Angaracris Bey-Bienko, 1930

Storozhenko, S. Yu. & Benediktov, A. A., 2020, AN ANNOTATED CHECK-LIST OF ORTHOPTERA OF TUVA AND ADJACENT REGIONS. PART 3. SUBRODER CAELIFERA (ACRIDIDAE: GOMPHOCERINAE: GOMPHOCERINI; LOCUSTINAE), Far Eastern Entomologist 402, pp. 1-36 : 26-28

Genus Angaracris Bey-Bienko, 1930

27(91). Angaracris barabensis (Pallas, 1773)

Bryodema barabense: Miram, 1907: 8 .

Bryodema barabense var. rhodopa: Miram, 1907: 8

Angaracris barabensis: Berezhkov, 1951: 21 ; Sergeev et al., 1995: 97–98; Benediktov, 1997: 117;

Benediktov, 1998: 792; Storozhenko et al., 2017.

Angaracris rhadopus (sic!): Berezhkov, 1951: 21

Angaracris rhodopa: Sergeev et al., 1995: 97–98 ; Benediktov, 1997: 117.

Angaracris rodopa (sic.!): Pavlov, 2004: 65.

MATERIAL. Tuva: W Sayan Mts., Alash Plateau, Ak-Sug River, 51°24ʹN, 90°28ʹE, 1075–1077

m, terraces, steppe and meadow, 12.VIII 2016, 2 ♂ ( MS); W Sayan Mts. , Alash Plateau, Ak-Sug River ,

51°24ʹN, 90°27ʹE, 1079–1085 m, southern slope, stony semi-desert, 13.VIII 2016, 5 ♂ ( MS); W Sayan

Mts., Alash Plateau, Ak-Sug River, 51°25ʹN, 91°04ʹE, 800–810 m, southern slope and upper terrace,

semi-deserts, 13.VII 2016, 9 ♂, 3 ♀ ( MS); Sayan Mts. , Alash Plateau, Ak-Sug River, 51°31ʹN, 90°12ʹE ,

1275 m, southern slope, steppe, 14.VIII 2016, 6 ♂ ( MS); W Sayan Mts. , Alash Plateau, Ak-Sug River ,

51°30ʹN, 90°12ʹE, 1250–1252 m, lower terrace and upper flood-plain, meadows, pebbles, 14.VIII 2016,

8 ♂, 1 ♀ ( MS); Turan-Uyuk Intermountain Basin, Begreda River , 51°59ʹN, 94°18ʹE, 830–845 m, southern slope and piedmont plain, semi-deserts, 12.VIII 2018 , 1 ♂, 2 ♀ ( MS); E Tuva, Academician Obruchev

Range , southern slope, Koptu (Khapto) River, 51°37ʹN, 95°21ʹE, 788–843 m, southern slope and upper terraces, steppes, 11.VIII 2018, some specimens observed ( MS); Chaa-Khol River, 51°34′N, 92°23′E ,

570–600 m, dry steppes, 14–18.VII 1962, 2 ♂, 1 ♀, 8 larvae ( IS); 32 km SW Kyzyl City, Elegest River ,

51°29ʹN, 94°10ʹE, steppe, 22.VII 2014, 2 ♀ ( SS); Ulug-Khem Intermountain Basin , 25 km SW Kyzyl

City, Ulug River, 51°35ʹN, 94°09ʹE, 630–830 m, mountain slopes and piedmont plain, semi-deserts with

Nanophyton grubovi , 19.VII 1995, 1 ♂ ( MS); Ulug-Khem Intermountain Basin , 25 km SW Kyzyl City ,

Ulug River, 51°35ʹN, 94°08ʹE, 608 m, upper terrace, semi-deserts with Nanophyton grubovi , 20.VII 1995,

1 ♂, 1 ♀ ( MS); Ulug-Khem Intermountain Basin , 25 km SW Kyzyl City, Ulug River, 51°34ʹN, 94°10ʹE ,

800–900 m, southern slope and piedmont plain, semi-deserts with Nanophyton grubovi , 22.VII 2003, 5 ♂,

1 ♀ ( MS); Ulug-Khem Intermountain Basin , Elegest River, middle part, 51°22ʹN, 94°04ʹE, 695 m, upper terrace, dry steppe, 16.VI 2017, a specimen observed ( MS); Kaa (Malyj Yenissei) River, right side, near

Boyarovka settlement, 51°32ʹN, 95°21ʹE, 703 m, plain, dry steppe, 12.VIII 2018, 2 ♂, 1 ♀ ( MS); SE

Ulug-Khem Intermountain Basin, Shuurmak River, 25 km S Balgazyn settlement, near Kuran settlement,

50°47ʹN, 95°17ʹE, 1030–1050 m, southern slope, steppe and meadow, 10.VII 1978, 2 larvae ( MS); W

