Somatina virginalis Prout, 1917

Sihvonen, Pasi, 2005, Phylogeny and classification of the Scopulini moths (Lepidoptera: Geometridae, Sterrhinae), Zoological Journal of the Linnean Society 143 (4), pp. 473-530 : 514-518

publication ID	https://doi.org/ 10.1111/j.1096-3642.2005.00153.x
persistent identifier	https://treatment.plazi.org/id/03E78792-E10E-2F58-D4A6-8ECBFE3FF969
treatment provided by	Diego
scientific name	Somatina virginalis Prout, 1917
status

Treatment

[ Somatina virginalis Prout, 1917 View in CoL should probably be combined with

Problepsis . Wings have weakly ocellated discal spots with iridescent scales not seen in Somatina . S. virginalis has two areoles on its forewing and this has been considered a feature of the genus. My cladistic analysis shows that this character is homoplastic and that

Problepsis , among others, has one or two areoles. Female genitalia (slide BMNH GEO 4275) also suggest affinity with Problepsis . Somatina virginalis is for the time being retained in Somatina , however, as

I have not yet prepared type specimen of this species, deposited at BMNH .]

wiltshirei Prout, 1938

Zythos

Zythos Fletcher, 1979 . Type species: Nobilia turbata Walker,

1862. Malaysia. [Replacement name for Nobilia Walker, 1861 ]. Nobilia Walker, 1861 . Type species: Nobilia turbata Walker, 1862 . Malaysia. [Junior homonym of Nobilia Gray, 1855 (Mollusca) .]

aphrodite (Prout, 1932) *

ssp. rooki (Prout, 1938)

avellanea (Prout, 1932) *

clypeata Yazaki, 1996*

cupreata (Pagenstecher, 1888)

nebulosa (Warren, 1897)

erotica (Prout, 1932) *

fastigata (Prout, 1938) *

modesta Yazaki, 1996 *

molybdina (Prout, 1938)

ssp. tombarensis (Prout, 1938)

obliterata (Warren, 1897) *

simplaria (Snellen, 1899)

strigata (Warren, 1896) *

ssp. rubescens (Prout, 1938)

turbata (Walker, 1862) *

strigularia (Snellen, [1886])

Lipomelia

Lipomelia Warren, 1893 . Type species: Lipomelia subusta Warren, 1893 . India.

Defoa Swinhoe, 1893 . Type species: Defoa ustata , a junior synonym of Lipomelia subusta Warren, 1893 . [ India].

subusta Warren, 1893 *

ustata (Swinhoe, 1893)

Problepsis

Problepsis Lederer, 1853 . Type species: Caloptera ocellata Frivaldszky, 1845 . [ Greece]. [Replacement name for Caloptera Frivaldszky, 1845 .]

Argyris Guenée, [1858] . Type species: Argyris ommatophoraria Guenée, [1858] , a junior synonym of Problepsis ocellata Frivaldszky, 1845 . [ Lebanon.]

Caloptera Frivaldszky, 1845 . Type species: Caloptera ocellata Frivaldszky, 1845 . [ Greece]. [Junior homonym of Caloptera Gistl, 1834 (Coleoptera) .]

Problepsiodes Warren, 1899 . Type species: Problepsis conjunctiva Warren, 1893 . India.

achlyobathra Prout, 1928 *

ssp. emphyla Prout, 1938 *

ssp. violescens Prout, 1934

aegretta Felder & Rogenhofer, 1875 *

egretta Hale-Carpenter, 1932 [misspelling of aegretta ]

ssp. insculpta Prout, 1917

albidior Warren, 1899 *

ssp. matsumurai Prout, 1938

apollinaria (Guenée, [1858]) *

ssp. aphylacta Prout, 1938

ssp. candidior Prout, 1917

ssp. deparcata Prout, 1925

ssp. hemicyclata Warren, 1897

ssp. wilemani West, 1930

argentea Warren, 1900

asira Wiltshire, 1982 *

borneamagna Holloway, 1997*

centrophora (Prout, 1915) comb. nov. *

[ Somatina centrophora Prout, 1915 is transferred from Somatina to Problepsis .]

