.; RLM SBV138537 • 1 ♂; ibid.; 10 Sep. 2001; Robert L. Minckley leg.; RLM SBV140236 • 5 ♀♀; ibid.; 31 Mar. 2002; Robert L. Minckley leg.; RLM SBV014562 , SBV014574 , SBV014584 , SBV014627 , SBV014645 • 1 ♂; ibid.; 12 Oct. 2002; A. Romero leg.; RLM SBV045036 • 1 ♂; ibid.; 5 Jul. 2005; Robert L. Minckley leg.; RLM SBV132892 • 2 ♀♀; Rancho San Bernardino ; 31.333° N, 109.2404° W; 5 May 2001; Robert L. Minckley leg.; PCYU SBV019684 , SBV019705 • 1 ♀; ibid.; 20 Aug. 2001; Robert L. Minckley leg.; RLM SBV136885 • 1 ♀; Rancho San Bernardino ; 31.333° N, 109.265° W; 6 May 2001; Robert L. Minckley leg.; RLM SBV012473 • 2 ♀♀; ibid.; 1 Aug. 2004; Robert L. Minckley leg.; RLM SBV026112 , SBV026114 • 1 ♀; ibid.; 21 Jul. 2005; N.M. Williams leg.; RLM SBV051759 • 1 ♀; Rancho San Bernardino ; 31.3333° N, 109.2403° W; 14 Apr. 2005; Robert L. Minckley leg.; ex Physaria gordonii ; RLM SBV046922 • 1 ♀; ibid.; 3 May 2005; Robert L. Minckley leg.; ex Phacelia crenulata ; RLM SBV048951 • 1 ♂; ibid.; 22 May 2005; Robert L. Minckley leg.; ex Cryptantha sp ?; RLM SBV050175 • 1 ♂; Rancho San Bernardino ; 31.3333° N, 109.2597° W; 20 Aug. 2003; Robert L. Minckley leg.; ex Baccharis sarothroides ; RLM SBV034744 • 3 ♀♀; ibid.; 7 Apr. 2006; Robert L. Minckley e t al. leg.; RLM SBV057021 , SBV057037 , SBV057053 • 1 ♀; ibid.; 13 May 2006; Robert L. Minckley leg.; RLM SBV059628 • 1 ♀; Rancho San Bernardino ; 31.3335° N, 109.2601° W; 19 Apr. 2001; Robert L. Minckley leg.; RLM SBV208319 • 1 ♂; ibid.; 20 May 2004; Robert L. Minckley leg.; RLM SBV024660 • 1 ♀; Rio San Bernardino and Highway 2, 28 km east of Agua Prieta ; 31.322° N, 109.266° W; 7 May 2001; Robert L. Minckley leg.; RLM SBV008588 • 1 ♀; Tónichi ; [28.6° N, 109.57° W]; 23 Apr. 1990; R. J. McGinley leg.; USNM .