Tannu-Ola Mts., 5 km N Khandagajty settlement, 50°48ʹN, 92°05ʹE, 1300–1350 m, piedmont plain,

stony steppes with Caragana , 16.VIII 1985, 27 ♂, 11 ♀ ( MS); W Tannu-Ola Mts. , 9 km N Khandagajty settlement, 50°51ʹN, 92°04ʹE, 1700–1900 m, mountain steppes, 19.VIII 1985, 1 ♂, 1 ♀ ( MS); W Tannu-Ola

Mts., 9 km N Khandagajty settlement, 50°50ʹN, 9°05ʹE, 1637 m, mountain steppes, 19.VIII 1985, 6 ♂, 1

♀ ( MS); W Tannu-Ola Mts. , NW Khandagajty settlement, near Solchur settlement, 50°46ʹN, 92°01ʹE ,

1150–1200 m, agricultural fields including abandoned and plots along canal and roads, 21.VIII 1978, 34 ♂,

32 ♀ ( MS); W Tannu-Ola Mts. , 5 km N Khandagajty settlement, 50°47ʹN, 92°03ʹE, 1193 m, piedmont plain, lower terrace of stream, meadow, 21.VIII 1985 , 1 ♂ ( MS); W Tannu-Ola Mts. , 5 km N

Khandagajty settlement, 50°47ʹN, 92°05ʹE, 1200–1250 m, piedmont plain, lower part, stony steppes with

Caragana and dry meadow, 21.VIII 1985, 12 ♂, 1 ♀ ( MS); W Tannu-Ola Mts. , 5 km N Khandagajty settlement, 50°48ʹN, 92°04ʹE, 1336 m, foothills on piedmont plain, stony steppe, 21.VIII 1985, 3 ♂, 1 ♀

( MS); W Tannu-Ola Mts. , 5 km N Khandagajty settlement, 50°50ʹN, 92°05ʹE, 1500-1550 m, piedmont plain, upper part, steppe, 21.VIII 1985, 16 ♂, 8 ♀ ( MS); W Tannu-Ola Mts., 4 km E Khandagajty settlement, 50°45ʹN, 92°09ʹE, 1150–1200 m, southern slope and piedmont plain, stony semi-desert with

Nanophyton grubovii , 24.VIII 1985, 6 ♂, 11 ♀, 3 larvae ( MS); W Tannu-Ola Mts. , 4 km E Khandagajty settlement, Mugur River, 50°44ʹN, 92°08ʹE, 1100–1150 m, low terrace and upper flood-plain, meadows ,

24.VIII 1978, 1 ♀ ( MS); W Tannu-Ola Mts. , near Khandagajty settlement, 50°43′N, 92°04′E , 1170–1200

m, piedmont plain, desert with Nanophyton grubovii , 24.VIII 1985, 3 ♂ ( MS); W Tannu-Ola Mts. , near

Khandagajty settlement, 50°43′N, 92°03′E, 1200–1250 m, piedmont plain and northern slope, semi-desert with Caragana bushes, 24.VIII 1985, 10 ♂, 7 ♀ ( MS); E Tannu-Ola Mts. , Shivelig River, 50°45ʹN ,

94°34ʹE, 1330–1340 m, flood-plain and terraces, steppe and short meadow between shrubs and Larix trees, 28.VI 1978, 8 larvae ( MS); E Tannu-Ola Mts. , Shivelig River, 50°45ʹN, 94°34ʹE, 1330–1340 m ,

terraces, steppe and short meadow between shrubs and Larix trees, 17.VII 1978, 3 larvae ( MS) ; the same locality, upper terrace, short meadow and steppe, 12.VIII 1978, 43 ♂, 12 ♀, 7 larvae ( MS) ; the same locality, lower terrace, steppe, 12.VIII 1978, 1 ♂ ( MS) ; E Tannu-Ola Mts. , Shuurmak River, 5 km N

Shuurmak settlement, 50°41ʹN, 95°19ʹE, 1173 m, local southern slope and piedmont plain, steppe,

10.VII 1978, 2 larvae ( MS); E Tannu-Ola Mts. , Shuurmak Pass, 50°37ʹN, 95°11ʹE, 1495 m, southern slope and upper terrace, mountain steppe , 12.VII 1978, 1 larvae ( MS); N Uvs-Nuur Intermountain Basin ,

Iribitej River , 50°45′N, 93°09′E, 1030–1040 m, upper terraces, dry meadows and steppes, 29.VII 1978 , 2

♂, 1 ♀, 2 larvae ( MS); N Uvs-Nuur Intermountain Basin, Iribitej River , 50°44′N, 93°08′E, 938 m, lower terrace and flood-plains, steppe, 30.VII 1978 , 13 ♂, 2 ♀ ( MS); N Uvs-Nuur Intermountain Basin, Iribitej