clemens Lucas, 1890 *

margaritata Warren, 1896

conjunctiva Warren, 1893 *

ssp. subjunctiva Warren, 1893

craspediata Warren, 1897

ssp. frosti Prout, 1938

ssp. rotifera Prout, 1916

crassinotata Prout, 1917 *

deducta Herbulot, 1962

deliaria Guenée, [1858] *

delphiaria Guenée, ([1858]) *

ssp. auriculifera Warren, 1897

argentisquama (Warren, 1899)*

diazoma Prout, 1938

digammata Kirby, 1896 *

digammata Warren, 1897

[Junior primary homonym and a junior synonym of P. digammata Kirby, 1896 . P. digammata Warren, 1897 was erroneously listed as a junior synonym of P. aegretta aegretta Felder & Rogenhofer, 1875 (Vári, Kroon & Krüger, 2002).]

digrammata Prout, 1933 [misspelling of digammata ] discophora Fixsen, 1887 *

superans coreana Bryk, 1949

ssp. kardakoffi Prout, 1938 *

erythra Wiltshire, 1982 *

eucircota Prout, 1913 *

evanida Prout, 1932

exanimata Prout, 1935

flavistigma Swinhoe, 1904 *

ssp. dilatistigma Prout, 1917

herbuloti Viette, 1968

insignita Prout, 1938

korinchiana Rothschild, 1920 *

latonaria Guenée, ([1858]) *

longipannis Prout, 1917

lucifimbria (Warren, 1902)

magna Warren, 1906 *

maxima Thierry-Mieg, 1905

meroearia Saalmüller, 1884 *

mayottaria (Oberthür, 1923)

metallopictata (Pagenstecher, 1888)

venus Thierry-Mieg, 1905

minuta Inoue, 1958 *

mitis Joannis, 1932

neumanni Prout, 1932

ocellata (Frivaldszky, 1845) *

ommatophoraria Guenée, ([1858])

ssp. cinerea (Butler, 1886) *

ochripicta Warren, 1901

paredra Prout, 1917*

phoebearia Erschov, 1870 *

[Replacement name for Argyris deliaria Bremer, 1864 .]

deliaria (Bremer, 1864)

[Junior primary homonym of Argyris deliaria Guenée, [1858] .] plagiata (Butler, 1881) *

riminota Prout, 1938

plenorbis Prout, 1917 *

rorida Prout, 1932 *

sancta Meyrick, 1888

shirozui Inoue, 1986 *

similinotata Prout, 1917 *

subreferta Prout, 1935

superans Butler, 1885 *

ssp. summa Prout, 1935

transposita Warren, 1903

triocellata Bastelberger, 1908 . comb. rev. *

[ Problepsis triocellata (Bastelberger, 1908) is transferred from Somatina to Problepsis .]

ssp. scenica (Prout, 1938)

vulgaris Butler, 1889 *

attenuata Warren, 1909

Scopula

Scopula Schrank, 1802 . Type species: Phalaena paludata Linnaeus, 1767 , a junior synonym of Phalaena ornata Scopoli, 1763 . [ Portugal].

Acidalia Bruand, 1846 . Type species: Geometra strigaria Hübner, [1799] , a junior synonym of Geometra virgulata [Denis & Schiffermüller], 1775. [ Austria]. [Junior homonym of Acidalia Hübner, [1819] 1816 ( Lepidoptera : Nymphalidae ).]

Acidalia Treitschke, 1825 . Type species: Geometra strigaria Hübner, [1799] , a junior synonym of Geometra virgulata [Denis & Schiffermüller], 1775. [ Austria]. [Junior homonym of Acidalia Hübner, [1819] 1816 ( Lepidoptera : Nymphalidae ).] Acidalina Staudinger, 1898 . Type species: Acidalina decolor, Staudinger, 1898 . Algeria.

Aletis Hübner, [1820] . Type species: Papilio helcita Linnaeus, 1763 . Indiis [incorrect locality, probably tropical Africa]. syn. nov.

Anacosymbia Prout, 1913 . Type species: Anacosymbia perstrigulata Prout, 1913 . [ South Africa] syn. nov.

Antanepsia Turner, 1908. Type species: Autanepsia poliodesma Turner, 1908 . Australia [missspelling of Autanepsia ]

Antilycauges Prout, 1913 . Type species: Emmiltis pinguis Swinhoe, 1902 . [ Taiwan]. syn. nov.

Antitrygodes Warren, 1895 . Type species: Macaria divisaria Walker, 1861 . [ India]. syn. nov.

Autanepsia Turner, 1908 . Type species: Autanepsia poliodesma Turner, 1908 . Australia. syn. nov.

Calothysanis Hübner, [1823] 1816 . Type species: Geometra imitaria Hübner, [1799] 1796 . Europe.