UNITED STATES – Arizona • 1 ♀; Cochise Co., ~ 25 mi. S of Willcox, Highway 191; 31.9064° N, 109.8118° W; elev. 1342 m; 15 Aug. 2007; Jason J. Gibbs, C. Sheffield leg.; PCYU PYU-6814 • 1 ♀; Cochise Co., 1 mi. E of Apache ; [31.69° N, 109.11° W]; 17 Aug. 1995; Jerome G. Rozen, S.A. Budick leg.; AMNH • 1 ♀; Cochise Co., 1 mi. E of Douglas ; [31.347° N, 109.507° W]; 20 Aug. 2010; H. Campbell leg.; AMNH • 1 ♀; ibid.; 24 Aug. 2011; WRME • 1 ♀; Cochise Co., 10 mi. E of Douglas ; [31.35° N, 109.35° W]; 22 Aug. 1994; Jerome G. Rozen, John S. Ascher leg.; AMNH • 1 ♀; Cochise Co., 10 mi. N of Portal Foothills Road ; [32.04° N, 109.18° W]; 30 Aug. 2010; H. Campbell leg.; AMNH • 1 ♀; Cochise Co., 13 mi. S of Apache ; [31.55° N, 109.27° W]; 20 Aug. 2010; H. Campbell leg.; AMNH • 1 ♀; Cochise Co., 13 mi. SW of Apache ; [31.55° N, 109.27° W]; 24 Aug. 2011; WRME • 1 ♀; Cochise Co., 2 mi. E of Apache ; [31.69° N, 109.1° W]; 19 Aug. 1998; Jerome G. Rozen, K. Franklin leg.; AMNH • 1 ♀; Cochise Co., 2 mi. NE of Portal ; [31.93° N, 109.12° W]; 30 Aug. 1994; Gretchen LeBuhn leg.; AMNH • 1 ♂; Cochise Co., 2 mi. W of Fort Huachuca ; [31.55° N, 110.38° W]; 28 Jun. 1983; W.J. Pulawski leg.; CAS • 1 ♀; Cochise Co., 23 mi. SW of Apache at Silver Creek Road; 31.475° N, 109.395° W; 6 Sep. 2012; Jason J. Gibbs leg.; CUIC • 1 ♀; Cochise Co., 27 mi. E of Douglas ; [31.35° N, 109.06° W]; 26 Aug. 1995; Jerome G. Rozen, S.A. Budick leg.; AMNH • 1 ♀; Cochise Co., 3.45 mi. S by E of Willcox ; 32.2262° N, 109.7822° W; elev. 1273 m; 1 Sep. 2009; Terry L. Griswold leg.; BBSL985430 • 1 ♀; Cochise Co., 30 miles SE of Willcox ; [31.94° N, 109.47° W]; 25 Jul. 1970; W.F. Chamberlain leg.; TAMU-ENTO X1495276 • 1 ♀; Cochise Co., 37 km E of Douglas ; 31.3713° N, 109.1484° W; 28 Mar. 2009; W. Radke leg.; RLM SBV084592 • 1 ♀; Cochise Co., 4 mi. E of Willcox ; [32.25° N, 109.76° W]; 14 Aug. 1994; Jerome G. Rozen leg.; AMNH • 1 ♀; ibid.; 27 Aug. 2007; John S. Ascher, C. Dong leg.; AMNH • 1 ♀; ibid.; 27 Aug. 2007; PCYU • 1 ♂; ibid.; 24 Aug. 2010; H. Campbell leg.; AMNH • 2 ♀♀; Cochise Co., 4–5 mi. NE of Douglas ; [31.39° N, 109.49° W]; 26 Aug. 1995; Jerome G. Rozen, S.A. Budick leg.; AMNH • 10 ♀♀, 1 ♂; Cochise Co., 6 mi. NE of Douglas ; [31.41° N, 109.47° W]; 6 Sep. 2011; Jerome G. Rozen, Eli S. Wyman leg.; AMNH • 1 ♀; Cochise Co., 7 mile marker Portal Road; [31.9° N, 109.1° W]; 28 Aug. 2004; PCYU • 1 ♀; Cochise Co., Chiricahua Monument / Pinery Canyon Road fork; [32.01° N, 109.39° W]; 3 Sep. 2009; N. Erwin leg.; AMNH • 1 ♂; Cochise Co., Chiricahua Mountains, Desert Scrub Site ; [31.9° N, 109.3° W]; J.L. Neff, A. R. Moldenke leg.; CAS • 1 ♀, 4 ♂♂; Cochise Co., Community Road , 9 mi. NE of Portal; [32.01° N, 109.09° W]; 7 Sep. 2011; Jerome G. Rozen, Eli S. Wyman leg.; AMNH • 1 ♀; Cochise Co., Flys Peak Trail ; [31.874° N, 109.286° W]; 2 Jun. 1999; Laurence Packer leg.; PCYU • 1 ♀; Cochise Co., near Willcox, Blue Sky Road ; 32.232° N, 109.7781° W; elev. 1279 m; 14 Aug. 2007; Jason J. Gibbs, C. Sheffield leg.; PCYU • 1 ♀, 1 ♂; Cochise Co., Portal ; [31.91° N, 109.14° W]; 27 May 1983; W.J. Pulawski leg.; CAS • 2 ♀♀; ibid.; 28 May 1983; W.J. Pulawski leg.; CAS • 1 ♂; Cochise Co., San Bernardino National Wildlife Refuge ; [31.34° N, 109.26° W]; 24 Aug. 2001; Robert L. Minckley leg.; RLM SBV137438 • 1 ♀; ibid.; 27 Aug. 2001; Robert L. Minckley leg.; RLM SBV137723 • 1 ♂; Cochise Co., San Bernardino National Wildlife Refuge ; 31.334° N, 109.2372° W; 23 May 2001; Robert L. Minckley leg.; RLM SBV030550 • 1 ♂; ibid.; 5 Aug. 2003; A. Romero leg.; RLM SBV032144 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3342° N, 109.2332° W; 5 May 2001; Robert L. Minckley leg.; RLM SBV011905 • 1 ♂; ibid.; 12 May 2001; A. Romero leg.; RLM SBV029391 • 4 ♀♀; ibid.; 18 Jul. 2005; T. Pasqual leg.; RLM SBV051064 , SBV051076 , SBV051077 , SBV051088 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3342° N, 109.2367° W; 24 Apr. 2001; Robert L. Minckley leg.; RLM SBV209689 • 1 ♀; ibid.; 5 May 2001; Robert L. Minckley leg.; RLM SBV012278 • 1 ♀; ibid.; 12 May 2001; Robert L. Minckley leg.; RLM SBV013102 • 1 ♂; ibid.; 10 Jul. 2001; Robert L. Minckley leg.; RLM SBV133129 • 1 ♀; ibid.; 18 Jul. 2005; T. Pasqual leg.; RLM SBV051210 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3342° N, 109.2678° W; 1 May 2001; Robert L. Minckley leg.; RLM SBV011088 • 1 ♀; ibid.; 24 Aug. 2001; Robert L. Minckley leg.; RLM SBV137469 • 1 ♂; ibid.; 24 Aug. 2001; Robert L. Minckley leg.; RLM SBV137464 • 2 ♂♂; ibid.; 1 Sep. 2001; Robert L. Minckley leg.; RLM SBV138693 , SBV138694 • 4 ♂♂; ibid.; 10 Sep. 2001; Robert L. Minckley leg.; RLM SBV140255 , SBV140287 , SBV140297 , SBV140298 • 1 ♂; ibid.; 21 Sep. 2001; Robert L. Minckley leg.; RLM SBV019052 • 6 ♀♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3344° N, 109.2681° W; 1 May 2001; Robert L. Minckley leg.; RLM SBV011246 , SBV011249 , SBV011253 , SBV011268 , SBV011272 , SBV011278 • 1 ♀; ibid.; 10 May 2001; A. Romero leg.; RLM SBV028542 • 1 ♀; ibid.; 22 May 2001; Robert L. Minckley leg.; RLM SBV042187 • 1 ♂; ibid.; 22 May 2001; Robert L. Minckley leg.; RLM SBV042190 • 1 ♂; ibid.; 3 Aug. 2001; Robert L. Minckley leg.; RLM SBV134968 • 1 ♂; ibid.; 13 Aug. 2001; Robert L. Minckley leg.; RLM SBV136627 • 1 ♀; ibid.; 1 Sep. 2001; Robert L. Minckley leg.; RLM SBV138751 • 1 ♂; ibid.; 1 Sep. 2001; Robert L. Minckley leg.; RLM SBV138771 • 2 ♂♂; ibid.; 10 Sep. 2001; Robert L. Minckley leg.; RLM SBV140009 , SBV140016 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3345° N, 109.2524° W; 21 Aug. 2001; Robert L. Minckley leg.; RLM SBV137163 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3349° N, 109.2529° W; 22 Jun. 2001; Robert L. Minckley leg.; RLM SBV031450 • 4 ♂♂; ibid.; 7Aug. 2001; Robert L. Minckley leg.; RLM SBV135650 , SBV135682 , SBV135688 , SBV135692 • 1 ♂; ibid.; 21 Aug. 2001; Robert L. Minckley leg.; RLM SBV137040 • 4 ♂♂; ibid.; 29 Aug. 2001; Robert L. Minckley leg.; RLM SBV138068 , SBV138071 , SBV138078 , SBV138083 • 1 ♂; ibid.; 25 Sep. 2001; Robert L. Minckley leg.; RLM SBV019433 • 6 ♀♀; ibid.; 3 Apr. 2002; A. Romero leg.; RLM SBV016297 , SBV016304 , SBV016320 , SBV016337 , SBV016396 , SBV016398 • 3 ♀♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.335° N, 109.2539° W; 14 May 2004; Robert L. Minckley leg.; RLM SBV022974 , SBV023006 , SBV023008 • 3 ♀♀; ibid.; 31 Jul. 2004; Robert L. Minckley leg.; RLM SBV026041 , SBV026050 , SBV026052 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3351° N, 109.279° W; 24 Jul. 2004; R. G. Hatfield leg.; ex Leucophyllum frutescens ; RLM SBV055184 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3353° N, 109.2514° W; 13 Apr. 2001; Robert L. Minckley leg.; RLM SBV208488 • 1 ♀; ibid.; 21 May 2001; Robert L. Minckley leg.; RLM SBV012957 • 1 ♂; ibid.; 21 May 2001; Robert L. Minckley leg.; RLM SBV012985 • 2 ♂♂; ibid.; 22 Aug. 2001; Robert L. Minckley leg.; RLM SBV137207 , SBV137228 • 1 ♂; ibid.; 31Aug. 2001; Robert L. Minckley leg.; RLM SBV138310 • 2 ♀♀; ibid.; 8 Sep. 2001; Robert L. Minckley leg.; RLM SBV139965 , SBV139968 • 1 ♂; ibid.; 8 Sep. 2001; Robert L. Minckley leg.; RLM SBV139761 • 3 ♀♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3353° N, 109.2517° W; 19 Jul. 2005; A. Romero leg.; RLM SBV051568 , SBV051569 , SBV051571 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3353° N, 109.2668° W; 14 May 2001; A. Romero leg.; RLM SBV029725 • 1 ♂; ibid.; 10 Aug. 2001; Robert L. Minckley leg.; RLM SBV135785 • 1 ♂; ibid.; 22 Aug. 2001; Robert L. Minckley leg.; RLM SBV137235 • 3 ♀♀; ibid.; 31 Jul. 2004; Robert L. Minckley leg.; RLM SBV026077 , SBV026081 , SBV026089 • 3 ♀♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3353° N, 109.2681° W; 13 May 2004; Robert L. Minckley leg.; RLM SBV022438 , SBV022449 , SBV022456 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3353° N, 109.27° W; 20 Apr. 2005; A. Romero leg.; RLM SBV047699 • 3 ♀♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3357° N, 109.2737° W; 6 Apr. 2006; Robert L. Minckley et al. leg.; RLM SBV056602 , SBV056605 , SBV056631 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3358° N, 109.2629° W; 20 Apr. 2005; Robert L. Minckley leg.; RLM SBV047896 • 2 ♂♂; ibid.; 20 Apr. 2005; Robert L. Minckley leg.; RLM SBV047860 , SBV047878 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3358° N, 109.2702° W; 10 May 2001; A. Romero leg.; RLM SBV028531 • 1 ♂; ibid.; 22 May 2001; Robert L. Minckley leg.; RLM SBV030252 • 1 ♂; ibid.; 13 Aug. 2001; Robert L. Minckley leg.; RLM SBV136507 • 3 ♂♂; ibid.; 1 Sep. 2001; Robert L. Minckley leg.; RLM SBV138473 , SBV138482 , SBV138508 • 4 ♀♀; ibid.; 31 Mar. 2002; Robert L. Minckley leg.; RLM SBV013792 , SBV013796 , SBV013815 , SBV013830 • 3 ♀♀; ibid.; 13 May 2004; Robert L. Minckley leg.; RLM SBV022135 , SBV022150 , SBV022187 • 1 ♂; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3363° N, 109.2675° W; 23 Apr. 2001; A. Romero leg.; PCYU SBV209390 • 1 ♀; ibid.; 1 May 2001; Robert L. Minckley leg.; RLM SBV011223 • 1 ♂; ibid.; 26 Jun. 2001; A. Romero leg.; RLM SBV017613 • 1 ♂; ibid.; 5 Jul. 2001; Robert L. Minckley leg.; RLM SBV132918 • 2 ♀♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3364° N, 109.2675° W; 23 Apr. 2001; A. Romero leg.; RLM SBV209394 , SBV209400 • 1 ♂; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3369° N, 109.2436° W; 20 Aug. 2001; Robert L. Minckley leg.; RLM SBV136955 • 1 ♀; ibid.; 2 Apr. 2002; A. Romero leg.; RLM SBV015592 • 5 ♀♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3372° N, 109.2311° W; 3 May 2001; Robert L. Minckley leg.; RLM SBV011519 , SBV011542 , SBV011545 , SBV011548 , SBV011551 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3374° N, 109.231° W; 15 Apr. 2001; Robert L. Minckley leg.; RLM SBV006291 • 1 ♀; ibid.; 23 May 2001; Robert L. Minckley leg.; RLM SBV030369 • 1 ♀; ibid.; 24 Jul. 2001; Robert L. Minckley leg.; RLM SBV134599 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3375° N, 109.2625° W; 13 May 2001; A. Romero leg.; RLM SBV029658 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3381° N, 109.2605° W; 30 Apr. 