Hippasella alhue, Piacentini, Luis N., 2011

Piacentini, Luis N., 2011, Three new species and new records in the wolf spider subfamily Sosippinae from Argentina (Araneae: Lycosidae), Zootaxa 3018, pp. 27-49 : 29-33

publication ID	https://doi.org/ 10.5281/zenodo.278639
DOI	https://doi.org/10.5281/zenodo.6189973
persistent identifier	https://treatment.plazi.org/id/03ED87C7-C234-7E24-A5D5-FC99FBC9FB61
treatment provided by	Plazi
scientific name	Hippasella alhue
status	sp. nov.

Treatment

Hippasella alhue View in CoL sp. nov.

( Figs 1–4 View FIGURE 1 View FIGURE 2 View FIGURE 3 View FIGURE 4 , 18 View FIGURE 18 c, 19c, 20)

Type material. Male holotype from Argentina: Neuquén: Cerro Bayo, S 40.74852° W 71.60021° elev. 1404 m GPS, III.2005, Werenkraut, V. (pitfall traps cod. M3S6M05) (MACN-Ar 22101). Three male paratypes (MACN-Ar 20471) from the same locality, data and collector, and three female paratypes (MACN-Ar 20467; temporary preparations LNP-00539, LNP-00540 and LNP-00548) from the same locality and collector, I.2005 (pitfall traps cod. M3S6M05).

Other material examined. ARGENTINA: Río Negro: Cerro Chall-huaco, S 41.23982° W 71.2878°, elev. 1123 m GPS, III.2006, Werenkraut, V. (pitfall traps cod. M5S3M06), 3 males [1 juvenile] (MACN-Ar 22090; temporary preparation LNP-00536); same locality and collector, III.2005 (pitfall traps cod. M5S3M05), 1 male (MACN-Ar 21958); Cerro Chall-huaco, S 41.25967° W 71.29105°, elev. 1404 m GPS, I.2005, Werenkraut, V. (pitfall traps cod. M5S6E05), 1 female (MACN-Ar 21771); same locality and collector, III.2005 (pitfall traps cod. M5S6M05), 2 males (MACN-Ar 21974); Cerro Chall-huaco, S 41.26248° W 71.29703°, elev. 1509 m GPS, III.2006, Werenkraut, V. (pitfall traps cod. M5S7M06), 2 males (MACN-Ar 20442); same locality and collector, I.2005 (pitfall traps cod. M5S7E05), 1 male 1 female (MACN-Ar 21773; temporary preparations LNP-00599 and LNP–00600); same locality and collector, III.2005 (pitfall traps cod. M5S7M05), 1 female (MACN-Ar 21977); Cerro López S 41.09845° W 71.54952°, elev. 1398 m GPS, III.2006, Werenkraut, V. (pitfall traps cod. M1S7M06), 1 male (MACN-Ar 20241); same locality and collector, III.2005 (pitfall traps cod. M1S7M05), 3 males (MACN- Ar 21622); Cerro López S 41.10307° W 71.56223°, elev. 1700 m GPS, I.2005, Werenkraut, V. (pitfall traps cod. M1S10E06), 1 male [1 juvenile] (MACN-Ar 20453); same locality and collector, III.2006 (pitfall traps cod. M1S10M06), 2 males (MACN-Ar 22063); same locality and collector, III.2005 (pitfall traps cod. M1S10M05), 4 male (MACN-Ar 22064). Neuquén: Cerro Pelado S 40.93134° W 71.33495°, elev. 1318 m GPS, III.2005, Werenkraut, V. (pitfall traps cod. M2S6M05), 1 male (MACN-Ar 21670; temporary preparation LNP-00583); Cerro Bayo, S 40.74895° W 71.60832°, elev. 1510 m GPS, I.2005, Werenkraut, V. (pitfall traps cod. M3S7E05), 1 female (MACN-Ar 20476); San Martín de los Andes, Cerro Chapelco (ca. S 40.223600° W 71.288900° -+ 2140 m), II.1961, Galiano, M. E. 1 female [1 juvenile] (MACN-Ar 5318).

Diagnosis. Males of this species can be distinguished from those of H. guaquiensis by the well-developed lateral apophysis of the conductor ( Fig. 2 View FIGURE 2 e). Females differ by the more elongated and squared median septum, not rounded as in H. guaquiensis , and in the triangular head of spermatheca, which is rounded in H. guaquiensis ( Fig. 4 View FIGURE 4 , 18 View FIGURE 18 c and figs 7–9 in Alvares and Brescovit 2007).

Description. Male (holotype). Colour in ethanol ( Fig. 1 View FIGURE 1 ): carapace brown with dark radial pattern, light brown median band narrowing posteriorly, pale submarginal bands enhanced by white setae. Chelicerae, endites and labium, dark brown. Sternum, and coxae, reddish-brown. Femora and patellae light brown with dark spots. Tibiae and metatarsi reddish brown, with light spots. Scopulae dense, on legs I and II from ventral distal sides of tibiae through tarsi, on legs III and IV from metatarsi. Abdomen brown, with two lines of white setae in posterior half, connected by five dark stripes; venter light brown.

Pedipalp ( Fig. 2 View FIGURE 2 ), tibia slightly curved. Cymbium without distal spines. Tegulum with an apical projection in ventral view and sperm duct with sigmoid shape in the ventral part ( Fig. 2 View FIGURE 2 e). Lateral apophysis of conductor well developed, median apophysis small and membranous ( Fig. 2 View FIGURE 2 f).