Cartaletis Warren, 1894 . Type species: Aletis libyssa Hopffer, 1858 . Mozambique. syn. nov.

Chlorocraspedia Warren, 1899 . Type species: Chlorocraspedia ansorgei Warren, 1899 . Uganda.

Cinglis Guenée, [1858] . Type species: Fidonia humifusaria Eversmann, 1837 . [ Russia]. syn. nov.

Craspedia Hübner, [1825] 1816 : Type species: Phalaena ornata Scopoli, 1763 . [ Italy].

Csopula Fischer von Röslerstamm, 1842 . [misspelling of Scopula ].

Cymatida Sodoffsky, 1837 . Type species: Geometra strigaria Hübner, [1799] 1796 , a junior synonym of Geometra virgulata [Denis & Schiffermüller], 1775. [ Austria]. [Unnecessary replacement name for Acidalia Treitschke, 1825 ].

Cymatoides Agassiz, 1847 . [Emendation of Cymatida Sodoffsky, 1837 ].

Dasybela Turner, 1908 . Type species: Emmiltis achroa Lower, 1902 . Australia. syn. nov.

Dasybella Turner, 1908 [misspelling of Dasybela ]

Dosithea Duponchel, 1829 . Type species: Phalaena ornata Scopoli, 1763 . [ Portugal].

Dosithoea Rambur, 1833 [misspelling of Dosithea ] Epicosymbia Warren, 1897 . Type species: Epicosymbia perrufa Warren, 1897 , a junior synonym of Acidalia ? dentisignata Walker, [1863] . [ South Africa]. syn. nov.

Eucidalia Sterneck, 1941 . Type species: Phalaena immorata Linnaeus, 1758 . Europe.

Glossotrophia Prout, 1913 . Type species: Acidalia confinaria Herrich-Schäffer, 1847 . Hungary. syn. nov.

Holarctias Prout, 1913 . Type species: Haematopis sentinaria Geyer, 1837 . [ Canada].

Ignobilia Prout, 1932 . Type species: Ephyra urnaria Guenée, [1858] . Borneo. syn. nov.

Induna Warren, 1897 . Type species: Induna rufisalsa Warren, 1897 . [ South Africa].

Isoplenia Warren, 1897 . Type species: Isoplenia trisinuata Warren, 1897 . [ South Africa]. syn. nov.

Leucoxena Warren, 1900 . Type species: Leucoxena lactea Warren, 1900 . [ Kenya]. syn. nov.

Leptaletis Warren, 1894 . Type species: Aletis variabilis Butler, 1878 . [ Angola]. syn. nov.

Leptomeris Hübner, [1825] 1816 . Type species: Geometra umbelaria Hübner, [1813] 1796 . Europe.

Lipocentris Warren, 1905 . Type species: Lipocentris rubriceps Warren, 1905 . Angola.

Longula Staudinger, 1892 . Type species: Longula extraordinaria Staudinger, 1892 , a junior synonym of Lycauges donovani Distant, 1892 . [ Lebanon].

Lycauges Butler, 1879 . Type species: Lycauges lactea Butler, 1879 , a junior synonym of Acidalia emissaria Walker, 1861 . Japan.

Oar Prout, 1913 . Type species: Phalaena pratana Fabricius, 1794 . [ North Africa ]. syn. nov.

Phyletis Guenée, [1858] . Type species: Phyletis silonaria Guenée, [1858] . [ Ethiopia].

Pigia Guenée, [1858] . Type species: Pigia infantularia Guenée, [1858] , a junior synonym of Phalaena opicata Fabricius, 1798 . [ Sri Lanka].

Pleionocentra Warren, 1898 . Type species: Scopula plionocentra Prout, 1920 . [ Geometra minorata sensu Warren , nec Boisduval 1833]. Niger.

Prasonesis Meyrick, 1889 . Type species: Prasonesis microphylla Meyrick, 1889 . [ Papua New Guinea]. syn. nov.

Pseudocinglis Hausmann, 1994 . Type species: Glossotrophia eurata Prout, 1913 . Turkmenistan. syn. nov.

Psilephyra Bastelberger, 1909 . Type species: Psilephyra bilineata Bastelberger, 1909 , a junior synonym of Induna curvimargo Warren, 1900 . [ Tanzania].

Pylarge Herrich-Schäffer, 1855 . Type species: Idaea commutata Freyer, [1832] 1833 , a junior synonym of Scopula ternata, Schrank, 1802 . [ Germany].