2001; Robert L. Minckley leg.; RLM SBV010982 • 3 ♂♂; ibid.; 10 Aug. 2001; Robert L. Minckley leg.; RLM SBV136366 , SBV136368 , SBV136379 • 1 ♂; ibid.; 19 Sep. 2001; Robert L. Minckley leg.; RLM SBV018739 • 1 ♀; ibid.; 30 Mar. 2002; Robert L. Minckley leg.; RLM SBV013559 • 1 ♀; ibid.; 16 May 2004; Robert L. Minckley leg.; RLM SBV023798 • 4 ♀♀; ibid.; 30 Jul. 2004; A. Romero leg.; RLM SBV055370 , SBV055371 , SBV055382 , SBV055383 • 3 ♀♀; ibid.; 19 Jul. 2005; A. Langley leg.; RLM SBV051581 , SBV051583 , SBV051594 • 1 ♂; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3385° N, 109.2531° W; 29 Aug. 2001; Robert L. Minckley leg.; RLM SBV137988 • 1 ♀; ibid.; 14 May 2004; Robert L. Minckley leg.; RLM SBV022727 • 2 ♀♀; ibid.; 31 Jul. 2004; Robert L. Minckley leg.; RLM SBV026002 , SBV026014 • 2 ♀♀; ibid.; 18 Jul. 2005; A. Romero leg.; RLM SBV051428 , SBV051447 • 1 ♀; ibid.; 18 Jul. 2005; T. Pasqual leg.; RLM SBV051388 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3406° N, 109.2575° W; 6 May 2001; A. Romero leg.; RLM SBV028430 • 1 ♂; ibid.; 31 Aug. 2001; Robert L. Minckley leg.; RLM SBV138128 • 1 ♀; ibid.; 30 Mar. 2002; A. Romero leg.; RLM SBV013183 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3407° N, 109.253° W; 26 Apr. 2001; Robert L. Minckley leg.; RLM SBV209934 • 1 ♂; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3411° N, 109.2705° W; 4 May 2002; N. de la Torre leg.; RLM SBV066792 • 1 ♂; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3415° N, 109.2728° W; 22 Apr. 2002; Robert L. Minckley leg.; RLM SBV064909 • 1 ♂; ibid.; 15 Aug. 2002; Robert L. Minckley leg.; RLM SBV044453 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3417° N, 109.2528° W; 20 Apr. 2005; A. Romero leg.; RLM SBV047614 • 2 ♂♂; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3417° N, 109.2582° W; 31 Aug. 2001; Robert L. Minckley leg.; RLM SBV138148 , SBV138159 • 1 ♂; ibid.; 8 Sep. 2001; Robert L. Minckley leg.; RLM SBV140144 • 2 ♀♀; ibid.; 30 Jul. 2004; A. Romero leg.; RLM SBV055324 , SBV055340 • 1 ♀; ibid.; 19 Jul. 2005; A. Langley leg.; RLM SBV051491 • 2 ♀♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3425° N, 109.27° W; 23 Apr. 2001; Robert L. Minckley leg.; RLM SBV209375 , SBV209376 • 1 ♂; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3425° N, 109.2745° W; 25 Apr. 2008; Robert L. Minckley leg.; RLM SBV082423 • 3 ♀♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3426° N, 109.2701° W; 1 May 2001; Robert L. Minckley leg.; RLM SBV011126 , SBV011133 , SBV011173 • 1 ♂; ibid.; 1 May 2001; Robert L. Minckley leg.; RLM SBV011181 • 1 ♂; ibid.; 22 May 2001; Robert L. Minckley leg.; RLM SBV042177 • 2 ♂♂; ibid.; 5 Jul. 2001; Robert L. Minckley leg.; RLM SBV132596 , SBV132949 • 2 ♂♂; ibid.; 24 Aug. 2001; Robert L. Minckley leg.; RLM SBV137375 , SBV137390 • 1 ♂; ibid.; 12 Oct. 2002; A. Romero leg.; RLM SBV044986 • 3 ♀♀; ibid.; 13 May 2004; Robert L. Minckley leg.; RLM SBV022250 , SBV022262 , SBV022332 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3427° N, 109.2746° W; 1 May 2001; Robert L. Minckley leg.; RLM SBV011057 • 4 ♂♂; ibid.; 1 Sep. 2001; Robert L. Minckley leg.; RLM SBV138410 , SBV138411 , SBV138412 , SBV138460 • 1 ♀; ibid.; 13 May 2004; Robert L. Minckley leg.; RLM SBV022007 • 2 ♂♂; ibid.; 13 May 2004; Robert L. Minckley leg.; RLM SBV022002 , SBV022009 • 1 ♀; Cochise Co., San Bernardino National Wildlife Refuge ; 31.3428° N, 109.2747° W; 23 Apr. 2001; Robert L. Minckley leg.; RLM SBV209431 • 1 ♀; Cochise Co., Sunsites ; [31.94° N, 109.83° W]; 23 May 1992; W.F. Chamberlain leg.; TAMU-ENTO X1495775 • 1 ♀; Cochise Co., Willcox ; [32.25° N, 109.83° W]; 2005; Laurence Packer leg.; PCYU • 1 ♀; Graham Co., Roper Lake State Park 6 mi. S of Safford; [32.75° N, 109.7° W]; 26 May 1983; W.J. Pulawski leg.; CAS • 5 ♀♀; Maricopa Co., 1100 W Mission Drive, Chandler ; 33.346° N, 111.8599° W; elev. 367 m; 27 Mar. 2017; S.J. Hall et al. leg.; ASUHIC 0112713, 0112714, 0112717, 0113161, 0113165 • 1 ♀; Maricopa Co., 115 E Country Gables Drive, Phoenix ; 33.6223° N, 112.0718° W; elev. 412 m; 12 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 113541 • 10 ♀♀; ibid.; 18 May 2017; S.J. Hall et al. leg.; ASUHIC 0113089, 0113091, 0113092, 0113093, 0113094, 0113095, 0113100, 0113103, 0113106, 0113108 • 1 ♀; Maricopa Co., 1414 E Libra Drive, Tempe ; 33.3656° N, 111.9163° W; elev. 365 m; 3 May 2017; S.J. Hall et al. leg.; ASUHIC 112608 • 1 ♂; ibid.; 3 May 2017; S.J. Hall et al. leg.; ASUHIC 112455 • 1 ♂; Maricopa Co., 15 km W of Maricopa ; [33.06° N, 112.21° W]; 3 May 1994; Robert L. Minckley leg.; ex Larrea tridentata ; SEMC • 3 ♀♀; Maricopa Co., 1615 E Fremont Drive, Tempe ; 33.3823° N, 111.9128° W; elev. 361 m; 4 May 2017; S.J. Hall et al. leg.; ASUHIC 0114782, 0114786, 0114787 • 2 ♀♀; Maricopa Co., 1738 E Palmcroft Drive, Tempe ; 33.4044° N, 111.9099° W; elev. 361 m; 16 May 2017; S.J. Hall et al. leg.; ASUHIC 0113366, 0113377 • 2 ♂♂; ibid.; 16 May 2017; S.J. Hall et al. leg.; ASUHIC 0113390, 0113392 • 1 ♀; Maricopa Co., 1863 E Geneva Drive, Tempe ; 33.3945° N, 111.9058° W; elev. 362 m; 26 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 112673 • 1 ♂; ibid.; 26 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 112684 • 10 ♀♀; Maricopa Co., 1926 E Calle De Caballos, Tempe ; 33.3401° N, 111.9069° W; elev. 362 m; 10 May 2017; S.J. Hall et al. leg.; ASUHIC 0113142, 0113145, 0113146, 0113148, 0113150, 0113152, 0113153, 0113154, 0113156, 0113157 • 3 ♂♂; ibid.; 10 May 2017; S.J. Hall et al. leg.; ASUHIC 0113187, 0113196, 0113197 • 1 ♀; Maricopa Co., 2031 S Sierra Vista Drive, Tempe ; 33.406° N, 111.9308° W; elev. 357 m; 3 May 2017; S.J. Hall et al. leg.; ASUHIC 112733 • 1 ♀; Maricopa Co., 2721 E Downing Circle, Mesa ; 33.4287° N, 111.7723° W; elev. 390 m; 19 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 113170 • 1 ♀; ibid.; 11 May 2017; S.J. Hall et al. leg.; ASUHIC 113625 • 2 ♀♀; ibid.; 22 May 2017; S.J. Hall et al. leg.; ASUHIC 0113572, 0113574 • 12 ♀♀; Maricopa Co., 2814 N 11 th Avenue, Phoenix ; 33.4801° N, 112.0868° W; elev. 335 m; 4 May 2017; S.J. Hall et al. leg.; ASUHIC 0113504, 0113506, 0113509, 0113514, 0113522, 0113525, 0113527, 0113530, 0113534, 0113535, 0113538, 0113539 • 3 ♂♂; ibid.; 4 May 2017; S.J. Hall et al. leg.; ASUHIC 0113578, 0113588, 0113589 • 10 ♀♀; ibid.; 22 May 2017; S.J. Hall et al. leg.; ASUHIC 0112824, 0112834, 0112837, 0112838, 0112841, 0112842, 0112843, 0112853, 0112854, 0112860 • 9 ♀♀; Maricopa Co., 302 E Fairmont Drive, Tempe ; 33.3964° N, 111.9346° W; elev. 358 m; 30 Mar. 2017; S.J. Hall et al. leg.; ASUHIC 0113443, 0113444, 0113445, 0113447, 0113451, 0113453, 0113454, 0113455, 0113457 • 14 ♀♀; ibid.; 27 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 0113461, 0113462, 0113463, 0113466, 0113468, 0113469, 0113470, 0113471, 0113480, 0113482, 0113483, 0113484, 0113485, 0113490 • 1 ♀; Maricopa Co., 3346 E Downing Street, Mesa ; 33.4293° N, 111.7585° W; elev. 395 m; 3 May 2017; S.J. Hall et al. leg.; ASUHIC 112592 • 1 ♀; Maricopa Co., 8201 E Windsor Avenue, Scottsdale ; 33.4783° N, 111.9038° W; elev. 373 m; 19 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 113567 • 5 ♀♀; ibid.; 11 May 2017; S.J. Hall et al. leg.; ASUHIC 0112599, 0113114, 0113116, 0113130, 0113140 • 4 ♀♀; Maricopa Co., 8433 E Palm Lane, Scottsdale ; 33.4686° N, 111.899° W; elev. 369 m; 19 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 0113546, 0113548, 0113549, 0113551 • 5 ♀♀; ibid.; 16 May 2017; S.J. Hall et al. leg.; ASUHIC 0112624, 0112635, 0112636, 0112647, 0112648 • 1 ♀; Maricopa Co., 862 E 9 th Place, Mesa ; 33.4321° N, 111.812° W; elev. 381 m; 11 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 112686 • 17 ♀♀; Maricopa Co., Estrella Mountain Regional Park; 33.3617° N, 112.3414° W; elev. 316 m; 29 Mar. 2017; S.J. Hall et al. leg.; ASUHIC 0112967, 0112969, 0112970, 0112973, 0112975, 0112976, 0112980, 0112981, 0112982, 0112983, 0112988, 0113412, 0113414, 0113418, 0113419, 0113420, 0113421 • 2 ♂♂; ibid.; 29 Mar. 2017; S.J. Hall et al. leg.; ASUHIC 0112567, 0112568 • 7 ♀♀; Maricopa Co., McDowell Mountain Regional Park; 33.6423° N, 111.7061° W; elev. 511 m; 4 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 0113422, 0113423, 0113424, 0113425, 0113428, 0113431, 0113432 • 1 ♂; ibid.; 4 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 113426 • 3 ♀♀; ibid.; 16 May 2017; S.J. Hall et al. leg.; ASUHIC 0112918, 0113009, 0113437 • 1 ♂; ibid.; 16 May 2017; S.J. Hall et al. leg.; ASUHIC 113435 • 11 ♀♀; Maricopa Co., McDowell Sonoran Preserve ; 33.644° N, 111.8567° W; elev. 516 m; 12 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 0112456, 0113288, 0113289, 0113290, 0113291, 0113293, 0113295, 0113297, 0113298, 0113300, 0113302 • 3 ♀♀; ibid.; 16 May 2017; S.J. Hall et al. leg.; ASUHIC 0113000, 0113002, 0113007 • 1 ♂; ibid.; 16 May 2017; S.J. Hall et al. leg.; ASUHIC 113010 • 19 ♀♀; Maricopa Co., Mountain View Park ; 33.5787° N, 112.0799° W; elev. 401 m; 7 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 0112875, 0112877, 0112880, 0112881, 0112882, 0112883, 0112887, 0112889, 0112890, 0112892, 0112893, 0112895, 0112896, 0112900, 0112901, 0112902, 0112903, 0112904, 0112906 • 4 ♂♂; ibid.; 7 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 0112879, 0113011, 0113012, 0113013 • 8 ♀♀; ibid.; 18 May 2017; S.J. Hall et al. leg.; ASUHIC 0112922, 0112923, 0112924, 0112931, 0112942, 0112945, 0112950, 0112962 • 5 ♀♀; Maricopa Co., Papago Park ; 33.4571° N, 111.9424° W; elev. 391 m; 30 Mar. 2017; S.J. Hall et al. leg.; ASUHIC 0112466, 0112558, 0112561, 0113236, 0113241 • 4 ♂♂; ibid.; 30 Mar. 2017; S.J. Hall et al. leg.; ASUHIC 0112557, 0112559, 0113234, 0113235 • 44 ♀♀; ibid.; 3 May 2017; S.J. Hall et al. leg.; ASUHIC 0112458, 0112462, 0112463, 0112464, 0112467, 0112474, 0112476, 0112477, 0112478, 0112481, 0112482, 0112486, 0112493, 0112495, 0112496, 0112498, 0112502, 0112505, 0112507, 0112508, 0112513, 0112519, 0112520, 0112522, 0112523, 0112525, 0112534, 0112535, 0112537, 0112538, 0112541, 0112544, 0112545, 0112547, 0112552, 0112553, 0112556, 0112565, 0113242, 