Tomoglossa arizonica Gusarov, 2002

Gusarov, Vladimir I., 2002, A revision of Nearctic species of the genus Tomoglossa Kraatz, 1856 (Coleoptera: Staphylinidae: Aleocharinae), Zootaxa 30, pp. 1-19 : 14-15

4. Tomoglossa arizonica Gusarov , sp. nov. ( Figs. 32 , 38­42 )

Type material: holotype ,, UNITED STATES: Arizona: Graham Co., Roper Lake State Park , 5 miles S Safford, [32º45'N 109º42'W], 900m, 12.v.1986 (L.Herman) ( AMNH) .

Diagnosis. Tomoglossa arizonica differs from T. ornatella , T. improvisa and T. decora in having transverse antennal articles 4­5 ( Figs. 29­32 ) and shorter body. Compared to T. floridana , in T. arizonica antennal articles 4­10 are less transverse, article 11 is as long as 9 and 10 combined ( Figs. 32­33 ) and the body is longer. Tomoglossa arizonica also differs from T. decora in having shorter apex of median lobe of aedeagus (in lateral view) ( Figs. 40 , 21 ).

Description. Length 1.8 mm. Head, elytra, abdominal tergum VI, antennal articles 2­ 11 and article 3 of maxillar palpus reddish brown; pronotum and abdominal segments III­ V and VII­VIII reddish yellow; apical quarter of elytra, legs, mouthparts and antennal article 1 yellow.

Head transverse, surface on disk glossy, without visible (at x100) microsculpture, with fine and dense punctures, distance between them equal to ½ their diameter. Antennal article 2 longer than article 3, 4­10 transverse, last article as long as 9 and 10 combined ( Fig. 32 ).

Pronotum transverse, width to length ratio 0.37 mm: 0.29 mm, wider than head (pronotum width to head width ratio 0.37 mm: 0.31 mm); surface glossy, without visible microsculpture, with puncturation as on head. Elytra measured from humeral angle shorter than wide (0.36 mm: 0.43 mm), glossy, without microsculpture, puncturation finer and denser than on pronotum, distance between punctures equal to ¼­½ their diameter.

Abdominal terga glossy, without microsculpture, with fine and sparse puncturation, puncturation becoming finer towards abdominal apex, on terga III­V distance between punctures within rows equal to their diameter and between rows 2­3 times their diameter, on tergum VI distance between punctures within rows equal to 1­2 times their diameter and between rows 2­4 times their diameter, on tergum VII distance between punctures within rows equal to 2­3 times their diameter and between rows equal to 3­6 times their diameter. Tergum VII with white edge. Median lobe of aedeagus with narrow apex ( Figs. 41 ).

Distribution. Known only from Arizona ( Fig. 47 ).

Natural History. No habitat information is provided on the holotype label.

Figures

FIGURE 47. Geographical distribution of Nearctic Tomoglossa. T. ornatella (Casey) (c), T. improvisa (Casey) (⌂), T. decora (Casey) (z), T. arizonica Gusarov, sp. nov. (S), T. floridana Gusarov, sp. nov. („).

FIGURES 29 ­ 33. Antennae of Nearctic Tomoglossa. 29, T. ornatella (Casey) (lectotype); 30, T. improvisa (Casey) (lectotype); 31, T. decora (Casey) (Pennsylvania); 32, T. arizonica Gusarov, sp. nov. (holotype); 33, T. floridana Gusarov, sp. nov. (holotype). Scale bar 0.1 mm.

FIGURES 38 ­ 42. Details of Tomoglossa arizonica Gusarov, sp. nov. (holotype). 38, male tergum VIII; 39, male sternum VIII; 40, median lobe of aedeagus, lateral view; 41, median lobe of aedeagus, ventral view; 42, internal sac of aedeagus, in packed state. Scale bar 0.2 mm (38 ­ 39), 0.1 mm (40 ­ 42).

FIGURES 21 ­ 28. Genitalia and terminalia of Tomoglossa decora (Casey) (Vermont (21 ­ 25), Pennsylvania (26 ­ 28 )). 21, median lobe of aedeagus, lateral view; 22, median lobe of aedeagus, ventral view; 23, internal sac of aedeagus, in packed state; 24, deeply concave sclerite of internal sac; 25, copulatory piece; 26, spermatheca; 27, left portion of female sternum IX; 28, female accessory sclerite. U ­ umbilicus. Scale bar 0.1 mm.