River, 50°44′N, 93°08′E, 938 m, upper terrace and piedmont plain, semi-desert with Nanophyton grubovii ,

30.VII 1978, 7 ♂, 7 ♀ ( MS); N Uvs-Nuur Intermountain Basin, 50 km E Amdaygyn-Khol , semi-desert with Nanophyton grubovii , 26.VII 1962, 2 ♂, 1 ♀, 3 larvae ( IS); NE Uvs-Nuur Intermountain Basin ,

Shivelig River , 50°43ʹN, 94°33ʹE, 1215 m, lower terrace, dry steppe, 15.VII 1978, 2 ♂ ( MS); the same locality and habitats, 11.VIII 1978, 41 ♂, 3 ♀, larvae ( MS); NE Uvs-Nuur Intermountain Basin, Shivelig

River, 50°39ʹN, 94°28ʹE, 990–995 m, flood-plain and terraces, dry steppes with Caragana , 18.VII 1978,

13 larvae ( MS); the same locality and habitats, 10.VIII 1978 , 57 ♂, 4 ♀ ( MS); NE Uvs-Nuur Intermountain

Basin, Shivelig River, 50°45ʹN, 94°34ʹE, 1280–1301 m, terraces, dry meadows, 8.VII 2017, some specimens observed ( MS); NE Uvs-Nuur Intermountain Basin, Shivelig River, 50°43ʹN, 94°33ʹE,

1200–1300 m, southern slopes and piedmont plain, steppes with Caragana , 24.VII 2003, 3 larvae ( MS) ;

NE Uvs-Nuur Intermountain Basin, Shivelig River, 50°43ʹN – 50°45ʹN, 94°33ʹE – 94°34ʹE, 1200–

1650 m, mountain slopes, piedmont plains, terraces, dry meadows and steppes, often with bushes and stones, 24.VI–12.VIII 1978,> 100 specimens, including larvae ( T. Myagkaya, A. Lee); Uvs-Nuur

Intermountain Basin, Tes River, 50°32ʹN, 94°31ʹE, 930–932 m, upper flood-plain, meadows, 13.VII

1978, 1 ♂ ( MS); the same locality, upper flood-plain and terraces, 11.VIII 1978 , 5 ♂ ( MS); DyttygKhem River , 12 km SW Samagaltai settlement, 50°38ʹ N, 95°19ʹ E, 17.VII 2014 , 2 ♂ ( SS); DyttygKhem River , 13 km SW Samagaltai settlement, 8–10.VII 2013 , 1 ♂, 1 ♀ (M. Prostshalykin, V.

Loktionov); Uvs-Nuur Intermountain Basin, Dus-Khol, 50°22ʹN, 94°52ʹE, 994–995 m, semi-desert with

Nanophyton grubovii , 24–26.VII 1962, 2 larvae ( IS); Uvs-Nuur Intermountain Basin , 7 km S Erzin settlement, Tes River, 50°11ʹN, 95°12ʹE, 1097 m, upper flood-plain, meadows , 21.VII 1978, 1 ♂ ( MS) ;

Uvs-Nuur Intermountain Basin, 7 km S Erzin settlement, Tes River , 50°11ʹN, 95°12ʹE, 1097 m, lower terrace and upper flood-plains, dry meadows and steppes, 22.VII 1978, 19 ♂, 7 ♀, 4 larvae ( MS); UvsNuur Intermountain Basin, 10 km S Erzin settlement, Tsuger-Els, 50°10ʹN, 95°11ʹE, 1154 m, sandy semi-desert, 23.VII 1978, 1 ♂ ( MS); Uvs-Nuur Intermountain Basin, 7 km S Erzin settlement, Tes River ,

50°11ʹN, 95°12ʹE, 1097 m, lower terrace, steppes, 23.VII 1978, 10 ♂, 4 ♀, 1 larva ( MS); Uvs-Nuur

Intermountain Basin, E Shara-Nuur, Yamalyg farwell rocks, 50°14ʹN, 94°45ʹE, 1150 m, piedmont plain,

dry steppe, 6.VII 2017, some specimens observed ( MS); 25 km SW Erzin settlement, Tes River,

50°05ʹN, 95°21ʹE, 14–15.VII 2014, 2 ♂ ( SS); 31 km NEE Erzin settlement, Erzin River , 50°21ʹN ,

95°34ʹE, 1300 m, northern slope, balka with Caragana , 18.VII 2014, 1 ♀ ( SS) .

DISTRIBUTION. Tuva: WSW, WSE, ET, MT, KHE, UKH, KKH, WTO, ETO, UVS,

SAN. – S Siberia, Amur Region; N Kazakhstan, Mongolia, N, NE China.

ECOLOGY. One of the most common and widely distributed species in the typical and dry steppes.