0113244, 0113248, 0113250, 0113258, 0113259 • 2 ♂♂; ibid.; 3 May 2017; S.J. Hall et al. leg.; ASUHIC 0112485, 0113254 • 2 ♀♀; Maricopa Co., Papago Park ; 33.4571° N, 111.9425° W; elev. 391 m; 24 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 0112822, 0113327 • 2 ♀♀; ibid.; 8 Jun. 2017; S.J. Hall et al. leg.; ASUHIC 0113200, 0113225 • 5 ♀♀; Maricopa Co., Phoenix Mountain Preserve ; 33.5679° N, 112.0121° W; elev. 452 m; 5 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 0113307, 0113308, 0113309, 0113310, 0113311 • 1 ♂; ibid.; 5 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 112756 • 15 ♀♀; ibid.; 18 May 2017; S.J. Hall et al. leg.; ASUHIC 0113015, 0113018, 0113020, 0113024, 0113027, 0113035, 0113044, 0113045, 0113049, 0113050, 0113051, 0113053, 0113067, 0113069, 0113082 • 1 ♀; Maricopa Co., South Mountain Park ; 33.3516° N, 112.1023° W; elev. 394 m; 18 May 2017; S.J. Hall et al. leg.; ASUHIC 112814 • 4 ♀♀; Maricopa Co., South Mountain Park ; 33.3565° N, 111.9891° W; elev. 406 m; 22 Mar. 2017; S.J. Hall et al. leg.; ASUHIC 0113266, 0113405, 0113406, 0113407 • 3 ♂♂; ibid.; 22 Mar. 2017; S.J. Hall et al. leg.; ASUHIC 0112819, 0113265, 0113267 • 14 ♀♀; ibid.; 12 Apr. 2017; S.J. Hall et al. leg.; ASUHIC 0112758, 0112759, 0112760, 0112764, 0112767, 0112768, 0112769, 0112770, 0112771, 0113269, 0113270, 0113400, 0113403, 0113411 • 7 ♀♀; ibid.; 10 May 2017; S.J. Hall et al. leg.; ASUHIC 0112774, 0112777, 0112781, 0112789, 0112797, 0112801, 0112802 • 1 ♂; ibid.; 10 May 2017; S.J. Hall et al. leg.; ASUHIC 112809 • 1 ♀; Maricopa Co., Usery Mountain Regional Park; 33.4696° N, 111.6171° W; elev. 588 m; 20 Mar. 2017; S.J. Hall et al. leg.; ASUHIC 112707 • 1 ♀; Maricopa Co., White Tanks Mountain Regional Park ; 33.6049° N, 112.4964° W; elev. 448 m; 22 Mar. 2017; S.J. Hall et al. leg.; ASUHIC 113276 • 2 ♀♀; ibid.; 8 Jun. 2017; S.J. Hall et al. leg.; ASUHIC 0113317, 0113318 • 3 ♀♀; Mohave Co., 10 mi. S of Lake Havasu City ; [34.38° N, 114.2° W]; 14 Mar. 1972; P.F. Torchio, B. Apperson leg.; ex Phacelia ; BBSL700790 , BBSL700791 , BBSL700793 • 2 ♀♀; Mohave Co., 15 mi. E of Oatman ; [35.03° N, 114.12° W]; 18 Oct. 1973; F.D. Parker, P.F. Torchio leg.; BBSL700785 , BBSL700786 • 1 ♀; Pima Co., 2 km NE of Oracle ; 32.626° N, 110.7362° W; elev. 1240 m; 20–24 May 2013; M.E. Irwin leg.; BBSL FDP152748 • 1 ♂; Pima Co., 7 km E of Robles Junction ; 32.0667° N, 111.25° W; elev. 790 m; 22 Apr. 1995; Robert L. Minckley, R. Gyuro leg.; ex Prosopis velutina ; SEMC • 1 ♂; Pima Co., 7 km E of Silverbell ; 32.3667° N, 111.4333° W; elev. 732 m; 10 Apr. 1994; Robert L. Minckley leg.; ex Sphaeralcea sp. ; SEMC • 1 ♀; Pima Co., Gardner Canyon Site ; [31.74° N, 110.7° W]; J.L. Neff leg.; CAS • 1 ♀; Pima Co., Madera Canyon ; [31.74° N, 110.89° W]; 21 Sep. 1982; W.J. Pulawski leg.; CAS • 4 ♀♀, 1 ♂; Pima Co., Mount Lemmon ; [32.4° N, 110.8° W]; elev. 1067 m; J.L. Neff leg.; CAS • 2 ♀♀; Pima Co., Sandario Road 7 km N of Highway 86; 32.1833° N, 111.2167° W; elev. 740 m; 2 Apr. 1994; Robert L. Minckley, S. G. Reyes leg.; ex Sphaeralcea ; SEMC • 1 ♂; Pima Co., Santa Catalina Mountains, Desert Scrub site; [32.4° N, 110.8° W]; J.L. Neff leg.; CAS • 40 ♀♀, 5 ♂♂; Pima Co., Silver Bell Bajada, Int. BioL. Program ; [32.4° N, 111.5° W]; J.L. Neff leg.; CAS • 1 ♀; Pima Co., Tucson, Pima Canyon ; 32.358° N, 110.933° W; 7 Mar. 2018; T. J. Wood leg.; WRME • 1 ♀; Pima Co., West of Quijotoa ; [32.13° N, 112.2° W]; J.L. Neff leg.; CAS • 1 ♀; Santa Cruz Co., 2 miles W of Highway 82 on road to Patagonia Lake Recreation Area ; [31.474° N, 110.846° W]; 23–24 Sep. 1982; W.J. Pulawski leg.; CAS • 2 ♀♀, 4 ♂♂; Santa Cruz Co., Patagonia ; [31.54° N, 110.76° W]; 16 Jun. 1984; S.L. Buchmann leg.; ex Sapindus saponaria var. drummondii ; SEMC • 1 ♀; Yavapai Co., Congress ; [34.16° N, 112.85° W]; elev. 1524 m; 1 Apr. 2002; R. S. Beal leg.; ex Sphaeralcea ; CSUC • 1 ♀, 1 ♂; Yavapai Co., Wet Beaver Creek Ranger Station ; [34.757° N, 111.764° W]; elev. 1219 m; 25 Apr. 2005; R. S. Beal leg.; CSUC • 1 ♂; 11 mi. NW of Fort Huachuca ; [31.67° N, 110.48° W]; 10 Jul. 1952; R. H. Beamer, L.D. Beamer, W.E. LaBerge, Cheng Liang leg.; SEMC • 1 ♀; 2 miles west-southwest of Prescott ; [34.51° N, 112.48° W]; elev. 1737 m; 13–14 May 2004; R. S. Beal leg.; ex Heteromeles ; CSUC • 1 ♀; 20 miles west of Tucson ; [32.22° N, 111.28° W]; 29 Apr. 1985; W.F. Chamberlain leg.; TAMU-ENTO X1313642 • 1 ♂; 23 mi. E of Kingman ; [35.18° N, 113.65° W]; 1 Jul. 1952; Cheng Liang leg.; ex Calochortus ; SEMC • 1 ♂; ibid.; 1 Jul. 1952; R. H. Beamer, L.D. Beamer, Cheng Liang, W.E. LaBerge leg.; SEMC • 7 ♀♀; 3 mi. N of Rock Spring [exact location unknown]; 7 Jul. 1952; R. H. Beamer, L.D. Beamer, W.E. LaBerge, Cheng Liang leg.; ex Eriogonum trichopes ; SEMC • 10 ♀♀; ibid.; 7 Jul. 1952; R. H. Beamer, L.D. Beamer, W.E. LaBerge, Cheng Liang leg.; SEMC • 1 ♀; 3 miles SW of Carrizo ; [33.96° N, 110.33° W]; 24 May 1997; W.F. Chamberlain leg.; TAMU-ENTO X1404715 • 4 ♀♀, 4 ♂♂; 5 mi. E of Kingman ; [35.19° N, 113.96° W]; 1 Jul. 1952; R. H. Beamer, L.D. Beamer, W.E. LaBerge, Cheng Liang leg.; ex Eriogonum trichopes ; SEMC • 2 ♂♂; ibid.; 1 Jul. 1952; R. H. Beamer, L.D. Beamer, W.E. LaBerge, Cheng Liang leg.; SEMC • 1 ♀; 7 mi. W of Seligman ; [35.32° N, 113° W]; 1 Jul. 1952; R. H. Beamer, L.D. Beamer, W.E. LaBerge, Cheng Liang leg.; SEMC • 6 ♀♀; Barfoot Park, Chiricahua ; [31.92° N, 109.28° W]; 12 Jun. 2017; BBSL 24778 , 30291 , 30296 , 30331 , 30333 , 30462 • 15 ♀♀; ibid.; 17 Jun. 2017; BBSL 30447 , 30459 , 30470 , 30472 , 30484 , 30486 , 30487 , 30489 , 30493 , 30494 , 30495 , 30496 , 30497 , 30499 , 30509 • 35 ♀♀; ibid.; 23 Jun. 2017; BBSL 21728 , 21735 , 21737 , 21738 , 21740 , 21741 , 21744 , 21747 , 21749 , 21750 , 21758 , 21759 , 21919 , 30338 , 30340 , 30341 , 30342 , 30344 , 30364 , 30367 , 30368 , 30371 , 30372 , 30375 , 30378 , 30383 , 30385 , 30392 , 30395 , 30396 , 30403 , 30404 , 30406 , 30421 , 30424 • 4 ♀♀; ibid.; 1 Jul. 2017; BBSL 24624 , 24630 , 24640 , 24654 • 2 ♀♀; ibid.; 7 Jul. 2017; BBSL 30246 , 30260 • 3 ♂♂; Huachuca Mountains ; [31.4° N, 110.3° W]; 10 Jul. 1952; R. H. Beamer leg.; SEMC • 1 ♂; Oak Creek Canyon; [34.91° N, 111.73° W]; 9 Jul. 1941; R. H. Beamer leg.; SEMC • 4 ♀♀; Rustler Park, Chiricahua ; [31.92° N, 109.28° W]; 17 Jun. 2017; BBSL 24042 , 24055 , 24060 , 24061 . – California • 1 ♀; Los Angeles Co., Chatsworth ; 34.26° N, 118.601° W; 1–8 Jul. 2017; LACM • 1 ♀; Los Angeles Co., Claremont ; 34.0894° N, 117.7111° W; 1–8 Jul. 2016; LACM • 1 ♀; Riverside Co., 18 mi. W of Blythe ; [33.61° N, 114.9° W]; 1 May 1992; Jerome G. Rozen leg.; ex Nama ; AMNH • 4 ♀♀; Riverside Co., Norco ; 33.912° N, 117.5702° W; 1–8 Jun. 2016; LACM • 1 ♂; San Bernardino Co., 20 mi. S of Needles ; [34.56° N, 114.61° W]; 3 May 1964; P.F. Torchio, G.E. Bohart leg.; BBSL700834 • 1 ♀; San Bernardino Co., Fort Piute , T12 N R19 E Sec. 18; [35.11° N, 114.97° W]; elev. 823 m; 29 May 1980; Terry L. Griswold leg.; BBSL FaunalSurvey No. 000094256 • 3 ♀♀; San Bernardino Co., Granite Mountains , T8 N R13 E Sec. 17; [34.78° N, 115.64° W]; elev. 1234 m; 22–25 May 1983; Terry L. Griswold leg.; BBSL FaunalSurvey No. 000093931, FaunalSurvey No. 000110478, FaunalSurvey No. 000110481 • 1 ♀; San Bernardino Co., Kelso Dunes , T10 N R13 E Sec. 30; [34.97° N, 115.66° W]; elev. 792 m; 12 May 1982; Terry L. Griswold leg.; BBSL FaunalSurvey No. 000094240 • 2 ♀♀; San Bernardino Co., NE of Granite Mountains, T9 N R13 E Sec. 33; [34.83° N, 115.62° W]; elev. 1160 m; 21 Sep. 1993; Terry L. Griswold leg.; ex Eriogonum deflexum ; BBSL FaunalSurvey No. 000093845, FaunalSurvey No. 000093846 • 3 ♀♀, 1 ♂; San Diego Co., Anza-Borrego ; 33.108° N, 116.4976° W; 20 Apr. 2013; K.J. Hung, Borchers, Nabors leg.; WRME • 2 ♀♀; San Diego Co., Anza-Borrego ; 33.1281° N, 116.4276° W; 23 Mar. 2012; K.J. Hung leg.; WRME • 1 ♀; ibid.; 2 Mar. 2013; K.J. Hung leg.; WRME • 1 ♀; San Diego Co., Anza-Borrego ; 33.2114° N, 116.4456° W; 12 Sep. 2013; K.J. Hung leg.; ex Pectis papposa ; WRME • 1 ♀; San Diego Co., Anza-Borrego ; 33.222° N, 116.4547° W; 12 Sep. 2013; K.J. Hung leg.; ex Chamaesyce sp. ; WRME • 1 ♀; La Mesa ; [32.77° N, 117.02° W]; 19 Jun. 2015; K.J. Hung leg.; WRME • 1 ♀; San Diego ; [32.71° N, 117.16° W]; 3 Jun. 2015; K.J. Hung leg.; WRME • 1 ♀; ibid.; 5 Apr. 2016; K.J. Hung, Lee leg.; WRME • 1 ♂; ibid.; 11 May 2016; K.J. Hung, Lee leg.; WRME • 1 ♀; ibid.; 2 Jun. 2016; K.J. Hung, Lee leg.; WRME • 1 ♀; ibid.; 14 Jul. 2016; K.J. Hung, Lee leg.; WRME • 2 ♀♀; Spring Valley ; [32.74° N, 117° W]; 20 Mar. 2015; K.J. Hung, Cen leg.; WRME • 1 ♀; ibid.; 24 Mar. 2016; K.J. Hung, Lee leg.; WRME • 1 ♀; ibid.; 14 Apr. 2016; K.J. Hung, Lee leg.; WRME • 2 ♀♀; ibid.; 2 May 2016; K.J. Hung, Lee leg.; WRME • 1 ♀; ibid.; 19 May 2016; K.J. Hung, Lee leg.; WRME • 2 ♀♀; ibid.; 8 Jun. 2016; K.J. Hung, Lee leg.; WRME. – Nevada • 2 ♂♂; Clark Co., 0.09 mi. NW of McCullough Pass ; 35.7362° N, 115.1704° W; 13 Sep. 2005; A. Portoluci leg.; ex Senecio sp. ; BBSL671758 , BBSL671759 • 1 ♂; Clark Co., 0.2 mi. E of Fletcher Spring ; 36.2733° N, 115.626° W; elev. 2296 m; 7 Jul. 2004; E. Ahlstrom leg.; ex Angelica kingii ; BBSL536927 • 1 ♂; Clark Co., 0.22 mi. W of Christmas Tree Pass ; 35.2617° N, 114.7466° W; 21 Sep. 2005; E. North leg.; ex Gutierrezia microcephala ; BBSL671304 • 1 ♂; Clark Co., 0.4 mi. E of St. Thomas Gap ; 36.4067° N, 114.0937° W; 8 Jun. 2005; S. Higbee leg.; ex Stephanomeria exigua ; BBSL670136 • 1 ♀; Clark Co., 0.54 mi. S of Paradise Gap ; 36.29° N, 114.1886° W; 24 Aug. 2005; A. Portoluci leg.; ex Baileya multiradiata ; BBSL676249 • 2 ♀♀; ibid.; 24Aug. 2005; O.I. Messinger leg.; ex Euphorbia sp. ; BBSL686060 , BBSL686062 • 1 ♂; ibid.; 24 Aug. 2005; O.I. Messinge leg.; ex Euphorbia sp. ; BBSL686063 • 1 ♀; Clark Co., 0.6 mi. WSW of Bills Spring ; 36.2931° N, 114.1735° W; 25 Aug. 2005; E. North leg.; ex Eriogonum sp. ; BBSL686664 • 1 ♀; Clark Co., 0.69 mi. N of Beehive Rock ; 36.4196° N, 114.5509° W; 23 Aug. 2005; E. North leg.; ex Croton wigginsii ; BBSL671661 • 1 ♂; Clark Co., 0.71 mi. ENE of Gass Spring ; 36.4292° N, 115.1526° W; 31 May 2005; S. Higbee