Runeca Moore, 1888 . Type species: Runeca ferrilineata Moore, 1888 . India.

Sarodria Sodoffsky, 1837 . [Unnecessary replacement name for Scopula Schrank, 1802 .]

Sarothria Agassiz, 1847 . [Emendation of Sarodria Sodoffsky, 1937 ].

Scopuloides Hausmann, 1994 . Type species: Acidalia fucata Püngeler, 1909 . [ Kyrgyzstan] syn. nov.

Sphecodes Hübner, 1822 . Type species: Geometra arcuaria Hübner, [1799] 1796 . [Junior homonym of Sphecodes Latreille, 1804 (Hymenoptera) ]. [ Germany].

Stigma Alphéraky, 1883 . Type species: Stigma kuldschaensis Alphéraky, 1883 . [ China]. syn. nov.

Synelys Hulst, 1896 . Type species: Acidalia enucleata Guenée, [1858] , a junior synonym of Phalaena limboundata Haworth, 1809 . [North America].

Trichoclada Meyrick, 1886 . Type species: Trichoclada epigypsa Meyrick, 1886 . [ Fiji].

Triorisma Warren, 1897 . Type species: Triorisma violacea Warren, 1897 . [ India].

Ustocidalia Sterneck, 1932 . Type species: Acidalia adelpharia Püngeler, 1894 . [ Israel].

Zygophyxia Prout, 1916 . Type species: Zygophyxia tornisecta Prout, 1916 . Somalia. syn. nov.

ablativa (Dognin, 1911)

abolita Herbulot, [1956]

abornata (Guenée, [1858])

accentuata (Guenée, [1858]) *

accenturiata (Walker, 1861) [misspelling of accentuata ] dentigerata Walker, ([1863])

exgiuaria Prout, 1932 [misspelling of exiguaria]

exiguaria (Walker, 1860)

exiquaria Janse, 1917 [misspelling of exiguaria]

rudisaria (Walker, 1861)*

accessaria (Herrich-Schäffer, 1852)

recessaria (Guenée, [1858])*

[Unnecessary replacement name for Acidalia accessaria Herrich- Schäffer, 1852.]

acentra (Warren, 1897) *

acharis Prout, 1938

achroa (Lower, 1902) comb. nov. *

[ Emmiltis achroa Lower, 1902 is transferred from Dasybela to Scopula .] achrosta Prout, 1935 *

acidalia ( Holland, 1894)

acinosa (Prout, 1932) comb. nov.

[ Antitrygodes acinosa Prout, 1932 is transferred from Antitrygodes to Scopula .]

actuaria (Walker, 1861) *

ssp. nigranalis (Warren, 1896)

parumnotata (Warren, 1898)

ssp. sheljuzhkoi Wiltshire, 1967

acutanellus Herbulot, 1992*

acyma Prout, 1932 *

addictaria (Walker, 1861) *

adelpharia (Püngeler, 1894) *

ssp. pharaonis Sterneck, 1933 *

adenensis (Wiltshire, 1986) comb. nov. *

[ Glossotrophia adenensis Wiltshire, 1986 is transferred from Glossotrophia to Scopula .]

adeptaria (Walker, 1861) *

eximia (Warren, 1898)

tainanensis (Wileman & South, 1917)

ssp. tenuipes (Turner, 1914)

aegrefasciata Sihvonen, 2001*

aemulata (Hulst, 1896) *

tawneata (Cassino, 1931)

aequidistans (Warren, 1896)

aequifasciata (Christoph, 1881)

aequifasciaria (Hedemann, 1881)

[Unjustified emendation of Acidalia aequifasciata Christoph, 1881 .] aetheomorpha Prout, 1917

afghana (Ebert, 1965) * africana Berio, 1937 * agglomerata Herbulot, 1992 * agnes (Butler, 1886) agrapta (Warren, 1902) agrata (Felder & Rogenhofer, 1875) comb. nov. * [ Trygodes agrata Felder & Rogenhofer, 1875 is transferred from Antitrygodes to Scopula .] agutsaensis Vasilenko, 1997 alargata (Dognin, 1901) alba (Hausmann, 1993) comb. nov. * [ Glossotrophia alba Hausmann, 1993 is transferred from Glossotrophia to Scopula . Junior secondary homonym of Aletis alba Druce, 1896 , requiring a replacement name.]