TL 8.53, CL 4.33, CW 3.27, CH 1.67. Eyes: AME 0.13, ALE 0.15, PME 0.33, PLE 0.27, POQ length 0.50, POQ posterior width 0.78, POQ anterior width 0.25. Anterior eye row slightly procurved, ALE on small tubercles. Chelicerae: 1.42 length, three promarginal teeth, the median largest; retromargin with three teeth of same size. Abdomen: length 4.20, width 2.73. Legs: length of segments (femur + patella/tibia + metatarsus + tarsus = total length): pedipalp 1.33 + 1.47 + - + 1.20 = 4.00, I 2.80 + 3.67 + 2.40 + 1.53 = 10.40, II 2.67 + 3.40 + 2.33 + 1.40 = 9.80, III 2.60 + 3.40 + 2.40 + 1.47 = 9.87, IV 3.13 + 4.00 + 3.40 + 3.87 = 14.40. Leg formula 4132. Spination pattern: femur I p 0-0-d2 d 1-1, II p d1-0-d1 d 1-1-1 r 0-d1-d1, III p d1-0-d1 d 1-1-1 r d1-0-d1, IV p 0-0-d1 d 1-1-1 r 0-0-d1; patella III p 1 r 1, IV p 1 r 1; tibia I p 0-0-1 r 0-0-1 v 2-2 -2ap, II p 1-0-1 r 0-0- 1 v r1-r1-2ap III, p 1-1 d 1 r 1-1 v 2-2 -2ap, IV p 1-1 d 1 r 1-1 v 2-2 -2ap; metatarsus I p 0-d1-1ap r 0-1-1ap v 2-2 -0, II p 0-1-2ap r 0-1-1ap v 2-2 -1ap, III p 1-1-2ap r 1-1-2ap v 2-2 -1ap, IV p 1-1-1ap r 1-1-2ap v 2-2 -2ap.

Female (MACN-Ar 20467). Colour in ethanol ( Fig. 3 View FIGURE 3 , 19 View FIGURE 19 c): as in male.

Epigyne ( Fig. 4 View FIGURE 4 ), median septum wide, covered by small setae, with lateral projections covering part of atrium. Internal genitalia ( Figs 4 View FIGURE 4 b, 18c) spermatheca with stalk long and sigmoid, vulval chamber rounded. Head of spermatheca wide and triangular. Fertilization duct small, membranous, located at posterior margin and curved dorsally.

TL 9.47, CL 4.67, CW 3.40, CH 1.33. Eyes: AME 0.17, ALE 0.15, PME 0.32, PLE 0.27, POQ long 0.55, POQ posterior width 0.92, POQ anterior width 0.25. Anterior eye row slightly procurved, ALE on small tubercles ( Fig. 19 View FIGURE 19 c). Chelicerae: 1.87 length, three promarginal teeth, the median largest; retromargin with three teeth of same size. Abdomen: length 4.80, width 3.20. Legs: (femur + patella/tibia + metatarsus + tarsus = total length): pedipalp 1.33 + 1.33 + - + 1.47 = 4.13, I 2.80 + 3.60 + 1.87 + 1.47 = 9.74, II 2.60 + 3.27 + 2.00 + 1.27 = 9.14, III 2.47 + 2.73 + 2.13 + 1.40 = 8.73, IV 3.07 + 3.73 + 3.47 + 1.53 = 11.80. Leg formula 4123. Spination pattern: femur I p 0- 1-d2 d 1-1, II p 0-0-2 d 1-1, III p 0-d1-d1 d 1-1-1 r 0-0-d1, IV p 0-d1-d1 d 1-1-1 r 0-0-d1; patella III p 1 r 1, IV p 1 r 1; tibia I v p1-2-2ap, II p 0-1- 1 v p1-2-2ap, III p 1-1 r 1-v 1 v p1-p1-2ap, IV p 0-1- v1 r 0-1- 1 v p1-2-2ap; metatarsus I p 0-0-1 r 0-0-1 v 2-2 -1ap, II p 0-1-2 r 0-1-2 v 2-2 -1ap, III p 1-1-2 r 1-1-2 v 2-2 -1ap, IV p 1-1-1 r 1- 1-2 v 2-2 -2ap.

Variation. Female (male) range, mean ± SD. TL 7.80 – 11.53, 9.92 ± 1.40; CL 4.13 – 5.47, 4.20 ± 0.45; CW 2.87 – 4.00, 2.63 ± 0.38; TCR0.94– 1.11, 1.04 ± 0.06; n = 7 (TL 7.20 – 10.13, 7.59 ± 0.78; CL 3.53 – 5.33, 4.39 ± 0.52; CW 2.87 – 3.80, 3.23 ± 0.28; TCR1.07 – 1.37, 1.07 ± 0.09; n = 9).

Etymology. The specific name is a noun in apposition that means soul, ghost or spectre of the dead in the Mapuche language.

Biology. Most of the material was collected by Victoria Werenkraut using pitfall traps in two different habitats: In lenga forest ( Nothofagus pumilo ) between 1100 and 1700 m altitude, and in Andean steppe at 1700 m. The presence of males and females in pitfall samples suggest that this is a wandering species

Distribution. South-western Argentina (Neuquén and Río Negro) ( Fig. 20 View FIGURE 20 ).