leg.; ex Eriogonum racemosum ; BBSL675489 • 1 ♀; ibid.; 14 Jun. 2005; S. Higbee leg.; ex Salvia dorrii ; BBSL603861 • 1 ♂; Clark Co., 0.9 mi. E of Indian Ridge ; 36.4765° N, 115.6351° W; elev. 1396 m; 16 Jun. 2004; E. Ahlstrom leg.; ex Sphenosciadium capitellatum ; BBSL535655 • 1 ♂; Clark Co., 1 mi. WNW of Christmas Tree Pass ; 35.2653° N, 114.7648° W; 11 Jul. 2005; S. Higbee leg.; BBSL678142 • 1 ♀; Clark Co., 1.5 mi. SE of Buck Spring ; 36.3243° N, 115.7625° W; 12 Jul. 2005; S. Higbee leg.; ex Penstemon palmeri ; BBSL638186 • 1 ♀; Clark Co., 1.6 mi. SSW of Spirit Mountain ; 35.2518° N, 114.7301° W; 9 Jun. 2005; BBSL637101 • 1 ♀; ibid.; 11 Jul. 2005; S. Higbee leg.; ex Eriogonum fasciculatum ; BBSL638147 • 2 ♂♂; ibid.; 6 Sep. 2005; Terry L. Griswold leg.; ex Gutierrezia microcephala ; BBSL669833 , BBSL669834 • 1 ♀; ibid.; 21 Sep. 2005; E. North leg.; ex Eriogonum deflexum ; BBSL669959 • 3 ♀♀; Clark Co., 1.6 mi. WNW of Whitney Pocket ; 36.5314° N, 114.1626° W; 25 May 2005; E. Ahlstrom leg.; ex Eriastrum diffusum ; BBSL669349 , BBSL669350 , BBSL669351 • 1 ♀; ibid.; 25 May 2005; S. Higbee leg.; ex Eriastrum diffusum ; BBSL670304 • 1 ♀; ibid.; 25 May 2005; S. Higbee leg.; BBSL638920 • 2 ♀♀; Clark Co., 1.61 mi. NE of Sheep Mountain ; 35.7699° N, 115.2673° W; 13 Sep. 2005; E. North leg.; ex Eriogonum sp. ; BBSL685864 , BBSL685870 • 1 ♀; Clark Co., 1.81 mi. N of Crescent Peak ; 35.5045° N, 115.1417° W; 12 May 2005; E. Ahlstrom leg.; ex Baileya sp. ; BBSL676449 • 2 ♂♂; Clark Co., 1.89 mi. E of Weiser Ridge ; 36.5872° N, 114.5295° W; 7 Jun. 2005; E. Ahlstrom leg.; ex Eriastrum diffusum ; BBSL669361 , BBSL669363 • 3 ♂♂; Clark Co., 1.92 mi. E of Weiser Ridge ; 36.581° N, 114.5314° W; 7 Jun. 2005; E. Ahlstrom leg.; ex Acacia greggii ; BBSL672213 , BBSL672214 , BBSL672215 • 1 ♀; Clark Co., 16.91 mi. SE of Devils Hole Hills ; 36.2433° N, 115.8991° W; 20 Jun. 2005; R. Andrus leg.; ex Stanleya sp. ; BBSL676677 • 1 ♀; Clark Co., 2.02 mi. NE of Spring Mountain ; 35.8181° N, 115.4594° W; 12 Sep. 2005; A. Portoluci leg.; ex Gutierrezia microcephala ; BBSL671151 • 1 ♂; ibid.; 12 Sep. 2005; A. Portoluci leg.; ex Gutierrezia microcephala ; BBSL671148 • 2 ♀♀; Clark Co., 2.15 mi. N of Piute Valley ; 35.554° N, 115.0758° W; 22 Sep. 2005; E. North leg.; BBSL678639 , BBSL678657 • 1 ♀; Clark Co., 2.16 mi. SW of Wechech Basin ; 36.4559° N, 114.1519° W; 12 May 2005; S. Higbee leg.; ex Phacelia indecora ; BBSL677594 • 2 ♀♀; Clark Co., 2.34 mi. SE of The Narrows ; 36.6064° N, 114.4987° W; 3 May 2005; S. Higbee leg.; ex Mentzelia involucrata ; BBSL650584 , BBSL650585 • 1 ♂; Clark Co., 2.46 mi. SW of Highland Range ; 35.5657° N, 115.0804° W; 22 Sep. 2005; E. North leg.; ex Gutierrezia microcephala ; BBSL671389 • 1 ♀; Clark Co., 2.49 mi. W of Grapevine Canyon ; 35.2643° N, 114.7772° W; 11 Oct. 2005; Terry L. Griswold leg.; ex Pectis papposa ; BBSL686226 • 2 ♀♀; Clark Co., 2.5 km NW of Callville Bay ; 36.157° N, 114.7438° W; 12 Mar. 1998; Terry L. Griswold leg.; BBSL277441 , BBSL277451 • 1 ♀; Clark Co., 2.5 mi. S of Wheeler Well ; 36.336° N, 115.8236° W; 12 Jul. 2005; E. Green leg.; ex Eriodictyon angustifolium ; BBSL637611 • 1 ♂; Clark Co., 2.5 mi. SE of Rainbow Mountain ; 36.0896° N, 115.4493° W; elev. 1138 m; 23 Aug. 2004; S. Higbee leg.; ex Gutierrezia microcephala ; BBSL542099 • 1 ♀; ibid.; 9 Sep. 2004; S. Higbee, Terry L. Griswold leg.; BBSL576731 • 1 ♂; ibid.; 21 Sep. 2004; S. Higbee leg.; ex Gutierrezia microcephala ; BBSL533881 • 1 ♀; Clark Co., 2.85 mi. SE of Rainbow Mountain ; 36.0804° N, 115.4587° W; 24 Jun. 2005; S. Higbee leg.; BBSL650810 • 1 ♀; Clark Co., 2.93 mi. NE of McCullough Range ; 35.5692° N, 115.124° W; 22 Sep. 2005; E. North leg.; ex Gutierrezia microcephala ; BBSL671378 • 1 ♂; Clark Co., 3.1 mi. SE of Black Butte ; 35.8524° N, 115.6717° W; elev. 820 m; 4 May 2004; S. Higbee leg.; ex Prosopis glandulosa var. torreyana ; BBSL544625 • 3 ♂♂; Clark Co., 3.5 mi. SE of Hiko Spring ; 35.1252° N, 114.6432° W; elev. 235 m; 27 Apr. 2004; E. Ahlstrom leg.; BBSL577372 , BBSL577379 , BBSL577382 • 2 ♀♀; Clark Co., 3.5 mi. W of Christmas Tree Pass ; 35.2658° N, 114.8021° W; 21 Apr. 2005; D. Allen leg.; BBSL639553 , BBSL639555 • 17 ♀♀; Clark Co., 3.9 mi. SSW of Whitney Pocket ; 36.4651° N, 114.1537° W; 26 May 2005; R. Andrus leg.; BBSL650741 , BBSL650743 , BBSL650744 , BBSL650745 , BBSL650763 , BBSL650764 , BBSL650765 , BBSL650768 , BBSL650770 , BBSL650773 , BBSL650774 , BBSL650775 , BBSL650777 , BBSL650790 , BBSL650791 , BBSL650793 , BBSL650794 • 1 ♂; Clark Co., 3.94 mi. SE of Piute Wash ; 35.2685° N, 114.8198° W; 11 Oct. 2005; Terry L. Griswold leg.; ex Pectis papposa ; BBSL686123 • 1 ♀; Clark Co., 4 mi. W of Bunkerville ; [36.77° N, 114.2° W]; 14 Jun. 1989; P.F. Torchio leg.; BBSL510006 • 1 ♂; Clark Co., 4.21 mi. SW of Roman Wash ; 35.2658° N, 114.8021° W; 21 Sep. 2005; A. Portoluci leg.; ex Eriogonum deflexum ; BBSL672337 • 1 ♂; Clark Co., 5 mi. NE of Spring Mountains ; 35.8167° N, 115.5022° W; 30 Aug. 2005; A. Portoluci leg.; ex Chamaesyce fendleri ; BBSL676269 • 3 ♀♀; Clark Co., 5 mi. S of Searchlight ; [35.39° N, 114.92° W]; 23 Sep. 1983; P.F. Torchio, D.M. Torchio leg.; BBSL510019 , BBSL510020 , BBSL510021 • 3 ♀♀; Clark Co., Arizona Road ; 36.5253° N, 114.0522° W; elev. 1330 m; 9–26 Jun. 2018; G. R. Ballmer leg.; UCRC ENT 519887 , 519897 , 519902 • 1 ♀; Clark Co., Black Mountain , S.; 35.4838° N, 114.7497° W; 14 Aug. 1998; C. Shultz leg.; BBSL313537 • 1 ♀; Clark Co., Blue Point Spring ; 36.3894° N, 114.4295° W; elev. 463 m; 20 Apr. 2005; R. Andrus leg.; ex Mentzelia involucrata ; BBSL569732 • 1 ♀; Clark Co., Bunkerville ; [36.77° N, 114.13° W]; 3 Jun. 1993; P.F. Torchio, Trostle leg.; ex Baileya ; BBSL510000 • 2 ♀♀; Clark Co., Callville Bay Road; 36.173° N, 114.7653° W; 12 Mar. 1998; Terry L. Griswold leg.; BBSL276200 , BBSL276201 • 1 ♂; Clark Co., Christmas Tree Pass ; 35.26° N, 114.7418° W; 10 Jun. 1998; C. Shultz, K. Receveur, K. Keen, M.Andres leg.; BBSL304776 • 1 ♂; ibid.; 10 Jun. 1998; M. Andres, C. Shultz leg.; BBSL291092 • 1 ♂; Clark Co., Clark Canyon ; 36.3252° N, 115.7617° W; 20 Jul. 1998; M. Andres, C. Shultz leg.; ex Chaenactis ; BBSL304958 • 1 ♀; Clark Co., Cottonwood Cove ; 35.4887° N, 114.6943° W; 14 Aug. 1998; C. Shultz, Terry L. Griswold leg.; BBSL316688 • 1 ♂; Clark Co., E of Christmas Tree Pass ; 35.2527° N, 114.7297° W; 10 Jun. 1998; M. Andres, C. Shultz leg.; BBSL291058 • 1 ♀; Clark Co., Elbow Canyon ; 36.7339° N, 115.0111° W; 10 Jun. 2005; S. Higbee leg.; ex Baileya multiradiata ; BBSL603733 • 1 ♀; Clark Co., Elbow Canyon ; 36.7422° N, 114.9916° W; 10 Jun. 2005; E. Ahlstrom leg.; ex Arctomecon merriamii ; BBSL672241 • 1 ♂; Clark Co., Gold Butte ; 36.2815° N, 114.1933° W; 18 Sep. 1998; W. R. Bowlin leg.; BBSL320873 • 5 ♀♀; Clark Co., Grand Gulch Road , 22 air mi. S of Mesquite; [36.49° N, 114.07° W]; 11–21 May 1983; F.D. Parker, J.H. Parker leg.; BBSL510017 , BBSL510018 , BBSL510022 , BBSL510023 , BBSL510024 • 1 ♀; Clark Co., Gypsum Spring ; 36.2093° N, 114.912° W; 12 Mar. 1998; Terry L. Griswold leg.; BBSL277461 • 1 ♀; Clark Co., Harris Spring Canyon; 36.2583° N, 115.493° W; 1 Jun. 2005; R. Andrus leg.; ex Penstemon palmeri ; BBSL639120 • 2 ♂♂; Clark Co., Hidden Valley ; 35.819° N, 115.2° W; 11 Aug. 1998; M. Andres, C. Shultz leg.; ex Sphaeralcea ambigua ; BBSL312600 , BBSL312601 • 1 ♂; Clark Co., Horse Spring Wash; 36.3292° N, 114.1457° W; 18 Sep. 1998; W. R. Bowlin leg.; BBSL319631 • 1 ♀; Clark Co., Lovell Canyon ; 36.1526° N, 115.5719° W; 7 Jul. 2005; Terry L. Griswold leg.; ex Heliomeris multiflora ; BBSL637833 • 1 ♂; Clark Co., Lovell Canyon ; 36.1648° N, 115.5758° W; elev. 1801 m; 16 Jun. 2004; D. Skandilis leg.; ex Verbena bracteata ; BBSL536691 • 2 ♀♀; Clark Co., Mesquite ; [36.81° N, 114.07° W]; 4 Oct. 1988; R. W. Rust, P.F. Torchio leg.; ex Eriogonum ; BBSL510007 , BBSL510008 • 8 ♀♀; Clark Co., Mesquite Valley , E.; 35.761° N, 115.5832° W; 10 Aug. 1998; M. Andres, C. Shultz, Terry L. Griswold leg.; BBSL312920 , BBSL312928 , BBSL312932 , BBSL312933 , BBSL312938 , BBSL312940 , BBSL312943 , BBSL312949 • 6 ♀♀; Clark Co., Mormon Well Road ; 36.6253° N, 115.1115° W; 16 Jul. 1998; M. Andres, C. Shultz leg.; BBSL266027 , BBSL266029 , BBSL266032 , BBSL266033 , BBSL269761 , BBSL269762 • 1 ♂; Clark Co., Mud Wash ; 36.433° N, 114.1588° W; 27 Aug. 1998; O.I. Messinge, C. Shultz, S. Messinger leg.; BBSL317239 • 2 ♀♀; Clark Co., N of Devils Throat ; 36.4427° N, 114.151° W; 6 Oct. 1998; Terry L. Griswold leg.; BBSL318911 , BBSL318930 • 1 ♂; ibid.; 6 Oct. 1998; Terry L. Griswold leg.; BBSL318916 • 1 ♂; Clark Co., NE of Little Virgin Peak ; 36.6097° N, 114.2167° W; 6 Oct. 1998; Terry L. Griswold leg.; BBSL318672 • 1 ♂; Clark Co., NE of Mica Peak ; 36.3348° N, 114.1445° W; 7 Jun. 1998; F.D. Parker, Terry L. Griswold leg.; BBSL278946 • 1 ♂; Clark Co., SE of Black Butte ; 35.8627° N, 115.691° W; 15 Sep. 1998; W. R. Bowlin leg.; BBSL324372 • 1 ♂; Clark Co., SE of Little Virgin Peak ; 36.59° N, 114.2167° W; 8 Jun. 1998; M. Andres, K. Receveur leg.; ex Eriogonum ; BBSL294853 • 7 ♀♀; Clark Co., Spring Mountain Ranch ; [36.07° N, 115.45° W]; 16 May–16 Jul. 1988; M.J. Verchick leg.; BBSL510009 , BBSL510010 , BBSL510011 , BBSL510012 , BBSL510013 , BBSL510014 , BBSL510016 • 1 ♀; Clark Co., St. Thomas Gap, T17 S R71 E Sec. 31; [36.45° N, 114.11° W]; 11 Apr.