ssp. africana (Hausmann, 1993) *

[Junior secondary homonym of Scopula africana Berio, 1937 , requiring a replacement name.]

ssp. brunellii (Hausmann, 1993) *

ssp. capriata (Hausmann, 1993) *

ssp. zahmi (Hausmann, 1993) * albiceraria (Herrich-Schäffer, 1847) *

sulphuraria (Freyer, 1847)

ssp. vitellinaria (Eversmann, 1851) *

mannerheimatia (Erschov, 1871) albida (Warren, 1899) *

pura (Swinhoe, 1909) albidaria (Staudinger, 1901)

ssp. sankana Prout, 1938 albidulata (Warren, 1897) albiflava (Warren, 1896) * albilarvata (Warren, 1899) albivertex (Swinhoe, 1892) comb. nov. * [ Idaea albivertex Swinhoe, 1892 is transferred from Epicosymbia to Scopula .]

ssp. ancillaria (Warren, 1895) albomaculata (Moore, 1888) alboverticata (Warren, 1895) aleuritis (Turner, 1908) alfierii (Wiltshire, 1949) comb. nov. [ Glossotrophia alfierii Wiltshire, 1949 is transferred from Glossotrophia to Scopula .] alma Prout, 1920 * alstoni Prout, 1919 * amala (Meyrick, 1886)

lacteisabulosa (Rothschild, 1915)

parasira (Meyrick, 1889)

[ Scopula (Acidalia) parasira Meyrick, 1889 was listed as a valid species in Scoble (1999) by mistake ( Linda Pitkin , pers. comm.).] amazonata (Guenée, [1858]) ambigua Prout, 1935 * amphiphracta Prout, 1938 amseli Wiltshire, 1967 * anaitisaria (Walker, 1861) anatreces Prout, 1920 ancellata (Hulst, 1887) *

ssp. catenes (Druce, 1892) andalusiaria (Wagner, 1935) comb. nov. * [ Cinglis andalusiaria Wagner, 1935 is transferred from Cinglis to Scopula . This species’ status needs to be revised. It is considered to be a valid species by Redondo & Gáston (1999) and Scoble (1999), whereas Müller (1996) treats it as a subspecies of Scopula [ Cinglis ] humifusaria (Eversmann, 1837) .] andresi (Draudt, 1912) angusticallis Prout, 1935 aniara Prout, 1934 anisopleura Inoue, 1982 * annae (Mentzer, 1990) comb. nov. * [ Glossotrophia annae Mentzer, 1990 is transferred from Glossotrophia to Scopula .] annexata Prout, 1938 * annubiata (Staudinger, 1892) * annularia (Swinhoe, 1890) *

nigropunctata (Guenée, [1858])

[Junior secondary homonym of Phalaena nigropunctata Hufnagel,

1767, requiring a replacement name.]

ssp. reducta Rothschild, 1920 *

anoista (Prout, 1915)

ansorgei (Warren, 1899) *

ansulata (Lederer, 1871) *

ssp. adulteraria (Erschov, 1874)

ssp. characteristica (Alphéraky, 1883)

ssp. eberti Wiltshire, 1967

[ Scopula eberti Wiltshire, 1967 was downgraded to a subspecies of Scopula ansulata (Lederer, 1871) by Hausmann & László (1999).] antankarana Herbulot, [1956]

antiloparia (Wallengren, 1863) *

[ Acidalia (Scopula) antiloparia Wallengren, 1863 is possibly a junior synonym of Geometra (Scopula) minorata Boisduval, 1833 , based on genital examination of type material at The Natural History

Museum, London, and Universitets Zoologiska Institut, Uppsala, Sweden. Prout (1929–35) suggested that A. antiloparia (Wallengren) could be conspecific with Scopula sincera (Warren, 1901) . The opinion was based on a description of A. antiloparia , not on examination of the types.]

anysima Prout, 1938

aphercta Prout, 1932

apicipunctata (Christoph, 1881) *

arenaria (Leech, 1897) *

apparitaria (Walker, 1861) *

approbata (Warren, 1900)*

atomaria (Warren, 1897) *

responsaria (Walker, 1861)

trias (Warren, 1904)*

floccularia (Herrich-Schäffer, 1870)

[ Acidalia floccularia Herrich-Schäffer, 1870 is considered to be a junior synonym of Scopula apparitaria (Walker, 1861) (Becker, 2002) .] arcuaria (Hübner, [1799])

arenosaria (Staudinger, 1879)