Anthobium latissimum Shavrin & Smetana, 2018

Shavrin, Alexey V. & Smetana, Aleš, 2018, A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group, Zootaxa 4508 (4), pp. 451-506 : 480-482

Anthobium latissimum Shavrin & Smetana , sp.n.

( Figs. 49 , 59 , 71–73 )

Type material examined: Holotype ♂: ‘ CHINA: Yunnan [ CH 07-24], Nujiang | Lisu Aut. Pref., Gaoligong Shan , valley 18 | km W Gongshan, 3020 m, 27°47'54''N, | 98°30'13''E, mixed forest, litter, moss, | wood sifted, 7.VI.2007, M. Schülke’ <rectangular label, printed >, ‘ HOLOTYPE | Anthobium | latissimum sp.n. | Shavrin A. & Smetana A. des. 2018’ <red rectangular label, printed> ( CSC) .

Paratype ♂ [specimen dissected; left side of pronotum with artificial dent]: same data as the holotype, with additional red rectangular printed label : ‘ PARATYPE | Anthobium | latissimum sp.n. | Shavrin A. & Smetana A. des. 2018’ ( CS) .

Description. Measurements (n=2): HW: 0.60; HL: 0.35; AL(holotype): 0.92; OL: 0.13–0.15; PL: 0.50–0.58; PW: 1.00–1.02; ESL: 1.15–1.17; EW: 1.17–1.23; AW: 1.07–1.08; MTbL(holotype): 0.60, MTrL(holotype): 0.27 (MTrL 1–4: 0.16; MTrL 5: 0.11); AedL(paratype): 0.62; TL: 2.56–2.95(holotype).

Body elongate, subconvex. Pronotum, elytra and antennomeres 6–11 yellow-brown; head reddish-brown; mouthparts, antennomeres 1–5, lateral and basal margins of pronotum and legs yellow. Lateral portions of head on infraorbital ridge with coarsely rugose sculpture, sometimes with distinct small wrinkles on posterior portions (holotype); dorsal part of neck with transverse microsculpture; abdominal tergites with fine isodiametric microsculpture. Frons and middle portion of head with irregular, sparse, small and deep punctation, denser on lateral parts of infraorbital ridges and between ocelli; middle part of neck with regular, small and deep punctation; pronotum with irregular and deep punctation, significantly denser than that on middle part of head; scutellum with several small punctures; punctation of elytra as that on pronotum, slightly denser on parascutellar area, not forming longitudinal rows, with indistinct elevations between punctures on middle part of each elytron; abdominal terites without distinct punctation. Habitus as in Fig. 49 .

Head transverse, 1.7 times as wide as long, slightly elevated in middle portion, with moderately flattened infraorbital ridges; middle part of head between posterior part of eyes with irregular shallow impression, more distinct in holotype; deep, narrow and moderately short grooves in front of ocelli reaching posterior third of length of eyes; posterior portions with distinct, moderately acute postocular ridges; anterior portion between antennal insertion and anterior margin of eye with superficial notch. Ocelli situated at level of postocular ridge; distance between ocelli distinctly longer than distance between ocellus and posterior margin of eye. Apical segment of maxillary palp elongate, about three times as long as preceding segment, markedly narrowed apicad from widest basal portion. Antenna slightly exceeding prescutellar area of elytra when reclined; length × width of antennomeres (holotype): 1: 0.13 × 0.06; 2: 0.10 × 0.05; 3: 0.08 × 0.04; 4: 0.08 × 0.05; 5: 0.08 × 0.07; 6–8: 0.07 × 0.05; 9: 0.07 × 0.06; 10: 0.06 × 0.07; 11: 0.11 × 0.07.

Pronotum wide and massive in general appearance, with widely rounded apical and basal margins, twice as wide as long, slightly narrower than elytra,1.6–1.7 times as wide as head, widest in middle, slightly more narrowed posteriad than anteriad; apical margin about as long as posterior margin; anterior angles widely rounded, distinctly protruded anteriad; posterior angles obtuse; lateral edges of pronotum with irregular small crenulation; pronotum with very wide middle elevation, with very small and indistinct longitudinal impression and irregular transverse impressions on mediobasal third; disc of pronotum with moderately strong small elevations between punctures in middle part.

Elytra narrow and elongate, indistinctly widened apicad from about middle, reaching basal margin or middle part (paratype) of abdominal tergite V, about as long as wide, more than twice as long as pronotum; each elytron with indistinctly elevated, small parascutellar and latero-apical portions.

Male. First four protarsomeres distinctly dilated. Apical margins of abdominal tergite VIII and sternite VIII slightly emarginated. Aedeagus ( Fig. 71 ) with median lobe wide, with wide, truncate apex; parameres each with widened apical portion, slightly assymetrical, exceeding apex of median lobe, with two short apical and preapical setae; internal sac elongate, spiralled in basal portion. Aedeagus laterally as in Fig. 72 .

Female unknown.

Comparative notes. Anthobium latissimum Shavrin & Smetana , sp.n. may be distinguished from all remaining species of the fusculum group by the shape of the massive pronotum, with markedly rounded apical and basal margins, and by the shape and internal structure of the aedeagus.

Distribution. The new species is at present known only from the type locality in the Gaoligong Shan range in Yunnan, China ( Fig. 59 ).

Bionomics. The specimen was taken by sifting litter, moss and wood in forest at elevation of 3020 m a.s.l. (locality DSCN7372: Fig. 73 ).

Etymology. The specific epithet is the Latin latissimus, -a, -um (the broadest). It refers to the very wide pronotum.

Figures

FIGURE 59. Distribution of Anthobium capitale (black circles), A. laozii (black squares) and A. latissimum (black triangle) in Yunnan, China.

FIGURE 73. Type locality of Anthobium latissimum in the Gaoligong Shan range in Yunnan, China (photograph: M. Schülke).

FIGURES 45–49. Habitus of Anthobium fusculum group: 45—A. confucii (paratype; CHINA: Hubei), 46—A. cuccodoroi (paratype), 47—A. denticulatum (paratype), 48—A. laozii (paratype; CHINA: Yunnan, Cangshan), 49—A. latissimum (holotype). Scale bar: 1.0 mm.

FIGURES 68–72. Aedeagus of Anthobium laozii (68–69: paratype; CHINA: Yunnan, Diancang Shan, 70: paratype; CHINA: Cangshan) and A. latissimum (71–72: paratype): 68, 71—parameral view, 69, 72—lateral view, 70—apical part of aedeagus, parameral view. Scale bar: 0.1 mm.