–10 May 1984; R. C. Bechtel, J.B. Knight leg.; BBSL510026 • 1 ♂; Clark Co., SW of Ireteba Peaks ; 35.571° N, 114.8535° W; 9 Jun. 1998; M.Andres, K. Keen leg.; ex Eriogonum fasciculatum ; BBSL295772 • 1 ♂; Clark Co., Virgin Valley ; 36.6868° N, 114.2643° W; 6 Oct. 1998; Terry L. Griswold leg.; BBSL319048 • 12 ♀♀; Clark Co., W of Beehive Rock ; 36.4088° N, 114.5803° W; 5 Aug. 1998; M. Andres, C. Shultz leg.; BBSL309738 , BBSL309748 , BBSL309749 , BBSL309765 , BBSL309767 , BBSL309773 , BBSL309775 , BBSL309777 , BBSL309783 , BBSL309787 , BBSL309803 , BBSL309807 • 1 ♂; Clark Co., W of Virgin Mountains ; 36.5768° N, 114.2042° W; 6 Oct. 1998; Terry L. Griswold leg.; BBSL318593 • 1 ♀; Clark Co., W of Wilson Pass ; 35.8705° N, 115.541° W; 5 Jun. 1998; F.D. Parker leg.; BBSL279754 • 1 ♀; Clark Co., West End Wash, Lake Mead NRA; [36.15° N, 114.72° W]; elev. 549 m; 31 Mar. 1993; Terry L. Griswold, V. J. Tepedino leg.; ex Arctomecon californicum ; BBSL FaunalSurvey No. 000031403 . – New Mexico • 1 ♀; Chaves Co., Bitter Lake NWR, trailer north of HQ; 33.4636° N, 104.4066° W; 20 Jul. 2004; D. Richman leg.; NMSU • 1 ♀; Colfax Co., Highway 64; 36.6369° N, 104.6775° W; elev. 1932 m; 6 Aug. 2007; Jason J. Gibbs, C. Sheffield leg.; PCYU • 2 ♀♀, 1 ♂; Doña Ana Co., Las Cruces ; [32.32° N, 106.76° W]; 12 Jun. 1950; R. H. Beamer leg.; SEMC • 2 ♀♀; ibid.; 17 Jun. 1950; L.D. Beamer leg.; SEMC • 14 ♀♀; ibid.; 15 Jul. 1952; R. H. Beamer, L.D. Beamer, W.E. LaBerge, Cheng Liang leg.; SEMC • 3 ♀♀; ibid.; 25 Apr. 1954; R. H. Beamer leg.; ex Salix ; SEMC • 1 ♀; ibid.; 25 Jun. 1962; S. R. Race leg.; ex cotton; NMSU • 1 ♀; Doña Ana Co., Leasburg Dam State Park ; [32.49° N, 106.92° W]; 25 Jun. 1983; W.J. Pulawski leg.; CAS • 1 ♀; Doña Ana Co., Mesilla Park ; [32.27° N, 106.76° W]; 29 Jun. 1962; S. R. Race leg.; ex alfalfa; NMSU • 1 ♀; Doña Ana Co., NMSU Leyendecker Farm, Route 28 at Rio Grande , 5 miles southeast Mesilla ; [32.2° N, 106.742° W]; elev. 1189 m; 15 Jul. 2004; ex Gossypium ; NMSU • 1 ♀; Eddy Co., 1.5 km SSW of Main Cave Entrance ; 32.1638° N, 104.4443° W; elev. 1152 m; 31 Aug. 2010; J.D. Herndon, R. Krauss leg.; BBSL CAVE21910 • 1 ♀; Eddy Co., 1.9 km SSW of Main Cave Entrance ; 32.1615° N, 104.4494° W; elev. 1161 m; 31 Aug. 2010; R. Krauss, J.D. Herndon leg.; BBSL CAVE21452 • 6 ♀♀; Grant Co., 0.5 km N of Hachita ; 31.9233° N, 108.3258° W; elev. 1440 m; 23 Aug. 2001; R. W. Brooks leg.; ex Chamaesaracha coronopus ; SEMC SM0273737 , SM0273739 , SM0273747 , SM0273756 , SM0273759 , SM0273764 • 1 ♀; ibid.; 23 Aug. 2001; R. W. Brooks leg.; ex Machaeranthera tanacetifolia ; SEMC SM0274237 • 4 ♀♀; Grant Co., 19 mi. N of Hachita ; 32.1822° N, 108.3158° W; elev. 1430 m; 24 Sep. 1999; R. W. Brooks leg.; SEMC SM0150372 , SM0150373 , SM0150374 , SM0150375 • 1 ♀; Hidalgo Co., 1 km N of Animas ; [31.96° N, 108.81° W]; 1 Aug. 1988; Bryan N. Danforth leg.; ex Sphaeralcea ; SEMC • 7 ♀♀; Hidalgo Co., 11.2 km E of Portal, 0.8 km W of Highway 80; 31.8706° N, 109.0433° W; elev. 1250 m; 23 Aug. 2000; R. W. Brooks leg.; SEMC SM0198518 , SM0198520 , SM0198521 , SM0198523 , SM0198524 , SM0198526 , SM0198530 • 1 ♀, 1 ♂; Hidalgo Co., 13 mi. N of Rodeo ; [32.02° N, 109.03° W]; elev. 1372 m; 16 Aug. 1964; Charles D. Michener leg.; ex “ Aplopappus spinulosus ”; SEMC • 2 ♀♀, 1 ♂; Hidalgo Co., 20 mi. S of Animas ; [31.66° N, 108.81° W]; 23 Aug. 2008; John S. Ascher, Jerome G. Rozen, M.A. Rozen leg.; AMNH • 4 ♀♀; Hidalgo Co., 21.7 km S of Animas ; [31.75° N, 108.81° W]; elev. 1400 m; 19 Aug. 2001; Charles D. Michener leg.; SEMC SM0270807 , SM0270808 , SM0270809 , SM0270810 • 1 ♀; Hidalgo Co., 22.6 km S of Animas ; [31.74° N, 108.81° W]; 13 Jul. 1987; ex Berlandiera ; SEMC • 1 ♂; Hidalgo Co., 23.5 mi. S of Animas ; 31.6184° N, 108.8674° W; 23 Aug. 2007; Jason J. Gibbs leg.; PCYU • 5 ♂♂; Hidalgo Co., 26 mi. S of Animas ; 31.585° N, 108.87° W; 18 Sep. 2012; Jason J. Gibbs leg.; CUIC • 1 ♀; Hidalgo Co., 28 mi. S of Animas ; [31.555° N, 108.876° W]; 19 Aug. 2010; H. Campbell leg.; AMNH • 1 ♂; ibid.; 19 Aug. 2010; H. Campbell leg.; AMNH • 2 ♀♀; Hidalgo Co., 28 mi. S of Animas ; 31.555° N, 108.876° W; 22 Aug. 2019; Jason J. Gibbs leg.; WRME 0500836 , 0500837 • 2 ♀♀; Hidalgo Co., 41.8 km S of Animas ; [31.58° N, 108.87° W]; elev. 1400 m; 19 Aug. 2001; Charles D. Michener leg.; SEMC SM0270825 , SM0270827 • 2 ♀♀; Hidalgo Co., 5.5 mi. E of Animas ; [31.96° N, 108.72° W]; 22 Aug. 1995; Jerome G. Rozen, S.A. Budick, Gretchen LeBuhn leg.; AMNH • 2 ♀♀; Hidalgo Co., 5.5 mi. E of Animas ; 31.95° N, 108.7114° W; elev. 1390 m; 22 Sep. 1999; R. W. Brooks, Charles D. Michener leg.; ex Euphorbia ; SEMC SM0149738 , SM0149741 • 1 ♂; ibid.; 22 Sep. 1999; R. W. Brooks, Charles D. Michener leg.; SEMC SM0149718 • 1 ♀; Hidalgo Co., 5.5 mi. E of Animas ; 31.9572° N, 108.7144° W; elev. 1219 m; 19 Aug. 2001; SEMC SM0262458 • 1 ♀; Hidalgo Co., 6 mi. N of Rodeo ; [31.92° N, 109.03° W]; 26 Aug. 2011; WRME • 5 ♀♀; Hidalgo Co., 6 mi. S of Animas ; [31.86° N, 108.81° W]; 19 Aug. 2010; H. Campbell leg.; AMNH • 1 ♀; Hidalgo Co., 8 km N of Rodeo ; [31.91° N, 109.03° W]; 6 Aug. 1988; Bryan N. Danforth leg.; SEMC • 3 ♂♂; Hidalgo Co., Animas ; [31.95° N, 108.81° W]; 9 May 1993; Robert L. Minckley, K. Toal leg.; ex Phacelia ; SEMC • 2 ♀♀; Hidalgo Co., Animas vicinity; [31.95° N, 108.81° W]; 28 Jul. 1976; M. Rothschild leg.; SEMC • 1 ♀; Hidalgo Co., Cienega Ranch ; [32.04° N, 109.04° W]; 20 Aug. 1998; Jerome G. Rozen, K. Franklin leg.; AMNH • 1 ♀; Hidalgo Co., E. Animas ; [31.95° N, 108.8° W]; 16 May 1987; Bryan N. Danforth leg.; SEMC • 1 ♀; Hidalgo Co., Rodeo ; [31.84° N, 109.03° W]; elev. 1268 m; 22 Aug. 1964; Charles D. Michener leg.; SEMC • 2 ♀♀, 1 ♂; Hidalgo Co., Rodeo ; [31.84° N, 109.03° W]; 31 Aug. 1970; D.J. Brothers leg.; SEMC • 1 ♂; ibid.; 11 Aug. 1975; M. Chabot leg.; SEMC • 5 ♀♀; Hidalgo Co., Rodeo vicinity; [31.84° N, 109.03° W]; 11–12 Jun. 1987; Bryan N. Danforth leg.; ex Berlandiera ; SEMC • 1 ♀; ibid.; 11 Jun. 1987; Bryan N. Danforth leg.; ex Euphorbia ; SEMC • 3 ♀♀; ibid.; 5 Aug. 1987; Bryan N. Danforth leg.; SEMC • 1 ♀; Hidalgo Co., San Simone cienega; [32.3° N, 108.9° W]; 17–24 May 1987; Bryan N. Danforth leg.; ex Prosopis ; SEMC • 1 ♀; ibid.; 1–9 Jun. 1987; Bryan N. Danforth leg.; SEMC • 1 ♀; Luna Co., Columbus ; [31.83° N, 107.64° W]; 21 Jul. 1982; W.J. Pulawski leg.; CAS • 1 ♀; ibid.; 26 Jun. 1983; W.J. Pulawski leg.; CAS • 1 ♀; Sierra Co., #4, 6 mi. W of Caballo ; [32.98° N, 107.38° W]; 30 May 1991; G. W. Byers leg.; ex Larrea ; SEMC • 1 ♀; Socorro Co., Bosque del Apache National Wildlife Refuge ; [33.8° N, 106.89° W]; 24 Mar.–7 Apr. 2008; TAMU • 1 ♀; ibid.; 16–30 Jun. 2008; TAMU • 1 ♀; Socorro Co., Sevilleta National Wildlife Refuge ; 34.3325° N, 106.6328° W; 17 Jun.–1 Jul. 2010; Karen Wetherill leg.; TAMU • 1 ♀; ibid.; 15–29 Jul. 2010; Karen Wetherill leg.; TAMU • 1 ♀; Socorro Co., Sevilleta National Wildlife Refuge ; 34.335° N, 106.7219° W; 13–27 Jun. 2012; Karen W. Wright leg.; TAMU • 2 ♀♀; Socorro Co., Sevilleta National Wildlife Refuge ; 34.3431° N, 106.7417° W; 21 Apr.–5 May 2009; Karen Wetherill leg.; TAMU • 1 ♀; ibid.; 19 May–2 Jun. 2009; Karen Wetherill leg.; TAMU • 2 ♀♀; ibid.; 15–29 Jun. 2009; Karen Wetherill leg.; TAMU • 2 ♀♀; 20 mi. W of Las Cruces ; [32.33° N, 107.1° W]; 13 Jul. 1952; R. H. Beamer, L.D. Beamer, W.E. LaBerge, Cheng Liang leg.; SEMC • 1 ♂; 35 mi. N of Hot Springs ; [36.16° N, 105.29° W]; 22 Jul. 1950; R. H. Beamer leg.; ex Baileya multiradiata ; SEMC • 1 ♂; Alamogordo ; [32.9° N, 105.96° W]; 26 Jun. 1940; L.J. Lipovsky leg.; SEMC • 1 ♂; Cloudcroft ; [32.96° N, 105.74° W]; 27 Jun. 1940; E.E. Kenaga leg.; SEMC • 1 ♂; Garfield ; [32.76° N, 107.26° W]; 16 Jul. 1952; R. H. Beamer, L.D. Beamer, W.E. LaBerge, Cheng Liang leg.; SEMC • 2 ♀; Hot Springs ; [35.65° N, 105.29° W]; 22 Jul. 1950; R. H. Beamer leg.; ex Chamaesaracha coniodes ; SEMC • 2 ♂♂; ibid.; 22 Jul. 1950; W.J. Arnold leg.; SEMC • 1 ♀; Mayhill ; [32.89° N, 105.48° W]; 21 Jun. 1967; W.F. Chamberlain leg.; TAMU-ENTO X1312641 • 2 ♀♀; Nogal ; [33.56° N, 105.71° W]; 11 Jun. 1959; R. H. Beamer leg.; SEMC • 1 ♀, 1 ♂; Organ ; [32.43° N, 106.6° W]; 3 Jul. 1940; D.E. Hardy leg.; SEMC • 1 ♂; ibid.; 3 Jul. 1940; L.C. Kuitert leg.; SEMC . – Texas • 1 ♀; Brewster Co., Big Bend National Park, Hot Springs ; [29.182° N, 102.992° W]; elev. 560 m; 14 Apr. 1986; Terry L. Griswold leg.; ex Chilopsis ; BBSL FaunalSurvey No. 000109944 • 1 ♂; ibid.; 14 Apr. 1986; Terry L. Griswold leg.; ex Chilopsis ; BBSL FaunalSurvey No. 000109943 • 8 ♂♂; ibid.; 14 Apr. 1986; Terry L. Griswold leg.; BBSL FaunalSurvey No. 000109935, FaunalSurvey No. 000109936, FaunalSurvey No. 000109937, FaunalSurvey No. 000109938, FaunalSurvey No. 000109939, FaunalSurvey No. 000109940, FaunalSurvey No. 000109941, FaunalSurvey No. 000109942 • 1 ♀; Brewster Co., Elephant Mountain WMA, campground & vicinity; 30.0482° N, 103.5698° W; 22–23 Apr. 2011; E.G. Riley leg.; TAMU-ENTO X0611399 • 1 ♀; Brewster Co., Terlingua Creek , 32 km N of Study Butte; [29.58° N, 103.69° W]; 16 Apr. 1986; R. T. Griswold leg.; BBSL FaunalSurvey No. 000109945 • 1 ♀; Culberson Co., Hurds Draw Road + Highway 10 intersection; [31.07° N, 104.28° W]; 31 May 1999; Laurence Packer leg.; PCYU • 1 ♀; El Paso Co., Franklin Mountains State Park, McKelligon Canyon Park ; [31.84° N, 106.485° W]; 15–30 Jul. 2005; N. Brown leg.; TAMU-ENTO X1359645 • 1 ♀; Hudspeth Co., Indio Mountains Research Station above Squaw Spring ; 30.7969° N, 105.0119° W; elev. 1280 m; 12Apr. 2002; A. R. Gillogly leg.; TAMU-ENTO X1413050 • 1 ♀; Hudspeth Co., Indio Mountains Research Station, Squaw Spring ; 30.7969° N, 105.0119° W; 12–13 Apr. 2002; M.J. Yoder leg.; TAMU-ENTO X1408409 • 2 ♀♀; Hudspeth Co., Indio Mountains Research Station, vicinity of Indio Ranch House ; 30.7769° N, 105.0161° W; elev. 1231 m; 12–13 Apr. 2002; M.J. Yoder leg.; TAMU-ENTO X1370805, X1417834 • 1 ♀; Jeff Davis Co., 9 mi. S of Kent ; [30.94° N, 104.22° W]; 29 May 1999; Laurence Packer leg.; PCYU • 1 ♀; Jeff Davis Co., Chihuahuan Desert Research Institute ; [30.54° N, 103.84° W]; 29 Apr. 2003; Laurence Packer, Fraser leg.; PCYU • 1 ♀; Val Verde Co., Devil’s River ; 29.7816° N, 100.9932° W; elev. 380 m; 27 Apr. 2019; Karen W. Wright leg.; ex Argemone ; TAMU-ENTO X1600145 • 1 ♂; Big Bend National Park, Santa Elena Canyon ; [29.165° N, 103.612° W]; elev. 640 m; 5 May 1959; W. R. M. Mason leg.; CNC • 1 ♀; [exact location unknown]; TAMU-ENTO X1388663. – Utah • 1 ♀; Millard Co., Delta , 2700 S 1700 E; 39.3123° N, 112.5448° W; 8–21 Jun. 2016; M.E. Christman leg.; BBSL • 1 ♀; Washington Co., 0.28 mi. NNE of Spendlove Knoll ; 37.3404° N, 113.1059° W; 13 Jun. 2006; B. Hays, F. Nicklen leg.; BBSL ZION22852 • 1 ♂; Washington Co., Beaver Dam Wash ; [37.1° N, 114.02° W]; 10–24 Jun. 1987; BBSL700822 • 1 ♀; Washington Co., Paradise Canyon; [37.15° N, 113.62° W]; 8–14 Jul. 1983; Dan Beck leg.; BBSL700674 .

Description

Female

COLOURATION. Head and mesosoma blue to blue-green; clypeus apex black; labrum reddish brown; mandible orange with black base and red apex; flagellum dark reddish brown dorsally, light brown ventrally. Pronotal lobe dark reddish brown; legs reddish brown; tegula translucent orange; wing membrane hyaline, veins with subcosta dark brown, otherwise brown to pale amber. Metasoma dark reddish brown with rims of terga and sterna broadly translucent yellow.

PUBESCENCE. Body pile colour white. Tomentum dense on paraocular area, frons lower half, gena, pronotal collar and lobe, space between pronotal lobe and tegula, metanotum anterior margin, metepisternum, T2–T3 basolaterally, and T4 throughout; sparse on clypeus upper half, supraclypeal area, preëpisternum, and mesepisternum. Mesoscutum pubescence thin to moderately plumose. Wing setae light and dark, short and dense. Acarinarial fan complete, sparse. T2 fringes sparse, T3 fringes sparse.

SURFACE SCULPTURE. Clypeus shiny, with punctures dense in basal half (IS <1 PD), large and irregularly sparse apically (IS <2 PD); supraclypeal area shiny, with punctures dense (IS ≤ 1 PD); paraocular area weakly imbricate around antenna socket, otherwise shiny, with punctures dense (IS <1 PD); frons shiny, with punctures crowded (IS = 0 PD); vertex shiny, with punctures dense laterally (IS <1 PD), fine and sparse medially (IS = 1–3 PD); gena shiny, with punctures dense (IS ≤ 1 PD); postgena shiny, becoming imbricate posteriorly. Tegula punctures absent; mesoscutum tessellate, becoming shiny posterolaterally, with punctures dense (IS <1 PD), becoming moderately sparse submedially and anteromedially (IS = 1–2 PD); scutellum tessellate, becoming shiny submedially, with punctures dense (IS <1 PD); metapostnotum weakly imbricate, becoming tessellate posteriorly, with rugae strong, anastomosing, nearly reaching posterior margin; preëpisternum shiny with crowded punctures (IS = 0 PD); hypoepimeron shiny, with punctures dense (IS <1 PD); mesepisternum shiny, with punctures dense (IS <1 PD); metepisternum ruguloso-lineate dorsally, ruguloso-punctate medially, tessellate ventrally; propodeum lateral surface tessellate, posterior surface tessellate. T1 anterior slope weakly coriarious, disc shiny, with punctures moderately dense (IS = 1–2 PD), impunctate in large subapicolateral boss and on rim medially; T2 disc shiny, with punctures moderately dense (IS = 1–2 PD), apical rim weakly coriarious, with punctures sparse (IS = 1–3 PD).

STRUCTURE. Face length/width ratio 0.81 (± 0.01 SD); clypeus apicolateral denticles rounded knobs; gena/eye width ratio 0.9 (± 0.06 SD). Pronotal angle slightly obtuse, nearly right-angled; intertegular span 0.84 (± 0.03 SD) mm; mesoscutum length/width ratio 0.85 (± 0.02 SD); mesoscutum/scutellum length ratio 2.86 (± 0.16 SD); scutellum/metanotum length ratio 1.58 (± 0.1 SD); metanotum/metapostnotum length ratio 0.68 (± 0.09 SD); forewing with 3 submarginal cells; tegula enlarged, reaching posterior margin of mesoscutum; inner margin straight; tegula length 0.37 (±0.03 SD) mm, width 0.19 (± 0.02 SD) mm; propodeum lateral carinae not reaching dorsal margin; oblique carina strong, straight. T2 depressed apical rim less than 50% of tergum. (n = 11)

VARIATION. Various parts of the integument including the clypeus apical margin, labrum, flagellum ventral surface, pronotal lobe, and leg joints can vary from black to orange. The metasoma can vary from black to reddish brown. The tegula is rarely opaque brown but still maintains an orange tint distinct from L. profundum sp. nov. (this may be due to liquid accumulating and drying under the tegula). The mesoscutum and mesepisternum are sometimes entirely dull, the mesoscutum is sometimes densely punctate submedially (IS ≤1 PD), and the metasomal terga can be shiny or coriarious.

Male

COLOURATION. Head and mesosoma blue to blue-green; clypeus apex reddish brown; labrum reddish brown to orange; mandible orange with black base and red apex; flagellum reddish brown dorsally, orange ventrally. Pronotal lobe dark reddish brown; legs reddish brown with femur-tibia joints and tarsi light brown to orange; tegula orange or reddish brown; wing membrane hyaline, veins with subcosta dark brown, otherwise brown to pale amber. Metasoma black or dark reddish brown with depressed apical rims of terga and sterna lighter reddish brown, downcurved lateral areas of terga becoming translucent yellow.

PUBESCENCE. Body pile colour white. Tomentum dense on face below median ocellus, gena, pronotal angle and lobe, and space between pronotal lobe and tegula; sparse on margins of mesoscutum, episterna, and T2–T3 basolaterally. Mesoscutum pubescence simple to moderately plumose. Wing setae light with dark intermixed apically, long and sparse. Sterna pubescence short (0.5–1.5 OD), moderately plumose, sparse.

SURFACE SCULPTURE. Clypeus shiny, with punctures dense (IS <1 PD); supraclypeal area shiny, with punctures dense (IS ≤ 1 PD); paraocular area shiny, with punctures dense (IS <1 PD); frons shiny, with punctures crowded (IS = 0 PD); vertex shiny, with punctures dense laterally (IS <1 PD), moderately sparse medially (IS = 1–2 PD); gena shiny, with punctures dense (IS ≤ 1 PD); postgena shiny to weakly imbricate. Tegula punctures absent; mesoscutum shiny, with punctures moderately sparse (IS = 1–2 PD), becoming dense marginally and on median line (IS ≤ 1 PD); scutellum shiny, with punctures dense (IS ≤ 1 PD); metanotum shiny to weakly imbricate and densely punctate (IS <1 PD); metapostnotum shiny to imbricate, with rugae strong, anastomosing, reaching or nearly reaching posterior margin; preëpisternum shiny with crowded punctures (IS = 0 PD); hypoepimeron shiny, with punctures dense (IS <1 PD); mesepisternum shiny, with punctures dense (IS <1 PD); metepisternum ruguloso-lineate dorsally, areolate ventrally; propodeum lateral surface shiny to imbricate, with punctures dense (IS ≤ 1 PD) to crowded (IS = 0 PD), posterior surface shiny with dense punctures (IS <1 PD). T1 anterior slope shiny, disc shiny, with punctures moderately dense (IS = 1–2 PD), sparse in small subapicolateral boss (IS = 2–6 PD) and absent on rim medially; T2 disc shiny, with punctures moderately dense (IS = 1–2 PD), apical rim shiny, with punctures absent.

STRUCTURE. Face length/width ratio 0.86 (±0.03 SD); gena/eye width ratio 0.77 (± 0.09 SD). Pronotal angle obtuse; intertegular span 0.71 (± 0.07 SD) mm; mesoscutum length/width ratio 0.89 (± 0.03 SD); mesoscutum/scutellum length ratio 2.67 (± 0.14 SD); scutellum/metanotum length ratio 1.87 (±0.17 SD); metanotum/metapostnotum length ratio 0.71 (± 0.07 SD); forewing with 3 submarginal cells; tegula enlarged, reaching or nearly reaching posterior margin of mesoscutum in dorsal view; inner margin weakly concave; tegula length 0.35 (±0.03 SD) mm, width 0.19 (± 0.01 SD) mm; propodeum lateral carinae not reaching dorsal margin; oblique carina absent. (n = 10)

GENITALIA. As in Fig. 76C . Gonocoxite about 1.3 times as broad as gonostylus. Gonostylus rounded on outer margin with no concavities, with sparse long setae dorsally and short setae along inner margin. Penis valves divergent apically, with short sparse setae dorsolaterally in basal half. Retrorse lobe rounded, about 2.5 times as long as broad, with sparse short setae apically.

VARIATION. The mesoscutum and scutellum punctures can vary from dense (IS ≤ 1.5 PD) to sparse (IS = 1–3 PD), the metapostnotum can vary from mostly shiny to mostly imbricate, and the propodeum lateral face can vary from shiny and distinctly punctate to imbricate and obscurely punctate.

Range

Sonoran, Chihuahuan, and Mojave Deserts ( Fig. 14 ).

Floral hosts

AMARANTHACEAE Juss. : Salsola L.: S. kali L.: S. k. subsp. tragus (L.) Čelak. • APIACEAE : Angelica L.: A. kingii (S.Watson) J.M.Coult. & Rose • Sphenosciadium A.Gray : S. capitellatum A.Gray • ASTERACEAE : Baileya Harv. & A.Gray ex Torr. : B. multiradiata Harv. & A.Gray ex Torr. • Berlandiera DC. • Chaenactis DC. • Erigeron • Gutierrezia : G. microcephala (DC.) A.Gray • Heliomeris Nutt. : H. multiflora Nutt. • Machaeranthera Nees : M. tanacetifolia (Kunth) Nees • Malacothrix DC. : M. fendleri A.Gray • Pectis L.: P. papposa Harv. & A.Gray • Senecio L. • Stephanomeria Nutt. : S. exigua Nutt. • BIGNONIACEAE Juss. : Chilopsis D.Don • BORAGINACEAE : Eriodictyon Benth. : E. angustifolium Nutt. • Nama L.: N. hispida A.Gray • Phacelia : P. indecora J.T.Howell • BRASSICACEAE : Lepidium : L. thurberi Wooton • Physaria (Nutt. ex Torr. & A.Gray) A.Gray : P. gordonii (A.Gray) O’Kane & Al-Shehbaz • Stanleya Nutt. • EUPHORBIACEAE : Croton : C. wigginsii L.C.Wheeler • Euphorbia : E. fendleri Torr. & A.Gray • FABACEAE : Acacia Mill. : A. greggii A.Gray • Medicago : M. sativa • Parkinsonia L.: P. aculeata L. • Prosopis L.: P. glandulosa Torr. : P. g. var. torreyana (L.D.Benson) M.C.Johnst. • P. velutina Wooton • LAMIACEAE : Salvia : S. dorrii (Kellogg) Abrams • LILIACEAE Juss. : Calochortus Pursh • LOASACEAE Juss. : Mentzelia L.: M. involucrata S.Watson • MALVACEAE : Gossypium • Sphaeralcea A.St. -Hil.: S. ambigua A.Gray • PAPAVERACEAE : Arctomecon Torr. & Frém. : A. californicum Torr. & Frém. • A. merriamii Coville • Argemone • PLANTAGINACEAE Juss. : Penstemon Schmidel : P. palmeri A.Gray • POLEMONIACEAE Juss. : Eriastrum Wooton & Standl. : E. diffusum (A.Gray) H.Mason • POLYGONACEAE Juss. : Eriogonum Michx. : E. abertianum Torr. • E. deflexum Torr. • E. fasciculatum Benth. • E. racemosum Nutt. • E. trichopes Torr. • ROSACEAE Juss. : Heteromeles M.Roem. • SALICACEAE Mirb. : Salix L. • SAPINDACEAE Juss. : Sapindus L.: S. drummondii Hook. & Arn. • SCROPHULARIACEAE : Leucophyllum : L. frutescens • SOLANACEAE Juss. : Chamaesaracha Franch. & Sav. ex A.Gray : C. coniodes (Dunal) Britton • C. coronopus (Dunal) A.Gray • Physalis L.: P. acutifolia (Miers) Sandwith • TAMARICACEAE : Tamarix : T. ramosissima Ledeb. • VERBENACEAE J.St. -Hil.: Verbena L.: V. bracteata Cav. ex Lag. & Rodr. • ZYGOPHYLLACEAE R.Br. : Larrea Cav. : L. tridentata (Sessé & Moc. ex DC.) Coville.

In addition to the above records, some of the floral hosts reported for L. perparvum in Moure & Hurd (1987) are likely attributable to L. deludens sp. nov.

DNA barcodes

Seventeen confirmed sequences available (BOLD process IDs: DLII753-07, DLII763-07, DLII771-07, DLII786-07, DLII793-07, DLII795-07, DLII802-07, DLII806-07, DLII809-07, DLII987-07, DLII1075-07, DLII1077-07, DLII1122-07, DIAL1099-07, DIAL1129-07, DIAL1174-07, DIAL1430-09). There is a moderate amount of divergence within these sequences (0.76% maximum intraspecific p-distance). They are closest in terms of p-distance to L. profundum sp. nov. (0.95% minimum interspecific p-distance). Two fixed nucleotide substitutions distinguish L. deludens sp. nov. from all other Nearctic species of the L. gemmatum complex: 346(C) and 471(C) (Supp. file 2).

Comments

Very common. In the Sonoran Desert, L. deludens sp. nov. is one of the two most abundant species of L. ( Dialictus ) along with L. microlepoides ( Ellis, 1914) . These two species are about equally abundant and together often comprise 50% or more of collections of L. ( Dialictus ) in this area. Many thousands of additional specimens are known to exist but were not examined in this study.

Most specimens determined as L. perparvum in various collections prior to this work were actually L. deludens sp. nov. This could be due to an initial misidentification which was then propagated by later workers. After the error was discovered, L. deludens was usually referred to as “ L. aff. perparvum ” or “ L. nr. perparvum ”. Alternatively, L. deludens could have been called “ L. nr. perparvum ” at first, and later workers disregarded the “near”. Approximate dates of identification (especially by George Eickwort, who determined many specimens) would be necessary to distinguish these scenarios. Morphology and DNA barcodes easily distinguish L. deludens and L. perparvum .

Figures

Fig. 12. Lasioglossum (D.) deludens sp. nov., ♀, holotype (PCYU SBV047813). A. Dorsal habitus. B. Lateral habitus. C. Face. D. Metasoma. Scale bars = 1 mm.

Fig. 13. Lasioglossum (D.) deludens sp. nov., ♂. A. Dorsal habitus. B. Lateral habitus. C. Face. D. Metasoma and propodeum. Scale bars = 1 mm.

Fig. 14. Georeferenced collection records of Lasioglossum (D.) deludens sp. nov. (black squares) and predicted distribution by maximum entropy ecological niche modelling in Maxent (colour shading). Warmer colours indicate higher cloglog probability of occurrence.

Fig. 76. Male genitalia illustrated in dorsal view and positioned such that the gonostylus is parallel to the plane of vision. Retrorse lobes are also illustrated separately to the bottom left of each figure in lateral view, positioned parallel to the plane of vision. A. Lasioglossum (D.) angelicum sp. nov.B. L. (D.) coactus (Cresson, 1872). C. L. (D.) deludens sp. nov. D. L. (D.) diabolicum sp. nov.E. L. (D.) gaudiale (Sandhouse, 1924). F. L. (D.) gloriosum sp. nov. (the retrorse lobe was not visible for illustration). G. L. (D.) helianthi (Cockerell, 1916). H. L. (D.) holzenthali sp. nov. I. L. (D.) indagator sp. nov. J. L. (D.) magnitegula sp. nov. K. L. (D.) perparvum (Ellis, 1914). L. L. (D.) profoundum sp. nov. M. L. (D.) pseudotegulare (Cockerell, 1896). N. L. (D.) rufodeludens sp. nov. This genital capsule was damaged during dissection and some details may be slightly distorted.O. L. (D.) stictaspis (Sandhouse, 1923) (holotype) (reproduced from Gardner & Gibbs 2022). P. L. (D.) tegulariforme (Crawford, 1907) (reproduced from Gardner & Gibbs 2022). Scale bars = 0.25 mm.

Fig. 78. Mesoscutum pubescence. A. Lasioglossum perparvum (Ellis, 1914), ♀, lateral view, with pubescence very short and sparse. B. L. magnitegula sp. nov., ♀, dorsal view, with pubescence very short and sparse. C. L. angelicum sp. nov., ♀, lateral view, with pubescence long and dense. D. L. deludens sp. nov., ♀, dorsal view, with pubescence long and dense. Scale bars = 0.5 mm.

Fig. 102. Face. A. Lasioglossum holzenthali sp. nov., ♂, with dense tomentum on lower paraocular areas and sparser tomentum between. B. L. ellisiae (Sandhouse, 1924), ♂, with dense tomentum below eye emargination. C. L. deludens sp. nov., ♂, with dense tomentum slightly above eye emargination. D. L. gloriosum sp. nov., ♂, with dense tomentum below ocelli. Scale bars = 1 mm.